Professional Documents
Culture Documents
TUBAL
RESEARCH INSTITUTE. NEW DELHI
/).. 9 7/b/tiC
TRANRAOTIONR
VOL, XLIV
(NEW ISSUE)
WI!lLLIN<:iTON, ~ Z
lOHN lIIACXAY, GOVERNMENT PRINTING OJl'FICF
WILLIA~' W)''1LI'Y AND SON, 118 JOsSBX STBBl.r, STIU,lIn, LI)NI)Ol! W0
',;,c,.'--\>'_-'
,
• ,
,
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... ''';
I'
Hoob.el II IllVt.'StiglltlOlll-, IJI 1'W\\ Zl',LI,.IIld hotn.llY l'\tolltlt.'el JUI Illtu tllt
"lIIlXtioR," \\hon hi~ "Hmlllhool, of the No\\ ZOll.lILlid 1"1otll" (IKtil-li7)
appea.red. 'l'hib WUR UO Illt'l C I ol'l"oelucl inll of hil'l 101"1111'1' works, 1"01
many cololtia1 collcl:tol"S Il11el 11t'1'1 1 hun1 at, "ork «'01('11"", 'l'I'nvl'rll,
Ila.ast, Hecto!", Buchanllll, &1.'.), I1m1 1\ vl1s1 Cjlllllltity of fI'c"h Illllle'l ia)
awaited examination. Itllk~'lI, Iho tusk \\118 olle of 110 IllOlLl1 IIIn~lIilllde.
and when the marvc]]clUs IIC'Cll1'UCY of the dOBCI'ipilOllS iM ccmbidl'rl'c! it
ill lU\I"d to helieve tho ract that Ihoy wel'O rll'n.wn up fl'OIll lwrh:u illlll
material alone. AMSIll"euly, well might IJal"win l.!A.clu illl , "Oh, my
heavens I to get up at second hand a New Zealand Born-that ill work."
How original the trea.tment was il'l flhown by Homsley's cOlllputation
that sixteen endemic genel'a anrl hali the species descrihecl havo thc'
aHh: " IIouk. f."
The indebtedness of Nell Zealaud science does not end ""ith nooker'r.
publishtlil work, To all r.e!'ious iJlvehtigators or the fiorl!. ho was n
friend, guide, and counst.'llor Thc!'t.' is, indeed, no nllollle of momeJlt
in the !att'r botany of the Dominion but is dee}>ly indebted to Hooker'lI
iufhwnce· mel I\lIl:1i",t,lllOO, gcn('l'ously given,
Almost to the In.st did the grcat hotnnist feel keen interest ill the
progress of knowlodge in tha.t fa r-ol'f r('~ion ",II ere hib spurs lwei ht·(·n
WOII, WI'iting to the Council of the (!:l1Ih'lbury hr:mch of tho Institllte
on the 24th June, 1910, he slLill, ill 1'(.'1'('1'01\1.'0 to the l'l'ccntly publi"lll·tl
., SuhUlltll1 ctic IRll1nns or N('" ZoalIt1ll1," "r was II.wnr(.' of tho lICi('ntific
expedition to the .\ucklnlld and ClImllhe>ll Jr.latub, OI'~lllIizorl in I!Hl7.
and WOR looking anxioll'Jly fo\' Romo records of itt! l'cHults." .Aft('r
explaining that, of courlle, ho was &pecilllly int('r('~t('Ci in the botany,
hE' I\dds, "Thero is really no section. hiological and gcologicnl, which
I can afford to overlook."
Hooker's work on New Zealand botany, to whioh tho above IIketch
does but Bcnnt justice, el.-OOnding over II. period of nearly forty years,
I'eflects but a. portion of that galli.UM 0.1111 uutiri.ng imlUl.Lr,~ whioh have
80 strongly infl.uenced botanical rCflearch thl'Ou~hont the Empire.
TA. (!()CR.\YNJa.
H. C. FIELD.
HINU Cr,AYI,ANDto. FI1~Ll.I was btlru at Holyboul'De, HllrnJlshh'!" 1~lIg1ll11d.
in 1825. He rcceived his OdUOlltioIl at 8tnckwcll r.I'UlnllIlLf S('}\olll :lfId
the Cit.y of J..ondon Sebool, a.lId completed the scliuluMtic p()rtiflll 11£ hill
life by a COUrRtl at King-'M College, LOllllon,
Being destined fOl' the p"ofebSion of u. oivil engineer, he was u.rticlu()
to Sir .Tohn Rcnnic, whose llame is well kno"n all over the worlc1 ill
connection with great works of engineering, and wllo WU.S t,he huileh·r
of the presont London Bridge.
At the age of twenty Mr. Field l'ntel'ed tho service of lID lenglish
nilway oompany, where he remained for ten years. The colonial! then
attrllCted him, and he came out to New Zealand in tho ship U Simla,"
nnd proceeded to Wanganui. The yuung settlement wus t.hen governed
by U Town Board, which appoillted Ml', Field its Clerk lLull EIlgiul"""
Sub"O!]nolltly he aot('11 a.lso as Engineer to th(.' nou.d BUBl'dK (ll tllt' (lill·
trict, I1ml Uil OOllslllting Engineel' i(l the Rangitikei ROllIl BOILl'd,
By the (,OUlltl uchion or good t'OlllllluniolthunY Mr, Field left lUI, 11111.1'1,
all over thc district, lJeing l'ol>pllnsible for the construction or no le~b
than ho tJlIIllbaud miles of l'llIlds, His name is perpetuated by .l!'ield'lI:
Track f,'oul Wall!:\,lInui to Kudol, thc half of "hich ntllll'C'<,t tu WunganUl
now forms pIli i of the Pal'apara Hond. This trllck hILs boon and is stilI
used uy ihOllMtndll, nnd is kuown nil ona of the beo;;i sUI'vayed and gl'a<ieci
roadways in the oountry, ('veil though pnlt of it nevel' got lwYOlltl the'
track stage. In li:!84 MI'. l!'iold I,(·til'ed flOm IIctive pm"mits.
Ho publisht'd severnl 11111)('1'1'1 on I'tciC'lltitic e.ubjects, and n book eutitll'lI
.. .l!'Ol'lU, of Now Z(·u.lalld," which giveh ,\ popul,tr ,LCCouut of the ff'l'ns II,
these hlandl:l find its imlllcdiuttl dCPC'llCll·llcieb. <IUd is noted for its gontl
descriptions and excelleut illul:!trations.
He was an Itrcleni ~uPl'Ol'tel' nnd ('xhibitol' of the 1I0lticuiturllI
Rociaty, and took a groat intel'eRt in harbour mattel'S and public dail'lI
gencrolIy.
He died at Aramoho. Wangallui, at the advanced Il~e of cightv-SPTeIl
OONTENTA.
L. BO'rANY.
'I'B~NIlA(·TION'I.
II. ZOOLO~ Y.
TI:I.ANKAI"l'IONII •
•\'RT. \'1. A HrI'Vl'tion eli tho (JlaHSifioatiOJl of the New Zealand Oal'4fltifHna.
By IcJ. Moyrick. B.A., F.R.S. ., 83-10'1
VIl. On the Nomenclature of the LepidopttrrJ of New ZeeJa.nd. By
O. 8. LongHtR.if, :M.A., M..D•• F.R.S. lOS-Ill)
vm. DebCl'iptionll of ThreI.· New Species of LepiiJoptera. By Alfred
Philpott.. .. .. .. .. .. 1115-116
IX. Descripti(J1\~ (If N,'w Zoaland lApidoplera. By E. Meyrick,
B.A., F.R.I::I. •• •• •• •• •• 117-126
X. Notes on HOme Dragon-.8ies from the Kerma.cleo Isla.nds. My
R. J. Till.ya.rd, M.A., F.E.S. •• .. •• 126-127
XI. MiMeellaneous Notes on lIome New Zee.Ia.nd Orustacetl. By
(,harles Chilton, M.A., M.B., D.So., F.LS. ]28-135
XII. Report on S'IUldry Invertebra.tes from the Kermadea IsJands.
:s.
By W. Benho.m, D.So., F.R.S. •• •. .• 13G-138
XX. Notes on New Zealand j1'il!hea: No.2. By l!lIlga.r R. Waite,
F.LB. 194-202
viii CO'lltelits.
ART. XXI. Now ~peoies ot IJI'pidO'}JtP.'TII, with Not&! on tho .Larvoe a.nd PUpa!'
. I·Af.I~h
of HOml.' Now Zco.lnnd Bntt.PrflitlJll. By Gt'orgl.' HOWL'",
F.E.H., T~.r•. R. 203-2011
,{XUl. The Ol'll~'l'arhil' lteilitiollliliiph 01 the liirdR of I.ord IIOWl',
Nol"(on., 111111 fJll' Kl.'rmudl'l' 1~I"ndR. By W. R. B. Oliver.. 214-221
XXV. Not.oM IlII N('ht. l.iCI."hi~t()r.v, mid n"bitK of ]Jliqn8 rfMillclllN, R
NI.'\\ Zl'ulnncl 'I'mptioor Spicll'!·. B~' .J. B. <lnh'nhy .. 234 ·240
'{XVI. SOIllI' Ji'I'lLhm'~ II[ Mil' <'il,(·IIIII1.(II), Ny~tl'lII IIC /lip/flirl/IIII "il'rlltu
FIII"Ml·. B) Pl'llfl"ohnr II. B. Kirl-, M..\ •• Vic·tnrlll ('()Ilc'g<'.
Wl'lIin~lIn .. 241-244
XXXVlll. UII I~ ('oll('otio.ll oJ: .J/IIUopilaga from tho Kl'rmlllil'l'h. By
T. Harvey Johnston, M.A .• 0.1'10., Quoollldllnd Univl'lhiLv.
Brillba.nc. Mel La.unt·('illt. HnJ'riI!on, SyrlllOY • !lO:i-373
XXXIX. VIlHOll1n.r System of Bipho/iUlitl obliqllaln Sowllrby. Uy A.•J.
Cot.trell. M.A •• M.Elo. •• .. •• •. •• 374 -379
XL. Descriptions of Nl'w Genera Ilnd Npll(oieil of Ooleopll!'l'fl. By Ma.jor
T. Broun, 1<' RR. .. .. .. .. 37!1-440
PBOOIUCJlJ NUll.
NotB on fihe HPCOil'h IIf I{J/,Im fOllnrl. in New Zl.'amnd. Br UilIK'rt
Archey .. Pn.rt I 2rl-28
Additions fill tho Ji'iRh ~'allna ot ~h(' Korma.d.·(· IHmndR. fly ICclAI\J' B. W,\i1A.'.
F.L.S., (.Jura fiar. Uallt.erbllry MIIMlUUl •. .• • . Pun I 2H-2CJ
NlltOio on tho NOlUollolatur(' IIf the Nt"w Zeal.md Gtumill-idl/~. with I>IlROrip'
tion of 0. Now RJK'ci(,H. n~ 'L. R. Prout .. !'nrf II 52-r»4
111. UI110LOUY.
i'BAl'I'SAOTIONB.
ABT. XIU• .Earthquake-origiUll in tho South-west. Pa,l'ifil' in 1910. By
George Hogbon, M.A., F.RS. 131t-142
XIV. Fluotuation... in the Lovel of the Water in KOIUO ArtoHian WelIH
in thl' Ch.riRI.ohur<'h Al't'o.. By ll'. W. HU~on.<1orr. M.A••
D.So. 142-159
X VI. Some- Rooks (If .l\luunt (Jal~l. l)ullIldin. By.r. A• .Ba.rtrullI. M.Sc. IU:t-179
XXII. The RaisE'd Bea.ch(·!1 of Cape 'J'ura.kira.o. By B. ('. AH~on, (".1.0.,
It.n.H. !!OM-213
XXIV. A Proliminary AI'('oUIIL or tltl' Lower Waipa.rn Unrgn. Hy B.
Speight, .M..A.. M.so., F.O.H. • • •• •• •• 221 233
XXVll. Note!! 011 Wllllilll,rWn Phyllio~rn.phy. By ('. A. (klt.tnlJ. i\Ull·.,
Victoria. (Jollog(' 2~1i-266
XXXlJl. Nopholin.it(l nook.. in New Zca.llLIld. By PJ'Uft'HKUf MIlJ'Hbo.ll.
D.8o •• F.G.S. .. •. 304-307
XXXI V. 'rho 'Discovery and Extent of Itormor (lIo.f.. ia.tiOIJ in tho 'l'Ofo.nlll
Ra.nges, North Island, Now Zoaltmd. By G. L. Adkill •• 30lh'U6
XXXV. The Geology of the Bluff, N~"W Zel\1and. By L. J. Wild. M.A... 317-339
P.aOcm'JDDLNGS.
Typioal Sections showing the Junotion of the Amuri Liluesfione and Web
Pass Stone at Web Pass. By C'. A. tJotton, M.So. (AbRtmct) .PM'!. rn R4--8lI
APl'lBNDIX.
Records of Milne l:!eismop}lhs. 1906-1911. By H. 8'. Slroy. B.Hc•• lind
G. U:ogben. M.A •• F.G.H... " 441-467
(}ontell t.,. ix
PRI)('l1lI!lJHNIJH.
v. M[SUJj}LLAN~JOnS.
TB\'IOI \CTfONh•
PlWU.I!IIIlDrNliI".
PROOEEDINGS.
[Following p. 457.]
PART L-Iasued 30th AugUbt, 1911.
U.-IsAued 8th January, 1912.
m.-IBs\IL'Ci 10th Juno, 1912.
LIST OF PLATES.
Plat.t>m-
~'ig. 1. 80pk0rt1 ,niN'Op/&yUtI
Fig. 2. PitIoBpcwum diuaricatu/II
Platt' IV.-Podoca'plIlI nivali.. "
Platu V-
Fig. I. Veronictj cAatllllmica 22
'Fij(. 2. VI'f'Onil'll 10flfl'llioidl'lI. 1'. I'IIHRinioidlo.s, and V. tl!lr.ltlllllfl 22
Plate VI-
I<'igs. I anrl 2. Nopoom Ieb-IIpklNI •.
Plate VII-
Fig. 1. AriBtml'lia /ruJiC08fI
Fig. 2. PlMnantia corymboBt,
Plate VlII.-Pitt08f1M_ IUI~lricn"'tu
POPPltl.WIIII.L,D. 1•. --
Plate IX.-
Fig. I. Ruggod Islamls. from the nortrh; Uod6sh lsland in the distance.. IiO
Fig. 2. Norfrhorn aspect of one of the Rugged Islands. ahowiJul; Oktwia
a1/.f]'ll8U/oUa and O. Oole.naoi .• .• •. •. tIO
Fi~. 3. Rugged [sIlmds (WI!'a.thf'r Aido) 80
AIITON,.8. ('.-
PlateXllJ-
li'ig. 1. ()orYIltJoor}l11I! .lHSOOia.tion• .Pa.lIiser Ba.y • . 2015
Fig. 2. OoryIIDr4f1lU8a.nd MueAletIbeckilJ~lemaasooiatiollS. 'BoMh No. 4. 2()8
Fie. 8. Pc)nd formed immedi&tely above Beaoh No. 2 208
JI'ig. 4. Beach No, 1, t'levo.ted a.t 1855 earthquake ., 208
Plate XlV-
.Fig. 1. Bea.oh No. 4) (95 ft. a.bove _I 208
.Fig. 2. Boulder Plain 'With No.8 Beach (60 it. above 110&) .. 208
J. B.-
GA'l'BI!IBY.
Plate :x.V.-Nests, &c., of Mil/all cUBtitI,.". a New ZHlalld trapdoor spider. , 240
xiI List of Plr"tS\.
KIi!.K, H. B.-
J{lLW"~
Plate XVI- 1'.1.0&
Ii'ig. I. Din.!(rlllnllllltic repr('M'lItnhon IIf thl' c·jT<'ldatory lIY'Ktcm of /lepta·
Ir~1It1/ rirratll .. 1!ol4
Il'ig. 2. Purl of !.he C'lorhl,1 vC'II""I14 ILncl Lhl' 1J('llhridinl lIy.ll'lIl. from th.,
t1ofR,,1 Hidu .. .. • ..' .. . 2~
l'III!.l· XYll. -
F~. 1. 'I'hl' C'!Tl·I'I.·lIt bruuc'hil~1 YChhl'J, 111111 tile nnLl'riol' (lILrt uf tilt" ciol'Hnl
,1Ort.il' hYhtl'III, ffllm f,(1l' cllll'hUI BHpIlf't 2401
Fig, 2. 'I'hl' afferent bruoc'hinl KyHLmll, fmm th(' velltral aMlwrL 2,1-1
Fig. 3. Right afferent brn.nl'hial V(·hhCI.... from tho rigbt I!itll' 214
Fig. 4-. Anterior part. (If PCIRI-clIorclinal hyllLl'm, Hhowing ('onnl'rtion of right
Hinull with pOrta.l heart. . .. .. .. . 244
Fig. 5. 1'hl' j1ll!:111Rl' HYflWIlI, di'4AC'ctl'lI frum venlnl lIoIIJIoct 244
AmUN. G. L.-
Plato XXll-
Fig. 1. Ganeml vio" of t.hl· .ul,lI'itllrt~1 Jlurl ttt P.lrk Vallc'y :108
Fig. 2. Waiohinll-iti V"Uoy :1011
Plato XXlII-(lJu.ciat.ec.1 bC'dd (lC 1',,1'(.. VII III'S :112
P'late XXTV-'I'be largetrl !.!;11I.I·inl hllRl(11l1l. \·nll(·y ill Pnl'k Va.lley :112
TRAN SAOTIO NS,
'p HAN SAC rr rON S
OF TIIF:
1911.
PJatc9 J-YIII.
T.\BLE OF (iON'1'ENTS.
I. lnh\llluGtion,
11. Elt'montary 6pooill'S.
HI. Variation.
IV. Mutation.
V. Epharmouy.
]. General.
2. Fixity of speoiet:l-pla~t.Joity:
3. ReRpOll'lO to ecological faotor~.
(n.) SIIil.
(h.) Liltht.
Cr.) Wind.
(d.) Water.
(c.) Altitude.
4. \ft<>r-oift'Ot of Htimuli.
5. Co11VOl'lJt'llt <'pharmouv.
(n.) The diva.rirating IIbru}) lorm.
(b.) Th(.' eu.mioll form.
(1'.) Th(.' ha.ne form.
(a.) ThE' pl'OIItratE' (orm.
II. Pt'Nlhtl'llt jU"f'nilp form~.
\' I. H.vbridizaiiol1.
\' 11. 'I'hl' Ittruggll' tor OXU.tE'lIC1e.
"111. J)i~ribution of H}lt'Oie04.
1. Di'1f.ribution in ~ICl·a.l.
2. lBOla.tion.
IX. }<Jvolution in t hI:' geJlIlM Veronit'll.
X. Colleludil1g roma.rkH.
XI. I,ist of litE'l""ture cited.
I. INTRODU('TION.
PLA.NT-ECOLOGY is concerned with the study of plants as Jiving orga.nisms,
not in the laboratory under artificial conditions, but in the field as they
grow naturally. Like every branch of a great science. its (:ontsut is not
bounded by any definite Jimits. but it intergrades \vith '\"arious departments
1- TrlouB.
Trallmctioll8,
• ~he oonvenient term "adaptation" is used lilroughout this p"PfT in 1.1 nOli-
teleo1oaioal sense.
t This statement applifllS rather to the eoology of p1&nt-dir.tributioll than to that
~ and more funda.inental stuclY of Ufe·reaotioD.II knOwn &1\ " biology" by German
lnveetigatortl. In this latter IleUse Darwin hirnaeIf stands pre-ominant &8 an eoologiM.
t The suba.ntarotio and the fIIUba.rotiO olimateR are by no means identioal, lntenlot'
cold plays no part in the first-named, it.. main characteristiet. being lAck of lIunshinc.
frequent cold gales, oow.taut bbowe:n.. and a 'low &vera~ temperature aJl1lbe yr.·ar, with
but little frost in winter.
('OC!iAYNE.-Erolo!Jlc·cil Studl(s m £I'01I1flOll. 3
500 I!m. and other parts less than 30 <:m.; the plant formations include
mall~rove swamp, rain forest, heaths of various kinds, subglacial fell- and
herh-fields, varied associations of rock Ilnd debris, subantarctic southem-
beech {orest. associations in and near hot springs, dunes. salt meadows.
st('pP!'s, swamps. and moors-in fact, for an equal variety an ecologist would
have to !·xplorr one of the hlr~('r eontiJwllts in its entirety. Further, the
isolation of the l'('giOll for a nst period of time fur from any other land-'
surfnC'!'; the abst'nce 01 graZillg animals. the moll. (IJinomiB) excepted:
the diverse floral clements (Malayan, Australian, Subantarctic, &c.); the
strollp, endemism; the numerous small islands where conditions are simpler
than on the larger ones; and, finally, the presellee of many areas whose vege-
tation has been ('hangt'd within a very feW' years through the farming
operations of the settler, and its components replaced by exotiC's of quite
difi(>rent growth-forms-a.ll these attributes much enhance the importance
of New Zealand for ecological research.
Now. although I welllmow that the final court of appeal ill evolutionarr
matters is experiment, still it seems to me that some few details having
a bearing on vllrious phases of the evolution question selee·ted from numerous
observatio1ls on a vegetation and a flora that olle may venture to designate
.. unique" may perhaps be worth the attention of students of descent.
• This is not very different, after all, from Darwin's view, who declartod that" a
well-marked VAriety may therefore be oODJSidered a.n incipient speoiea • • • the
lerm 'species' i~ ono arbitrarily given to a. at of individuals olo!Iely reAeDlbUng l'&Oh other•
• nll th8.t it does not eft"1\tially differ from the term ' va.riet,.'" (Darwin, 1899, p. 39.)
t And then a.oooptins thi'! M a sJlPOi~, it is tqSid 10 be' mreDlC'ly va.riabll'."
7'r(m~l/rfilJlI\
forms, &c., may iurilldtl ('VC'1l mort> than OtiC LIIlIH'ILll Hlll'C'll'l>, .till cLppt'cLn.
to be the caso with PitU)8porum riyidum Hook. f., lib dcfill!'d by Hook"I',
Kirk, and CheeselJlun. Pilitc I sbows this eab(' ('Il'ul'ly, wh('rl' th(' tYPI'
of P. ri(jfdul/I 011 th(' righl diffl'I'1:I most Ulul'kl'clly rl'Cltll lIw l'ummOIl Houth
IlIlAnd form on the Il'ft, wltidl LlUn naming P. tlivtlril'cullm. '"
In sornl' l'o,lIt>S til<' llifli{'ulty c", til Ilistillgllillhing-IIIlIlIl'H II> md h,~' tlH'
.. creation" u{ •• vllril'til'fI"; hut IltellC', ("!Inin. (Ir,' II/ I/llill' diffrrl'llJ ,'alU(>,~,
cma may betony to disti.nct billlClgiCftI C'a/I'(foriI'H. A rew e'xllmpll'H tclkl'lI
from the" Mauual of till' New 7..enllUid FIlii'll" «'h('I'Kt'mnll, 1\)06) will
explain my mea.niug.
1. II()heria pOpuZ1/Ra A. Cunll. (p. 78) il:l divided into till' 1hl'I'I' V.II'ic·ti('K
ta) 'Vulgaris Hook. f., (b) lal'leeoiata Hook, f., nud (e) MI!lltllli/()lia Hook. f.
There is no such plant in I'xisteucc U8 If. pOpUlYltfl, fOl' l,h(' u('tlcriptioll
includes the three vari('tics (fI), (b), untI (e), ('at'h oi whi('h, hOWCV!'I', if>
distinguished by a spedal diufl,llllbis, the VI\l'il'tit's (1'), (b), lind (e) TCllpl·(,tiv('ly
representil1fl, distinct ~roups of illdividulllK whil·II I'{'prodn('(' thl'nls('lv!'tI
true from s('ed.
2. ('c67mic1laf'/ift Eny8i! T. Kllk hUK II vllrlt'ly IIrbiclI/llllI '1'. KlI'k (p. L1l).
Both the' species l\lld its varit'tr url' d{'I'l('rihl'ci. Bul ill thiH I'UI>(' till' "pl'('ilil'
description rt'fers to Ollt' l:Iet of individualll POIISI'IIHinl-l ('('rtllin ('hluIH't!'rll,
which is (I. En!ls!'i, Pl'OP('I', und dot's not ilwlnd(' VUI'. ()I'bi('ulr~fc" whic'h ill to
be recognized throup,h ita hllVillg (lthl'!' c1l1n~I('tcl'S Iti1flC'ut in ('. R'»!llIii ]11'111'('1'.
which latter may then lx> tt'l'l111'd tlw t,ypl·.
S. Epilobiwm jU'Y£Ct'"m Sol. hlLl:I VIU'I:I. ritlere"m Hausskn" IlirtiUI'rIUl1
Hook. f., and maorophuUum Hu,ussku., t'uch 01 which is dofillc<l at c'onsidl'r-
aLle length (p. 175). But none (If tilt>tle names repl't'scnts a biologil'nl cutity,
for E. junceum, to quot!' il'OIll (1}1C'eseman, "ill I~n l'xtr!'ml'ly val'inblt·
plant, the numt'l'OUB forms of whie'h may be groupod ill tht' throc (ollowing
varieties "-i.e., as above. Furlh('r he writc.>a, "The t'xtreDll" Ktl\tes of th('
above varieties havc a very distinct appeal'lU1('C', n!ld might haw b(,l'1l
treated as species W(,l'e they llOt C'OllllC('ted hy I1Um(,I'OUS intermediate' fOl'llIH,
which make it quitl" impossihlc to draw stric,t liIl('s of demar<'lItion betw('('n
them." Here, tht'n, th(' desllriptioll of the spt'c'il.'s dot'S 1I0t indic'nt(' u ty}>c"
but it inoludcs till' Ihret> tl(ff"iefies nfl.({ fill fliP itltt'Nn('d,'nlc' /""''''11, while i Ill'
vn,rieti611 thems('lws a.re likcwiSl' 1101, diHtinl'1 I.'lItitit·K, t lIud lll'i<HIA to n
different biologic'ltll·u.tegory to lIw Vllf. Clrbwulrtfa of (', EI~!lsii.
4. (JaUltl,tl"ifl rUlJH'8tris R. Hr. (p. ·107) iK Ib tlil1lilll r I'Xlllllplc' to till' II'Ht,
boing said to bc' .. a highly vurin bit, p,lllllt. the nUI\Wl'OUti fm,otl of whit·1i
lUll best arrllllgcd ulld('r tW() la'nels '. lII\uwly, vnr, itltlCl'II/flftl ('1ll'l'Ill'UI.
and val'. ptl.rui,/oUa OhC'l:'lIelll.
5. In certain ot,h('l' ("\SCS, wh!'r(' tllt'I'{' IIrc II h()st of illh'l'gl'lIUitlA tormll,
tho most div(,fgt'nt n1'e trcl.lted aLi l:I('parate spt'dc's noiwithl:llluuling thltt thlly
aro connected by illtermediatel:l. An cxamp)(' of thil:l il:l VI'l'tmil'll p1'Y1{!ui/()lifl
Hook. f. and V. BuchfMl,(l.'f6i Hook. C., of whit'll I,\U~I' spoc'it'K CheCI:IOIDal1
writes (p. 527), .. Larger form" appl'Oach V. 'P~lJ,i/c)l1'(J so clol:lely that
it is di.fficult to draw a lille of demtJ.rcatioll betwet'n the two species.
My var. 'fMIjor might be re£t'rred to t'ither." Othel' examplefl of simill~r
• It seems poas.iblu also that P. iliva1',catum COllHU,tK of two eJAomonta.l:Y IlpeoiCll, fOWld
in the steppe and. fOl'llBt OlillUl.Wb of tho &uth L!Jand l't!'I))E'Otively. &e Pla.tt' VIll, and
oom.]l6l'8 it with the figure in Diole (1906).
t Biologioa11:r. SOlD.(! are OIIrtainly eliatinot ontiti(;R, &II, t.g., the varioty n_pllglllltll,
which Petrie ha.. • m.&<le " into a IlpooioH unclor tho name lff. Srffl661ll, ane1 whioh is greatly
on the inor.m" WllOro foro'lt i~ b3inl{ removell ill tho Waima.rino looality.
TBU!" N.Z IN'T. Vor•. XI,I V PL\T~ I
1""<"1 11./1
TRAN ... N Z INS1. VUL XLI V PLAU II
}<'lO. 1.-T11R1lK 1<'011"" ()I '1111 ' HI'I ell .. " V),b()NIC'A 1I0~!I'III,rA.
trclltnll'llt Ill'e Oleolla IIaasMI Hook. f. and O. uleifolia ']'. Kirk (p. 290),
Rammculull Sinclair;i Hook. f. and R. qracilipes Hook. f. (p. 18), and
POll lIeficlilmili Pl'tl'il' and P. pUIIllla Bl'rggr. (p. 905).
1>. l'mmica lJllzi/olia Bellth., ilK originally dl.'nned, prohably referred to
,~ quitt' dennit(' B~t of iudivieltULIII Evell hy Cheeseman (pp. 522, 523) the
IIpl'('i~1:I ill IIpoken of I\I! II .. plant," and not ... s ... varying lIeries of fOrIDs.
FUl'thl'l', thl' flpl'l'i('s is d('tilll'Ci al! "erert," nlld but OIlr variety is allowed.
III puint of fll<'t, howevel', t.ht' .. SPOCllll!" ill<lludCb th1'e~ distinct growth-forIDs,
.It any l'dott', two of which, thR prostrate, Imd flit' low, erect, sparingly bro.nched,
dore shown in PI~tc II, fig. I. The var. odora T. Kirk (patens Cheesem.)
ill of tht' ball-like Arowth-forlll. In tMs f'Xumple, then, a taw07lO1'nto species
inoludes plants belon!l;,ng to at least tlif" ,thsolutt'ly distinct bioloyical categories.
And, in addition, it is highly p1'oba hIe that d dozen or more distinct trul'-
brecdill~ entitil's mil!,ht easily be sl'paratl'd from the heterol!,eneous mass of
individuu.ls known 11.1:1 V. bIlJYI'/olio.
7. Many val'il'ti('s lIore of a quite difiercnt physiological value to othera.
Home, 8S III cllses 1, 2, and 6. rept'odue'e themselves true from seed. This
I ha\'e definitely proved ill II !lumbel' of iIUltanlles; tkt'y are, in fact, froe
el('f1PR'll.tary IJpt'OWIl. Others, again, are mf'rely environmental (unfixed ephar-
moni(')'" forms, SUdl I\S are dealt with further 011, of which notable examplos
lIrt' tht' var. prostralat Hook. f. of T.Jt'pt()8pf'TmWm 8coparium Forst. (p. 160),
tht' var. rhombifoUUilt Ilook. f. of RaMJ,nculus pinguis Hook. f. (p.12). and the
var. paupemtUS§ T. Kirk of RtibUH oillsoid"l1 A. OUJID. (p. 125). Finally, other
vari('ties rl'prt'sollt ~ IIl'ries of formll regarding the stability of which llothing is
known, bllt whi('h lUI' supposed, without any suffiril'nt reason, to be uIUltable.
Without gOiIlg iuto further dt·ta.ils, it is evident that the species of New
Zel\land taxonomis1s m'U rather the creation of man than of Nature. In
aaying this I am not. hyp01·critical. The main object of a flora is to enable
,\ plant to be rC'a.dily identified, Ilnd this, from the very nature of the case,
demands 1\ more 01' II'ss Il.l'tifieial classificatioll. Where such precise and
('opious informatiou I\S to vnria.tiOll is givell as in Cheeseman's most careful
dud .·Xf\(·t work tllcrc !leed be no mistake, ILnd the worker in the field knows
f'Dctly what h" ma.y expect. But, as a. rule, writers on evolution have
quite' negleCltOO. to distinguish betwel'll tllzonMnio and physiologicaZ species,
whielh In11;(·r I\lon.e al'l' their l·oll11l.'rn.l1
Although hreodill~-~xperjntents ('an alonc decide as to fixity of form,
ecolo,zy should k>1l IIOmethinp,. If tI. cl.'l'tain sot of illdividuals remain
ullc]ll\ng(>d IlVAr wid.' 81'l'lIfI, I:IU far as thl'iJ' spec:ific marks go, and under
\'nrying ('onditjonfl, it. may btl IISBUUlt'd with tolerablo confidence that they
reprodm'l.' their likl', Rllel art· thl.'l'efofl' spet·ies, e-Iementary or Linnean, as
the ca.se may b('. And perhaps it is allowable to go furtht'r, and sa~' that
if several allied plants grow in close proximity in sufficient numbers, and
preserve their distinguishing ('hara('ters, they are probably distinct, and
would romt' true from seed. A ('ase of this latter class is to be scen at
the low('r gorge of thl' River Wllimaka.rirJ, Canterhury Plain, wht're thl:'
vars. miN'0'}Ih1l11a Hook. f. and 'P"o81rata T. Kirk of Sophora tf'fraptera J. Mill.
I!.row sidt' hy side, and in this rase I hav(' proved t'xperimentally that both
vari(.'tics come true from secd. So, too, with certilin forms of A('«1'tI1I
SOIIlfI'Ui8orbae VahI. growing on subalpine fell-fields.
There is no nc(.'d to multiply instances such as tht' above; suffice It to
say that both from experiment and ecologica.l observations I am sa.tisfied
that elementary specie8 are very twmerou8 -m the NetD Zealand flora, especially
in certain genera-e.g., Oalamag'l'08tis, Dcmthrmia, Poa, Festuca, Scirpu8.
Uncinia, CUR'e:t, Lusula, ? Phor'fllitlm, Rcmunculus, Oardarnirte, Pittosporunl.
Rubus, .Acaena, Oa'f'tn.ickaelia, OzaUs, OoriUR'ia, .Aristotelia, Pimelea. Epl-
fob&um, Leytospen)I'IlIm, Ani80t0me, Aciphylla, Gaultheria, Dr(JC()phyl1U'lII.
Gefl.ti«M, MY080ti8, Veronica, Oop'l'08ma, OeZmisia, C'otula, C'raspedia. and
Senecio. On the other hand, many species vary to a slill.ht degree only.
sud are to hI.' recognizt'd at a glance.
III. VABlATION.
Apart from constant hereditary distinctions, there are . f the individUAl
differences," as Darwin ca.lled them (1899, p. 31), or "fluctuating varia-
tions," as they are now frequently designated. These are supposed to
depend upon a reaction of the organism to a change of environment.
Klebe (1910, p. 235) distinguishes two kinds, the one tt caused by different
external conditions during the production either of sexual cells or vegetative
primordia," and the other .. is the result of varying ex.terual conditions
during the development of the embryo into an adult plant." The two sets
of infl.uences ca.nnot as yet be sharply differentiated. The following case
illllstrates this difficulty.
Olearia semidefitatl& 'Dene. is a modetate·sized xerophyti(' shrub. which
is eonfined to the moors of the Chatham Isla.nds, where both thc ('limatil'
and edaphic ('onditions appear to be of great constancy (Co('kayne, 1902.
p. 288). The leaves va.ry on different individuals in size, shape, toothing.
n.nd tomentum, and plants grow sid(.' by side whi('h, 80 far as gent'fal llP-
pearanee goes. might easily be taken lor distinct speci(.'s. Probably here
the variations are genninal, but at the same time eaeh plant hRs its o\\o'll
rooting-place* and its individual physiologi('al chara('wr, 80 it ('annot ht"
denied but that each plant is subjected to slightly different stimuli to those
experit'nced by any other.
.A most important question is the heredity in :fiuctuating varia.tions
and the degree to which they can be accumulated. Darwin (1899, pp. 31,
32) considered them all-important. tt These individual differences," he
writes, " are of the highest importance for us, for they are often inherited.
as must be familiar to every one; and they thus afford materials for
natural selection to act on and acoumulate in the same manner as mall
... Romanes (1895, p. 125) writes, "Tb.a.t is to .y, it has 3QLl8 true to seed for the
laat forty ye&m." Roma.nes mentions this caae as an example in support of the heredity
of an acquired cha.ra.eter, but. Darwin (1905, p. 229) mentions it as a oase of" methodioa.l
seleotion."
t With .. speoies &uoh as this it reali, must be nearly impoSBible to judge under
European oonditions how far a supposed' wild" plaut may be reslly wild and not the
de~t of .. oultivated form.
~ The leaf-like oalyx of the primroses, &0., lmown 80S "Jack.iu-the.green" is a
remarkably persistent ob.a.ra.ater.
8 Tl'fln ~fl('fIOIiS 0
TVo MUTA'I'ION'!I/o
The)I' bc'ems to b(' 110 doubt but that De Vl"ieSlll.ll mutations llris(' fum,
tim(' to time. That slwb nfi"ord a bet1Rr material fo), preservation hy n.ltur.d
Kl:'lectioll Hum do small f1u(otuating variations is obvioUl!. Ullfol'tllnnh'ly,
the numh('1 of rl\lI(,R of vl'l'ltabll' mutants is small, while most hav(,OIoiginut{·d
ill cultivatioll. Thill lust fact discoullts the value of th(' mutation thllon"
in tlw opimoll ot m.tllY. t My own f{'eling, as all amatl'UI' I!.clld('uPloof m.m'·
yeaTS' bhllldill~, II nd as one who has C'ultivatC'd with his OWlI hands sevl'r.ll
thoUblind sped<·s of both wild and garden plants in an dlltipodclI.u p,.lfden
far from the- home of most, is that ordinary cultivation, without mUI1l1l'<'.
has little effect in producing variations of moment. In my gal'dell. plaut&
reproduced themselves :£rom seed freely and rame to maturity, hut 11(,\"1111<1
a number of daffodils and some, probably hybrid, dwarf phlox('t; (Ph'" I
IlIWulata L.) I remember nothing "new."
In estimating the origin of species by mutation, nothing but l'xperI-
ment can prove the heredity of the new character. All that el"Ology edn
do is to note striking varieties, their frequency, their environment. tIll:'
position of the individual possessinll. such variations with l'<'gal'd to Ilormal
individuals, and so on.
ThC' following examples of what may be- full or partial mutatIOns III
the Nt'w Zealand flora, indigt'lloUS and introduoed, roay bt' of illtt'l't'!!t ;-
1. The white forIn of M Y08otidi'lllmo nobile Hook.
The spt'cies is confined to tht' Chatham Islands, where it growl> on or
near the Bea-shore. In the normal form the oentral ha.lf of th(> l'orol1., illo
bright blue, which fades to purple, a.nd tho edSt's are more or 1t'Sf, whIte.
Mrs. Chudleigh, of Wharekauri, some years ago discovert'd one plant with
white :fI.owers growing wild ill the nOl'th of the main island, and although IIhe
is an excellent observer, and Myosotidiwm has been carefully noted ill itt;
habitat by Mr. Cox and others, no more white-flowered fOlIDB ha.ve h(.'l:'11
obsl'rvt'd. The plant in question is now fairly common ill (Ou1tivatioll, ,mdo I
lUld"rstancl, comes truf' from seNt So, too, does the normal bluE' fOl"Dl.t
• Soml>thing not wrv lhffl'ft'llt to the llIut.&LjUJ] theory waH propowldod by J. ll.
Armstrong, form(:'rly I)f tho ChriBtohurch Bota.nica.l Garden, in 1\ papor (l<'IIling with thl'
New .zc..a.ia.lld hJK!OieR of VCI'01&I"t'1l in 11181, in thl'h!' WOl'tlh: "I haVt' h<-<-u enabll'lJ to .,I!o.,('rvil
IIUmel't)U8 ~"rd(:,)l'RooilliugK IIf many of tho form~, and thl'y a..tmO!.t invlloliabiy l'IlIII.'mble
thl'lf pall'ntK. Hon\lltimos, ho\\,ovor. "POTU! appear, and whlln thih hapIlIlIl. .. tlwl"I.' h(I('mR
til bll 110 htl'Oll~ tenodl'no? 011 tho }lart of the "110ft to l'l'proIlUl'l' its!.'lf, and it npIlC;'a"" to
me loot it iK jUhl in thil! manner that the grea.t!'r numbt'r of our na.tiw fOrDlh hM l' IlCll"ll
pl'OducOO. •\t IIOW' v('ry dilltu.ut date thore Wl."l'l' prooo bly ouly t.wo 01· thl'CIl (pl'J haJ1l' unly
o
one) lI}JlOoi('ft oxiRting wit hill t h(' linuth of thl' oolony; but, 011 account of til<' I'xtl"l'mt'
ioeal va1'iatioll.l! of climatll and vari<><l geologIC!&! formation of tbl' RUrla.ce, 01.'1 tam \O,tria_
tions uccurrell, and a Ioport r:.o produoed, bOing III.'lf.f(·rtilC'. and ha.ving withill it...eif all
the o1emPJIts required for reproduction, naturally rt':vroduccd it.. likl' until all.oti1t'r 1111('11
sport ocourl't.'Cl. and thUII the formK gradually bt>l'aml" di&rontiatcd from th!' type, and
by a long 1lElri0ll of such sports one la.rgl· family of l'erom rllB hat. been formod. t' Thl'1l
o
hi! goes on to show how sim.i1ar muta.tion~ have taken pJa.ocs amongst spedE'lo of other
la.nds, and considers that the intermediates baVE' bI."en E'radioatcd " by man or the la.r~r
a.nimaJ6, leaving only in most _ thE' more widely diifel't'nt.ia.tod forma.'· But III
New Zealand man ha.R done little, and very many in~rml.'diate forms have b('E"Il. pre·
Rel'VOd.
t Klebs, however, writes (1910, p. 241). "Even if it is demonstratoo that ill' WOh
IlimpJy desling with the Hplitting-up of a hybrid, the facts adduced in no IIOU<oC lollf' their
very ~t value."
t Raising from IIeIld is, in fact, the only satildsctory method 01 Propalllltb11t I",th
1he type and the white-Bowered form.
('OIJK.nNIC.-Ecu1oglral 8tudu:b 111 E /)01I1flOlI. 9
• Mr. Cheeseman informs me that he also has raised the white form frtlln hOOd, &nrl
that none of the plants produoed flowers other than white.
t FifLY-lI8ven names are given in .. Phormiurn 'BftaQI &8 a Fib1'OUb Plant" ( Wt'llirJgtoIl,
1872). but it ill now known that many of tilem aro synonyms. Tllore are extenw.ve 001-
lections on somn of the- Uovurnmont experimental farms, where their hehaviour a& to
,·on~t,\noy. hybridization, &"., hi Ix>inll' studied.
10 7'ranllactions.
parts of New Zealand. The two best known bear thl' gllrden names of
L. Ohaptnanii* and L. Nickulsii* rl'spl'('tivl'ly. Seedlings in abundance
haVI' been raised from the latter by Messrs. Na.irn and Son, Christchurch, in
their nursery, and eve1'Y opporlwlity wall kindly afforded me of studying
their form, &e. (see CoC'lmyne, 1907A). Thl' colour of the original plant
is repeated more or less in tbe seedlings, but it "m'il's II good deal, and SODle
flowl'1:II are whitt'. Dark-('olourl'd lellVl!s, 1\ parl'lltal character, IL('company
the darker flowers.
III Ilo oase recorded by Cheese-man the phmt WIIS reported by its finde-r,
Mr. R. J. Gilberd, to come- true to colour (Cht't'seman, 1908, p. 275).
It is obvious that these crimsoll forms only appear o(·casiouallr. for
they are too striking in ('ontrast with the familiar white blossoms to be
overlooked by even a casual observer. Further, the change of colour is
deep-seated in the plant, since the leaves are also affected. In L. Nicholsit
Rort., too, the plant is of a weeping habit, as opposed to the normal erect
stature. Finally, it must be noted that the semi-mutants grew in widelr
sepa.ra.ted loca.lities, some in the SO\lth a.nd others in tht' North Island.
7. Double white torm of LPpt08pel"lnUtli scopanum Forst. (Myrtac.).
This was loWld growing wild on pumice soil in the Hot Lakes district
by Mr. E. Philipps Turller. The doubling is very complete, and, so far as I
could judge from much-damaged spe('imens, resulted from petalody of the
tltamtlllB. Probably it is unable to produce seed. This case is of further
illterest because double flowers, as De Vries has pointed out (1905, p. 489),
are exceedingly rare in the wild state, though so common in cultivation.
Only OM individual was noted. The mutation was evidently quite spon-
taneous, and cannot be attributed to any lIudden ('hange of soil-conditions.
Leptospermwm scopauium is 0. most variable plant. Doubtlee.b some
of the forms are good elementary species. The form ,vith pinkish :fio\'I"ers
and hairy leaves, &c., of northern Auckland, which occurs over wide areas
side by side with other forms from which it can be re('o!!,nized at II alallC!!.
is a ca.se ill point.
8. 01.earfa semidentota Dcne., 10J'm with white :fiorets.
The typo has brilliant purplt' ftower-ht'o.d!!, Thp \vhite form waR dln-
('overed growing wild by Captain A. A. Dorrien Smith. It is now in
('ultivation in the' gardt'll tit Trl'sc'o Ahhl'Y, tidily.
A similar <.'1IoB1.l is VILr. Detl.tiyi (10('I.o~ynt· of Olearia chat/lmmca T. KiJ'k,
lound on Pitt IsII\lld by Dr. A. Dendy, F.R.H., u.nd whit'h has purpll' fior('ts
Imd yellower denser tomentum on thl' undt'r-surfact' o{ the leaf tllllil the
typt" the lior(\ts of wltiC'h, 1ll0reoVl'r, !trt' whitt' fllding to purplish.
1I. lIletrosideros,llUJ'i,do, Menzies, form with white flowers.
The type has crimson flowers. The white-ftowt'r('d form has ll(~en founel
in two places, one plant which I have seen growing llear the hend of the
Otira Gorge, Westland, and the other lower down the valley.t
10. Metrosiileros to'l'llefltosa A. Rich., form with yellow flowers.
Mr. H. Carse (Cheeseman, 1906, p. 1137) discovered one specimen with
yellow flowers, those of the type being dark orimson, o.t RanQnlID\l
Harbour, northern Auckland.
• Because 1 u~e these garden na.m.os it mUllt n:>t be oonoluded that I conRider the
plarlts of the sarne biologiOaJ class as Linnea.n spec.iell, or oven elemeut&ry speciei.
t Iamm(lebtecl to.Mr. J. O'Malley. ofOtira, foroaJlingm.y attention to the latter plant.
(JooruYNJlI.-EcologMal Studies i'll Evolution. 11
l~lG. I.-OUTLINE Oll' LEAII' Oil' (1/) RIrBtrs BAUlIIBI, (II) R. PAllVIfil. X 5
Only one pln.nt was originally noted. Recontly I bave seen abundance of
Rubua parvus in various localities in Westland and under difYerl'nt <'Ondi-
tions, but it is remarkably constant in characters, and presented no trrmsi-
ticms towatds R. Bar1ceri. I kuow well that my action in "creating" a
taxonomic species in this case is open to adverse criticism, especially as
I believe that the original wild plant may be the only one in existence; but
if a species can originate by mutation there must be a time when there is
only OM individual, and if so, and its characteristic marks are of " specifio ..
importanoe. it is just as much a t< species" as if there were thousands of
similar individuals.
Red clover (Trifolium pratense L.) and cowgrl:lSS (the var, perenne) vary
to an astonishing extent in a small patch, chiefly self-soWll, in my garden.
Many of the forms are most distinct, and the new characters are diverse.
affecting colour of fiowers, stems, and foliage, fonn of inflorescence, degree
and kind of hairiness, genera] habit, &c. Hert> pure culture-methods and
Mendelian procedure would be needt.'<l to come to any reliable conclusions
as to variants such as these.
HOlell1 lanatus L. and Dactylis glomernta L., I am informed, vary at
times beyond their ordinary fluctuating capacity.
('apsella BUT8a-tpaJltoris Medic., a very variable species in its natural
habitat, and which has alrl"ady given rise to carlall} lllutants, varies to an
astonishing degree in New Zealand, ospecially in highly manured ground.
A ('aroful study of such yariatioll is certainly demanded.
V. EpHARMo:NY.
1. General.
It is when we come to epharmonic adapttitiolls that ecology presents
its most important contribution to the evolution questiOll.
III !\ttemptillf.!, to explain the origin of epharmonic adaptations it is
evident that, as ill the case of all explanations of evolutionary phenomena,
no absolute proof can be given without experiment, and, where such is
wanting, it seems l'eallOnable that the most probable explanation should
be accepted for the timn being, notwithstanding that other though less
pl'Obable explanations would fit the case. Geuerll.lly in polemical dis-
cussions on matters of evolution llatUl'&1 selectioll is assumed to be a vera
causa which ueeds no demonstration, and if any other reason is put for-
ward, however likely it may appear, it is considered of no moment, unless
it can be proved not merely to the hilt, but to the objector's satisfaotion.
Now, I am of opinion that in the hereditary epharmonic variations
cited below there is a much greater likelihood of their having been brought
about by the direct actio)] pf the various ecological factors than by tht'
continuous accumulative selection of fluctuating varieties, and in making
this statement I am merely echoing the opinion regarding a.nalogous phe-
nomena or Romanes (1895, pp. 122-32), Warming (1909, pp. 370-78),
MacDougal (1911, p. 57), HCllSlow (1895, 1908), Costantin (1898), (Scott-
Elliott, 1910), ulld many othor writers 011 evolution.
With the lIlu('h-dispnte-d Lalllardcian factors use and diBUBe, which are
so frequently the on I!! parb:! of tho doctrine dealt with by the zoological
(JpponAllts of modifipd Lalllan·kism, I have nothing to do. How far evolu-
tionary methods ('orl't'Spond ill the plant and animal kingdoms no one can
say, but it dol'S )lot seem unreasonable to imagine that they may have been
ill many respects differont. * At any rate. this pa.per is concerned only wit11
the hotanical sid(' of (·volutioll.
2. F1'Z'if!J 0/ Species-Plasticity.
Nothing has been brought out more clearly by ecological studies in
New Zealand than tho extreme ., plasticity" of many species and struc-
tures, and their rapid response to a change of environment. Thil is 8t)
• LIla,yitt (1907, p. 237) writes, "In no case ill it Rafe to reason deductively from
nne kingdom to tht' other. In the facloTl! affeoting tbPh' 8,·olution, plant« and animaJ.o..
(liffer vAl'tly.
14 Transactions.
great in numerou8 insta'1l.Ce8 that the idea of "normal" 'ose8 tts meaning.
Take the following examples :-
(0.) Leptospermutn acopariun! FOESt. (Myrtac.) lUay be a moderate-sized
tree, a tall shrub, a dwarf plant 2-8 cm. tall which :flowers and ripens seed,
and an absolutely prostrate plant which fonns a dense covering to the ground
and puts forth adventitious roots, although tho erect forms are exceedingly
difficult to a.rtificially strike as cuttings.
(b.) Certuiu shrubs are of tnt> xerophytic awaricating growth-Iorm when
growing ill the open, but of fl compOITati'flely loose, leafy, and meaophytic
habit when growing in the shade and shelter of the forest-e.g., Piitosporum
tlivarioatum* C'ockayne, Oorokia Ootoneaster Raoul, A.'7istotelia lruticosat
Hook. i, &C. In such a case, were the shade form alone in existence (see
Plate VII, fig. 1), there is no botanist but would consider it fixed and
normal, and yet it is the sun and wind form rather that is so considered.
(c.) Fuchsia ('olel&8oi Hook. f. (Onagrac.) is a twiggy shrub in the open,
but in the forest it is frequently a sorambUng liane.
(d.) HymenophyllJum multifi,dum Sw. (Filic.) when occupying wet rocks
in the A.uckland Islands has its fronds closely curled up, but when growing
in the forest interior of the same group they are quite flat. That the curled
fronds are not fixed 1 have shown by means of moist-air culture (1904:,
pp. 266, 267). Suppose that H. mufJ:ifi,dum was only found on B 'wind-
swept treeless island, such J.S Macquarie Island, no one would question
the cutled frond being normal and fixed.
(e.) Myoporum laetum FOESt. t. (Myoporac.) is nearly "lways 1'1 Btnafl
round-headed coastal tree b.avinl!, a distinct erect trunk, but on Moko Hinou
Island it is altogetktfl' 'P'08trate, and its branches far-spreading, cord-like,
and twiggy. Were it not that I have seen intermediate forms on some
parts o! the North Island coast I could hardly believe that the Moko Hinou
plant was not a stable form.
(I.) MyrluB pedtmculata Hook. f. (Myrtac.) is generally either a smalZ
(ree or a twiggy erect snrub,t but at an altitude of some 1,200 m. in the
Notkofagus forest of tho volcanic plateau, North lsland, it is Ireq:uemly
quik prosttOtle and wQt;ng. 8typ'keZw, fascicula"a Diels (1iJpa.crid.), although
nearly always an erect shrub as n forest-plant, behaves exartly as the last-
named in the same station. On dWlOS it is I~SO frequently prostra.te.
(g.) Dracophllilum poZitum Cockayne (Epacrid.) when growing 011 tlu'
mountains of Stewart Islalld is II tllrf-making shrub, a low 8prsaclitlg shrub
with stout horizontal bra.nches, or II mas8illll ball-like oU8hion pla'llt, accord-
ing to (·ircwllllbmces. So difft'Tellt are those various forms that I mn
hardly yet believe them to be merely nnvirollmontal UIlDX('() fOlms of one
.anothor§ nnd that my observation is not faulty.
(.n.) (JIRiakenia clicarpfl R. Br. and G. ~ Sw. (Filic.) diff('r speci-
fically in the former having the lnargins of the segments of the pinnae in-
curved so as to be- pO\1ch-shapod, whereas those of the latter are virtually
:fiat. But the same individual of tbe var. Mcistophylla Hook. i will possess
• Thls plant has been morgt'd with P. rigidlUll! Hook. f. A dlagnor,is hAS not yet
been published, owing to lack of auftioient material, but it is neoeBfl&ty he~ to use &
definite name. since P. rigillv.m and P. divarilltltwn are certainly distinct t'ntities. (See
Pla.te I.l
t For further deta.ils, see Cookayne, 1901, pp. 265-67, and Dials, 1906, pp. 66-611.
t It hi possible that tho tree and shrub are ditferent species, but 1 ha.rd1y think
so, though I have not seen intermediates.
§ See Cookayneo. 1909, p. 16, and photo No. 13, facing p. 17.
('0CKAYND.-Ecological Studies in EI·olutioll. 15
some pinnae wiJ,h poucl&eB and otM:rS quite flat, in accordance with the degree
of illumination to which they are exposed. In fact, here tke 8pecific dis-
tinctiOfJ does not hold-it is merely ephanllonic-and the latest name must
be abandoned; 110r call the two" species" be maintained even 88 " varieties."
(i.) Di8C08'ia townatou Raoul (Rhamnac.) when growing in positions
subjet. .t to tho a.ttacks of rabbits may form low green oushions made up of
leafy spinoless shoots. " Normally" it is a stiD branching 8kl"l0 furnished
with a.bundant spines.
Many more examples could be cited, but the above SMW clearly enough
how unstable species may be, even when growing under natural conditions.
When experimental methods are brought into play the e:ffecOi from plasticity
become still more strikinp:. For example, spine-production may be sup-
pressed in Disoaria toumatou,. true leaves may be produced in the whip-
oord veronicas and species of Oarmiokaelia (Legum.); rolled leaves made-
flat,· and vice versa i cushion plants opened out widely. Undoubtedly
a serios of experiments such as those of Klebs (1908) would yield results
equally surprising.
It can be seen from the above that this uncertainty 88 to .. nOl'mal"
form opens up room for great doubt in all discussions regarding the origin
of permat&ef6t adaptations, for it may quite well be as8erted that absolute
fi,zity does notwt. It seems to me all that can be done is to consider
as to llorma.l" those forms whirh predominate and represent the general
growth-fonn of the bulk of the individuals; but (/,88'118'edly in no lew oases
there i8 no normal/orm oJ all.
:t Response to Ecological Flwtol·S.
W8l'ming has IIwnmed up the state of knowledge on this head up t&
the date of pUblication of his admirable" Oecology of Plants" (Warming.
1909, pp. 16-81), so that only a few local examples are necessary here_
First of all, it lllust be emphatically pointed out that it is virtually
impossible in the field, where so many ecological factors are concerned,
to say whioh is the predisposing cause of the internal response of the-
plant. Generally more than one factor will be concerned.
(a.) Soil. ..
Exoess o£ salt leads to succulence, as in certain salt-meadow species
which become loss succulent as members of non-halophytio formations.
The introdu<'ed Silme angZicfJ L. develops more succulent leaves when
growing neal' tho sea tban inland. Miss Cross examined the anatomy of
certain salt-mt'adow plants and those of the same species grown in oxdinary
soil in a greenhouse. Her figures show considerable di:fferences in thick-
ness of leaves, but other factors besides want of salt doubtless affected
the result (1910, pp. 569-71).
The soil Ileal' hot springs containing excess of sulphur, &c., inhibits
the erect sbrnb form of Leptospermufn ericoides A. Rioh., which then occurs
only in the prostrate form.
Lack of nutritive salts in sand-plains near the mouth of the River Rangi-'
tiUj IUld elsewhere changes the leaf-form of Selliera 'laau:aM Cav. (Gooden).
This is in accordance with the much more carefully conducted observations
'" III the caso of Olearia cYM1:t'foli.n Hook. f. the much revolute, boat.shaped J.ea.n
become flat with mom-air oulturt', and what was considered an important speoifi.
charaoter, di'ltine,ui..hing the" Rpeci8lt " from O. nUI/UII'IIlarijolia Hook. f•• vanishes.
16 1'''(lII~acf/Onl!.
(b.) Llullt.
Tho bright light of dunes proba.bly leads to the red- or orange-Loloul'I'd
steDlil of the rush-like uptocarpu8 8implex A. Ri('h (Restiac.), whiuh 1),1'('
green in the shade, and ItS salt-swamp plants not nearly so brilliantly
coloured. It is a moot point how far the reddish, yellowish, or brownish
hue of certain true dUllt'-pln.nts may be considl.'l'cd fixed and hl'reditary
* Boo on thill Iltl&tl BnnUl, 19] J, Pl1. 121,124. Xerophytos are confined to (!('rtain
zones in the bogs studilKl, tIll' lal'gl!f>t OO~'BI'('Qq lx,illlJ hycll"l,hytil' or mUl-ophyti('.
('OOKAYNJ!..-Ecologlr(l[ 8tllliles III El'ullltltJII 17
(e.g., SOli pUB /l"(lIIdOSUS BlIllkt. & &1., ('opros/lla aenola A. CUUIl., (.Jurmern
rtren.nria Cheesem., Euphorlnn ,,[nUt'n FOl'bt. f.).
An mteretlting cast" is that of Lycopodium ramulosum T. Kirk, a plant
fOlming extt"usive patches on moors ill tht' west of the South Island and
~tewart Island, the sporophylls of whwh are absent 01 scantily produt'ed
In shade plants, but extremely abundant 1ll those growing in bright light.
Many youn~ trees ill the forest aStluIUt' u sperial form with a slender
ma.in stem and few branchos, whirh art" confined to its upper portion.
Similarly, the xerophytie fern Pterid&ulJI esculelltut1l Cockayne becomes in a
dim light a srrambling liane, Au examplt" obst"rved by Mr. H. Carse and
myself was growing a.mongst tall, slender Leptospenllum BCOPMiu»I on Reef
Point, north-wel:!t Auckland. Some of the fronds were more than 3 m. in
It''ngth. Pinnae were II bsent until the brighter light was gained. The final
portion of the rhachis was green Ilolld succulent, and the distance between
the pinnae 46 t'm. Tht'sl' latter were still coiled up and quite rudimt"ntary,
,Illihough tho largest ,vas 25 cm. 10nf2.. The rhnt'his was twisted-i.e., it
bhowed a tendency to tWllle.
I!'lhade-and herf.' proba.bly COUlt'S III mOIsture in the air-increases th('
Rlze of leaves, t'hllllges certaill xerophytes into mel>ophytes: e.g., species
of <.'armichaelia, D,scnri« fuulllntou. PodOt.'llrpU8 nil'alis Hook., as may be
plaillly seen from PInta IV, l't'spouds markedly to changes in illumina.tion,
tht' shadt' form resembling P. tolam much more than the speciel:! which it
really is. Th(' Spt'('inll'ntl were ('ollt.'(·tt'd within a few feet of one anothu.
The lie of the leaf is rt'gulnted by the light. Olf'aria ilISi!ln!B Hook. f.,
a shrub of dry l'ocks ill Marlborough, archt's its brant'hes upwards to a sur-
prising degree, thus bringing its leaf rosettes into a suitable position with
regard to the light. This hahit persists in plants raised from seed and
grown on Jla.t ground.
(0.) WInO.
Wind is a most impol'tant mctor in New Zeala.nd. First comes tht"
" wind-shearing" action, which is m part a physiological process; it is
wt"ll marked in trt'es and shrubs of oxposed positions, and may be frequently
seen in Podocarpus totara D. DOll., Leptospnlnutn scopari'U'In Forst., and
many other plants. The prostrate habit is encouraged by wind; but here
other mdors may enter in, as cold and acid soil. ('opro8'lIIa foetidi.ssimu
Forst. is usuallv a tall forcst-shrub, but wht'n a member of the tussock-
moor Ilssoriatioil of the Aucklnnd Islands (Cockayne, 190<JA, pp. 200, 201.
and 219) it is prostmh' and twiggy. The prostrate form of Leptospenll'Um
scoparium. 011 the subl~lpino moors of Stewart Island is another and remark-
I~ble example. Well-developed prostrate trunks are to be seen ill Ml'tro-
sii/eros luciila Mellz. (Myrta('.) in the Auckland Islands, Stcwal't Island, nnd
thl' Southern Alps, and ill Olearia ilioiloUa Hook. f. (Compos.) in some sub-
.l.lpille foresia of the Bouth Island. Reduction in sizt" of leaves must often
be attTibuted to wind-nction.
(d.) Wat~r.
Plants of still or slowly running water are subjected to a fairly con-
tltl1nt environment. * ('otu.la coroflopijolia L., as a land-plant, is a herb with
branched, prostrate, more or less rooting stems, the branches of which are
erect or semi-erect; the internodes are short; the leaves are rather
B.eshy, more or less lanceolate in outline. and pinnatifid, lobed, toothed, or
sometimes t"ntire; the roots are, at most, of a moderate length. As a
• Of OOUfIIC', the position of tho l)lant with roganl to the bUrf.OI', the nature of tht'
&ubcltra.1um. a.nd othe-r fa.ctoT'l ('Zl'roi'!l' 110 oon'lidera.ble inilUE"nCf'.
18 T /'(1,11 ~tlrtiollio.
water-plant, the stem il:! straight, unbl"lln("ht'd, and pt'rhllps 40 em. long; the
intel'l1odes are lone;; th'" leav('s linear and E.'l1tir(', and the roots numerous
and SO 40 cm. long; wilen the shoot risE.'s abovE.' the water-surfacE.' it
hrallt'hes, nnd the Il·t\V('S ,Ht' much as in the lu.nd-plant.
Not only th(' leuvl'1:! hut also the inflorescence diff(·J' gJ'catly in size In
thl' ltllld and wllt('r fonnl:! of thl" illtl'Odl1l'cd Radi(!'U/a NallfUl1itmr-aqttatioot1l
Brit. & Rcnd.
Specially moist air I'auses the production of aereul l'OotR 011 thl' IItems
of certain whipcord veronicas.
Bc1Ieflkra digitoJ..o, Forst. (Aralial'.), a low forest tree OJ' shrub, when
II,rowiug ill certain damp gullies of northern Auckland produces sometimes
leaves much more deeply cut than the normal.
The moist-gully form and the dry- or aeid-ground form of Blrckntllll i
capeme Schlcht. (Filic.) are so distinct in appearance that many might
('onsidcr them distinct spec·ies.
(e.) Altitude.
Altitude is l~ complex combination of faetors which sometimes product's
striking difierencrs in the same species, according to thl" height at which
the individuals grow.
A very common feature is diminution of stature with increase of alti-
tude, though this is not so with all species. The trecs Dacrydium cupre8-
stntlm Bal., Wl'inmannia racellW8a Linn, f., and Griselinia littoralis Raoul
are much reduced in size when forming a part of the mountain-scrub of
Stewart Island, the two latter eventually becoming small shrubs.
On the other hand, if the lowlands can offer an equivalent environmont
to tha.t of the mountain&-thougb, of course, it can never be actually
identical-a.lpine plants may oceUl' at sea-level, their forms differing not
at aU n'Om those a.t all altitude of 600 m., 900 m., or considerably higher.
The lowland moor of Stewart Island ('ontsins vaIious alpine plants ot
this ('hal'acter-I'.g., Celnm'sia argenlRa T. Kirk, Astelia li'llRariN Hook. f.,
DracophylluJIl p"litum (iockaYllt', Carpha alpina R. Br., Drmatia novae-
zelnntiilU' Hook. f., {falthn. l&ooae-ze/MuHae Hook. f.. (/aimardia ciliafa
Hook. {. (For full list, Sl"e CO('kaytll", 1909. p. 27.)
4• .ljtl'l'-eUl'cl 0/ Stimuli.
It is most imp0l1ant with regard to thl' question of thl' ultinwh' hClreuity
of (·hallAc~ ill form and structul'l". &('., Jll'OUgbt nbout by 11.11 internal l'('-
spouse of thC' plUllt to stimuli from without to iuqlliJ'(' liS to ddbtite
examples where the> form, &;('., persists for a reasonably long time after
thl' stimulus is removed. Thr following cases lwar 011 this sllhjcc·t :-
1. A prostl"ate form of a ~pecies of Oopr08fl'la (Ruhiac.), which originally
grew on acid peat on the Chatham Island tableland. was cultivated by
me in a pot for three years, and then ill ordinary garden-soil ill a garden
for four years more, during the whole of which time tho prostrate habit
remained. But all 011 a sudden, durinp; tho eighth yc-ar, it commenced
to put fOl'th erec..-t shoots, and hut for its unfortunate destruction would
lmdoubtedly by this time have been 011 erect shnlb. So assured was I
that this plant would remaill prostrate or stunted tha.t I published cer-
tain remarks to that effect (1907, p. 878). Bo, too. with a stuntC:'d form
of Q.lloGhcr species of C'OF08flIG, pE'rhaps O. ouneata Hook. f., collected
by me ill 1903 in AntipodE'S Island. This WIlS grown on the rorkery Bt,
()ocruYNE.-Ecolog~cal Studies 1'''' EvofutlOri. 19
Canterbury College for six years and kept its habit, but later on it too com-
menced to put forth erect shoots.
2. Ooprosma Bauen Endl. when growing on a sea-cliff is a straggling
shrub, more or less closely :ll.attened to the rock-surface, and puts forth
nothing but long spreading horizontal shoots. Such plants bear flowers
and fruit. This growth-form of the species may be referred to wind, and
porhaps heat. But when C. Baueri grows in u. coa.stsl forest, or even when
isolated on loamy clay, it is a tree with a stout trunk. Plants whirh I raised
from seed, and whirh a.re now growing in the experiment-ground at Canter-
bury College, possess long spreading horizontal shoots---i.e., they are of
the shrub form, as above; but they are also developing erect shoots, and,
if permitted, they will eventually grow into trees (see Plate II, fig 2). Here
it is possible that the prostrate form is inherited from the rare of rock-
frequenting plants. But the sti.mulus has not been sufficient to make a
really permanent race, and so the prostrate form only occurs during an
early stage in the ontogeny of the individual. Similar cases of partial
heredity are dealt with further on when treating of prolonged juvenile
forms.
S. Oleana Lyallii Hook. f. (Compos.) forms a pure forest on some of
the New Zealand subantarctic islands. A striking feature is the prostrate
or semi-prostrate trunk, which may be referred to wind, a peat soil, and
perhaps a uniform low temperature. In the interior of the forest, no
matter how boisterous is the wind without, it is quite calm, and yet the
seedlings are nearly always more or loss prostrate at first. So, too, with
the seedlings of O. Oo'lensoi Hook. f. when growing on the mountains of
Stewart Island.
4:. The case of Suphora microphylla Ait. and S. prOBtral,a, Buchana.n:
This is fully discussed in this paper under the heading" Persistent Juvenile
Forms" (p. 25), to which it may be well perhaps for the reader to turn
a.nd consider the case in relation to the point under discussion.
It would be beyond the scope of this paper to mention in detail instances
of after-effect of stimu1i in places other than New Zealand, but it is we1l
to Lriell.,Y ~llum.erate ~ few of the more striking. Such a.re Schiibler'B
cereals, which, grown in a northern climate, ripened their seeds earlier even
when cultivated in southern countries; Cioslar's conifers, whose seeds,
collected in the Alps, when sown on the plains produced plants of slow
growth and small diameter j Klebs's Veronica and Semperoilvum, whose
striking a.bnormalities of inflorescence were repeated in plante raised from
seed j Blaringhem's races of maize and barley originating from planm pur-
posely damaged in a specific manner (Blaringhem, 1907) j Zederbauer's
experience with a form of OapBella Bwrsa-pastoriB from an altitude of
2,000-2,400 m. in Asia Minor, which through four generations in Vienna
maintained in part the special alpine stamp; and MacDougal's ovarial
treatments, where one new induced form has maintained its character, so
far, up to the fifth generation (see MacDougal. 1911, pp. 56, 57).
5. OO'llllJef'f]etR Epharmony.
From. what has gone before, it is plain that ,-arious growth-forms of
New Zealand plants may be referred with confidence to stimuli from outer
£actors. It has been seen also that of such forms SOlUe are merely environ-
mental; but there are others,' now to be dealt with. which are hereditary,
and remain constant, unless perhaps when exposed to such a change of
conditions as they would not encounter in nature.
20 Tr(m~actlolU
Every tra.nai.tion oxist.e between the open circuLu mat-like form sud
dense Ullyielding eushions. It is nteTely a question of degree in redu('-
tion of internodes and closeness of growth. The genus Ot'ZmisiG shows
... M. Aalcmi p(otrit', mo&t o1olo<'ly related to thl' Il&nl'. 1J/. romplsru, i. a dl...",rioating
phnlb.
TRANI. N:if, IN ...' , VOl XLIV PL\'!r III
• I:!ee '&Iso figs. 229, 230, Goebel, 19().), pp. 333, lIM.
o.
t qtUJd~ Col. is OJDltted, .., It seems to me merely a. va.ru.nt of O. tllAnUa
J. B. AnnstronJr:. Nor do 1 know a.nytlll~ roga.rdWg ~ V&1'I. d8fllJ"fIt"'Itll Hook. f.
and jrlio6cUtJ Klrk of O. 'fJfI7'fIiflm'a .A.. CuIUI.
'I'II\N' N ~ I", I \'11 \I.IV PLATL V.
.I.
II"1Q !l
L. VL:BONlUA LOGANIOIllCb 2. V. C.AbbIN,OlDll&. J ltL\ LBbIOIOo SUOO'r 01 V lLlll.AGONA.
4.. V TE'l'l!..o\GlONA X 3
Ji'au p II.J
TuAN... N Z IN-.'l, VOL XI,]V
'" 1 ..m llIJ)lined to tlunk it would be better to (lol1ll.lder tluK .. ~l'0(lIefo,. It di1ferl,
con"iderably from the typioa.l form. which grows in the Ea.1lt Cape distnct.
t HN.erobla.,tio development is a. world-wide phenomenon whioh !wi not received
nearly the atteIltion it deeerves from writerK on evolution. It is its OOC!urrlmCC' in RO
many ondemio speoies in New Zealand tha.t:ma.kee data from. tlwI rtogion of Iopecial iuterest.
In 1879 I. Bayley Balfour roc01'ded a. number of striking examplell from the Ibland of
Rodriquez-..6.f/., Olerodsftdron laci"iaeum BaJf. f., :reminding one of the N'bW Zealand
YolMpa'IIQII; Bt''lll'plefl: Seem.; PyroBWia tn'loculan8 Balf. f.; l'emelll1 ~tolla Lam., a
l'Ubiooeouli plant, evidently wheli juvenile somewhat of the dival'ica.ting t.hrub form;
a.nd .JlaawriM ~fI,oreI Balf. f. (Tumerao.). which haIIIQ!lg narrow juvenile and
broad adult loaves, as in J'M8I1I'I.8ia. 'lMterop'/ltJ'O.a A. Cunn. and othl'r New Zealand plants.
AltogothOl ISOvonteen. species of t:recs and 'lbrubs and one herb out of 170 Sl'eoies of
~permophyt.oP ahow marked. dimorphism. GOl'bel (1889-93) gives a number of t'Dmple&
of hN.erophylly, &0., referring the phenomena. in some insta.noes to direot outl'r stimuli,
a.nd he deals further with tlie matter in hiR "Organography of Pia.ntb" (1900-5) and
his .. Exporimentelle Morphologie It (1908). Diels (1906) goes into the IIl&tter at con-
sideAble 1ensrth. ueing many important illUlitrations from his ob&ervatlonB in We&tern
Australia. As for thti phenOmenon in Ne~ 1.caWld. Hooker WI.9 the first to refer to it,
m hla splondid .. Introductory Elsay to the New Zealand Flora " (1853, p.l). Kirk giVOEo
many cfetai.ll! in his "ForNI; Flora" (1889), and these are E.uppleJllentcd by Clheeot'man
In bill lfanual. FinaJIy, my own writing<! '\lure 1890 acmtain a. goon deal of 'lCatteJ'('(l
Information not previously publiehed.
Tr(f'nlutIfJII~.
form may persist, or there may be lIo combination of charucwrs, 1\1"1 ill tht'
transitiona.l forms. In any case, hl"Tedity comel:! in, and thiR hns uttuiUOll
to such a degree thu.t under normal conditions there is a juv('nilt' IItngr of a
('ertain average duration, a transitional stage, and an adult.. Diffor('nt
degrees of heredity have arisen, as I believe, in proportion to the lengtb
of time the original stimuli have functioned, combined with their intensity,
and abnormal increase or decrease of stimulus can in many iustances
hasten or retard the procession of events. There is in some measure, per-
haps, species-making going on before our eyes. This is best seen in those
cases wher~ the juvenile form. produces flowers, for if progressive develop-
ment should cease at this point what is virtually a species distinct from
('OC'KAY:IIJI..-J<J('(J109lr(/1 Sfllrlll ~ III J<J !'Ol,iflOll 20
the ooult has appe8rt'd, Should such II flowering juverule form be eph'il-
monk then, 8S Diels has shown, we are face to face with a case of onto-
genetic evolution (1906). In somp of the species the juvenile and adult
fonns Cdn both clearly be shown to be epharmonic (e.g., Veronica l!loopo-
dwides Hook. f., Oa,miollaelia 8ubulata T. Kirk, Discaria toumatou Raoul,
Potamogeton Ohee8elnMlii A. Bennett, Clematis a/oliata Buchanan); they
lim even be experimentally produced or prolonged. In other cases ephar-
mOllY can only be inferred (Sophora mi(Jfophy17a, Podocarpu8 dac'f!ldioidelJ,
Rubus schmidelioidea); alld ill othE'rs it is more or 11'88 obscure (Pa/'8(lnBla
heJerophylla, PBeudo]Jf1lflA1a cra88i/()1iu'R C. Koch, PittoaPO'f'IIAR patulum
Hook. f.). There is, thereforo, a gradual gl'Sdation from the known to the
unknown, but, as the main features are alike throughout, it is reasonable
to assume an epharmonic origin in most ('ascs, notwithstanding that con-
tradictory examples occur, and to consider that there is n relatiou between
the age of the form and lts relative stability. Here there is no attempt
to go thoroughly into the phenomenon under cousidcration; cel'tain typical
examples are alone discussed.
The significance of the divar;cating growth-form has been ah'eady noted.
It ma.y bt' remembered it is emillently xerophytic, extremely well dt'fined,
,md preSt'nt in various unrelatt'd families. But this form is not confined
to shrubs alone, but appears as a persistent juvenile stage in the life-histolj
of certaiu plants, which aro thus xerophytic shrubs for some years sud
finslly ordinary mesophytic forest-trees. The following are examples:
Pffl.tlantia oorymbosa Forst. (Icscinac.), Hoheria anguBtl10lia Raoul, PlaYI-
antkus betulifuu A. Ounn. (Malv&c.), Suphora mirrophlllln Ait. (Lel!,um.),
ElaeoCOlpus Hooke'fianus Raoul (Elat'ocorp.).
The csse of Sophora miorophyUa Ait. is the most instructive. It must
be considelC.'d along with the remaining species-So tetraptera* J. Mill.,
8. granrliflora Salisb., and S. preJBtrata Buchanan. All the species com-
men~" with hypogcal ('otyledons, and th" first, or first two, leayel:\ are
simple "nd arrested structures, but the RUI'cerding ones are pilluute clnd
of the adult type. The primary stem is pre(:t and sompwhat :flexuous (set'
Plate VI, fig. 2), except in the csst' of S. fl1(I1Uliflora. This speciel!
('()ntinuOB to grow erect, and in time devt'lops into 8 small tree. T.I!erc
is no heterophylly beyond the early simple leaves, and no hint eYen of
any xerophytic shrub stage. With S. tIIicrophyUa the progress of events
is very different. Here the early seedling 80011 develops into a xt'rophytic
divaricating shrub, and so it will remaiu for somt' ten years or more, and
attain a height of perhaps 1·4 m. before the more 01' less erect branches
shoot upwards, the forerunners of the mesophytic treP form (see Plate nI,
fig. 1). It is quite common to see a specimen which is shrub at the
base and tree above. Occasionally the upper part of the shrub fonn will
hlos80m. but I do not think this evPl' happens before the tree itsel£
fiowel'S. SophO'1a pr_ata neVer grows out of the shrub state; it is a (izfd
j!WffIt·lt form, whioh, moreover, rt'produ('es itself true from seed. Betwpen
B. tnicrophylla, and S. prastmta there lire illten11Miates. With regard
to S. tetraptera, the juvenile plant differs but little from the adult (see'
Plate VI, fig. 1), though it has for s time a few flexuous twigs. I have
• Under this 11&DlO [include thu llhatham IAland plant, 01. form in the neighbour-
hood of Auokland City. and the Chilian plant. As for the- Auokland pla.nt, I do not
know it. juvenile &ta.te well enough to speak with certainty. but in any case its behaviour,
jf diBeft'llt from that stated, woUld not in any way aft'eot my ~OIll'bl6iolls.
1'r(1II ~(/rf/()1I ~
80(,ll only Onll speCImen r<lised from Chili un seed,· and it rcsemble& rlo&ely
the Chatham Island plant. S. micl"Ophylla and S. prostrata grow sidt' hy
Sidt' itt the lo"er Waimak.mri Gorge, Canterhury Plain.
In the abo,,(' case of Sophora the adult form is probably the stem form,
.\Ild the xerophytic dlv.ulca.ting shrub form an epharmonic adaptatiol
whi('h uw':>(, during a probable period of drought on the east of the Routhertl
Alps at th(' time> of the glacl&l period (see Diels, 1896, and Cockayne, 19(0).
In tertain parts ot the problematical Greater New Zealand where tht'
ctimdt(' still relTldomed sufficiently wet the ancestral Sophora would ft'-
main unchanged; 80 we still see S. grrmrUflora in the East Cape distrirt
and S. tetrapt"a in tht" Auckland distrirt and the Chatham Islandb. III
the South Island there is only S. mA,crophylla and S. p1OBtrata, in the fonnel
of which the xerophytir stimulus never evoked an absolutely hereditary
form, whereo.s in the latter the effect of the stim.ulus is much more deep-
seated. To what extent such a stimulus can leave its mark is shown in
the forest-tree Elaenoarpus HookerialwuB, which at any age may put forth
reversion shoots high up the trunk or on the branches. The hetcromorphy
in the other species listed above may be similarly explained. There i~
first of a.ll a short-lived erect mesophytic stage, then a long-persisting
xerophytic stage, and a final adult me80phytic stage. Tht' first stap;e,
suitl'd illS it is to shelter by ground-plants, &c., is epharmonic j it may
also be coneidHed. a surVIval from the a.neestral plant. The second (XE.'l0-
phvtic) stage Wlt~ epharmonie during the steppe-clima.te period of thl'
eusWorn I:!outh Island, but is certainly benefioial no longer it and the adult
sta.ltE.' is morl' or lees a. return to the original form, but now ('aIled fortll hy
the pl"t.'Sl'nt mesophytlc ronditions. A('cording to this supposition, it i~
considered that the tendency to hoth xerophytie and me80phyti(' form
is latent in the plllnt, and that onf' or the other will appear 1111 1:10011 ,1M
1.11(' necossary intpnsity of stimulus is rl'llChed, Until that iR th(' ('aM',
whiohever form is thl' more heTl'ditury-i.e., the moT<' strolll(ly fiXt'd-
Will pl'rsi'it, (,Vl'lI though it is far from beinA epharmoni(·.
In ,I COllllidl'rab]e number of illstallOrs there is a lll(,llOphyti(' jU\'enill'
st.L£!,l' an<1 a xerophytir adult. III this 1'111.811 the prosont m<'Sophytic ('on-
ditions lire not lIufficil'ut to inhibit tht! strongly heroditary xllrophytir
form, which also in II. number of cases is in ha.rmony with the xerophyti('
stations ..fft'ctod by these plants. ThE.' following examples of this and othE.'f
p<'rllistent juvenile forms may be noted.:-
(1.) Shrubs which a.re leafy as juveniles, but leafless &8 adults, whell
they have B.a.t or terete green assimilating stems - e.g., specie'S of Oar-
miohal'lia,.~ Notospartium, and OoralZoB'partium. How unstablE.' really is
the xerophily of many species of Oarmiohaelia is shown by their abundant
production of leaves in shady stations.
(2.) Shrubs with an abundance of leaves, sometimes very thin, when
juvenile, but of the cupressoid form when adult-e,g., certain TaztJCl'fJE
(see Griffen, 1908), whipcord veronicas, a.nd some speoies of Helic1t,!/,um
b('lol~in~ to the section Ozothafllllit',
------------------ -
• 'l'ho IIeed was very kindly Bent to me by Dr. Eug. Autran, of Buenoll AyL'tlll, anll
tho seedlings were raised by Mr. T. W• .Ada.m.'I, to whom I UI greatly indebted.
t The divarioatinJr form of ~U8 BooirilntIB and the jUvenile PRtV40pa_
craNi/olt·ufII. with its thiok, narrow. BtHf, defleud leaves, oertaInly seem out of plaoe in
.. rain forest, where they &lI) assuredly IIDt epbArmonio struotures.
~ OtmJlic1aGelilJ gracili8 J. B. .Arm~rong i8 leafy in the adult; it is a aorambling
_ . and groWl in wet gtouud or swamps. O. grandlftom Hook. f. is deoiduOWl, but
8burula;ntlyleafy in spring a.nd Bummer. O. odortIfa Co1enso is also leafy.
PLAn VB
a.' .,
FIG. 2.-PCNN\NTTA. C'OllYlTllOM.
On left, adult; on uqht. "'Jm-ju~"ml€' In bluom
'l'R\N'I N 'I '1\"', \'0' \ 1.1" I'r\1I "'"
COOKAYNE.-Brologll'lll Sf11dlf'8 III EwllltlOlI 27
Tht.' juvcmlr stagE' in these plants, the Tazaccae excepted, does not
usually persist for any 101lg ~riod, .md may be compa.red to the first st.lge
III Sophllra and thl:' various diva.rlCatillg shrubs. But ill the verowcas, as
J IlllVt' hhown, it can b<.' made to P<'l"Bist artifit'ially for years, so long as
th(.' plont is kf'pt in moist air. Fllriher, reverSlOll shoots all' frt.'quent
('V(,ll on llul:'b a typical xerophyt<' ell> Hel1tiluY8um Belago, wlule it seems
probablc· that hereditary setnl-juvl:'mle races occur of Veronica fetl"aqo'1w:t.
Hook. .Illd V. 19copoiiAoides* Hook. 1. ThC'se are further dealt mth on
p. 45. In the C&ll' of Dacrydtttm lnxifoliUffl Hook. f., a pro&trate or
Buberl:'ct mountl.lm-shrub, gro,ving in wet ground or bogs, it is qwte com-
Dlcm to find juveUlle plants With the lax spreading lea.ves bloODllDg freely.
IIl1d nev!'r dovoloping into the cupressOld adult. The juvenile I>tage of
D. mtermedi'l1»1 T. Kirk frequently bpcome>s a tree, c\nd fiowels and frwts
nK ahundantly ciS the" normal" lldult growing ill the same swamp forest.
This flowering juvenile WIIS giv<'l\ thl' val'ietal name graetl?8 by Kirk.
(3.) N!'arly 1111 the clivaric'ating shrubs have a pnmaly juvelllie mebO-
phyti(' staAt'. This is gonerally but transitory. but I have already shown
ill the> ('ase (If Pitt08po1"t14f1 dioorioatum and Corokia Ootoneaster how the>
early sto.~e may persist ill the for<'st and reach its full stature. Rt'mi-
juv(,llllf' phmts of the Pitt08porum may also ftOWt'l •
.lriRWft'lta /rofioll8a Hook. f. (Elneo('arp.) is an mtercsting (,8se. The
edrly st'('dhng is eT<'ct, mesophytlc, a.nd, ('ompared with the adult, shows
a most remarkable variety of leaves. These are oftl'n more or less lanceo-
lnt('. toothed, lobed. or pinnati:fid (see Pla.te VII, fig. 1). La.ter on the>
c\ivaricating form Ilppears, which may finally be of the most intenl>e cha-
racte>r, the small frequently more or less oblong leaves being scanty. dnd
the> nltiruatl' sllOots almost spinous.t But this form is not truly stable.
plants J!,rowinp, ill an adjac<'Dt N othofagus forest being much. more meso-
phytil" Even whell quite in the open, thl're a.re forms still divaricating
to 8Om(' exWnt, it is tl'Ue, but juve>nile 80 £."Ir as leaf-form goes, and these
develop no furthor, and blossom. This semi-juvenile :fixed form should
hI' l'onsidt'rl'd olde>r than the :' norma.l" adult, olld it may represent the
lIT('-gluu",l plan1,.
Ntlttcmill IliVfm'cata Hook. t., 110 fdr .tI:! I have observed, has 110 set'dling
llwSOphytil' I:Itugt'. But c'v!'n thIS .. wl'lI-fix('d" spe(·jcs when p,rowing Oil
1,hc' POOl' Kuigh1B Mands hi,s l(,lIvf's thr('l' tim('s the size of those of the
utltll.ll stations.
(4.) Notllll'jlQMJl simple:r. 8I:'(,nl. 11Iltl N. Bclgerleyi, Harms. (Araluu:.) hove
.1111() I~ tn!'MOphytic' juvcni](' form, hut. tb(.' adult must be considered me>DO-
pllytic· Ukewiso. Both arl' rain-fof('st plants, while the fOrnlt'l" is found
J1KO ill certain subalpine s(.'rull. I have not full details regarding N. Edgf't'-
1('gi, the juvenile f(lrm of whil'h sometim<'s so closely resembles tb.a.t ot
N. H;mp7n tllat I, for one, l'l!.1Inot distinguish between them, so my remarks
lire> l'olmut'd to the latter species. The ea.rly stage has a fern-like. much.-
(ut, thin and largt' ll'af. This is sUl'Ceeded by a st'cond stage with ternate
lell.Vl't!, aud this by the Sllllpll'- I:llId thicker-lellvl'Ci IIdult, III SOIll(, IOl'dlitiel>
the mU('h-c'ut form it! &upprebt!rd to some extent, or almost rnhrely IIhtlellt
(Auckland Islaudt!; hut Sl.'(, Cockuyne, L904, p, 249,'" and pI. 11), Thl'
(,Iosely r('lated NothopotlflJ: part'UI/I Cocku.yne ILI&o s('rms to lnck a cut-
ll'lwcd tlt.age, N, anom(llmll &t'IIl.. tLlthough f!'equently IL f()fl·tlt-pluut,
ha.s '"' juvPllile ml'80phytil' form wi1h small terlULie Il'IIV1'S lind an ILd"lt
divltricutillg &hl'ub (mm l'olllw(·ting tht' t('rnllh'-It'lwed Coml of tht· ~1'IlUIi
with tht' divari('utiug shmbs,
(5.) In this I'last> <:onll' d l'ollblderll.ulc numul'!' of pillnts whit'h ('311110t
With A.uy confiden~ b(' referred in their different stages to &pl'l'iul OutCI'
factors, Take tho case of certain speoies of Pseudopam(JX (Arllli!lC',): two
(P, rrtllJstjolium C, KOell and P. jl"fOO: T. Kirk) have the I'urious 111"'1'0'"
deflexro juvenile leaves and unbranched stem, but ill p, N'fIRaFi' <.', Kodl,
d tlubalpine shrub, the virtually similar juvcuile leaves flrc ('reet; and ill
P. rhntll11nniC'lHII T, Kirk they ,trt' wanting .Iltogl'ther, tho juvenile I\nd
Adult Ie-aves not bl'in!!, very different.
The primury seedling ieaves of P. crassi/olwlII t~re somewhat simihH ill
form to the adult, hut, of COU1'8e, much smaller. Th('y are cred, und neVl'r
de:B.exed, p, /f'I'OX, 011 th(· contrary. comlll(,Il('eS with uMrow-liut'ur toothed
leav(>S of the st'l'ond btage, which nre not ~rect, hut horizontal for a tim(',
The smllll-l('uved juvt'nile und the iarge-I('ILVed udllit forms of the l'Oot-
climbing fel'n Blrclmwn fili,jorme Ettill~h, cannot be llxplainNl t'phar-
monically, thoUAh there probably is, or has been, l:Iome relation of th(· sor1,
SillCC the first-na.med is the commou ground form (ereeping fOl'm) a.ud tIlt'
large-leaved the climbing form, Nor can I suggest any explanation of th('
two juvenile led-forms of Parsonsia keterophylla and p, capB'Ularis R. Br.
(see fig. 2). In the former species the long nall'ow-Ieaved shoottl O('('osiOll-
ally flower. aUld ill the latter there is a fixed flowering juvenile raCl' oeclll-
ring in thc uplands of the South Island whirh I ronsider a. distinct specictl,
WeittmlmrWI rn.eem()fl(l L. f. and W, syZvicola Sol. (Ounonial'.) nr(' two
closely rl'lutE'd tlpec'il's who!!t· flowers IUl' virtually identical, lind whirh
diffel' mOl'('I,\" in thl' IItll1lt lell f of the fil'st-llamE'd being entire nnd I)f thl'
Qthl'I' l'OmpOlll1cl. 'rht' ('lU'ly 1:I('(.'dlings of both 1\1't' idontic!ll; tht')' art'
I.'r('rt, tlll'ir 1t'llvPs nrl' Kimpl(', toothlld, thill, anll hairy, Thon ('onu'S IL KOI'oud
stllgE', in whi('h ill lV, rnfflnOR(I the loavos IU't' terlllltl', Itnd ill W, 111I11Ii('olll
both h'l'Ilntl' 01' pilllllLk At this stuge, when hoth plants I~r(' ru(·rl·ly hushy
shrubs. th('y ('1111 tlO\VI'r, Im<l IlPpfl not dl'volop into t,f(l(>S, Fl'Pqul'utly Ull
thn h('oth lunchl of nIH,tht'I'1l Auckh\lld W. 8uwicI)l(l I\ttuins 3--~ m, in hl'ight ;
the lcavt's 1Ll'(' ItLl'gl', lind mwo many lelloflets, yl'llowilllt in ('oll)ur, allli
although MI', H, Oartll', myself, lind otlwrs hllVl' se~n hundrt'dll of thl'II(' tall
juvenilo pllluts WI' htlVe l1ev~r RI't'll them in flOWPl', Aokl'II!I' TOllaej(llifl
A, (\um. (Cullon.), if not Iletunlly a eompa.nion pla.nt, grows 11<'1101' hy 011
the forest's outskirts, &(' •• 1\11d its adult form so much l'etlembles this juv('nill'
Wf'inmMlMU that no Ollt' could distinguish flowerless examples Olll' from
tho Otht'l' without a kllowledgl' of certain quite obscure difierono6s. t The
adults of the two species of Wemmannia are lofty forest-trees, From tho
above it seE'ms l'eaSOllable to conclude that W. syl9Jioola is merely a fix<!'d
• 'L'hl'Ough a oLorioal elTOr ·'l.'lltiI.'U-It!&wd " i"l)rinh.'d lIl.·vcrsJ. time'> instoocl uf "~implc
leavetl." The leaves are mol'E' or 11..,.. HC.'rra.tu, hut compal'N1 with the juv('lIil.l· till'\' 110m
vil'tua.Ily .. ontire," ,
t The IliH1.lnotion~ giVl'1l by Kirk ill t hI' .. I''ol'll'-t IrlUl'8," p. 11:1, (Iu nut hulel in
pra.ctlQf', lID fllr a~ tho lo&f I.. oonroml.'ll,
('OUK_\YNE.-Kro[(){/I('(71 Siud/I'li lit }if 1·lIllltWII. 29
juvenile stage of W. racemOSIJ, or elst' that the former is tht' litem form and
W. 7acemosa a mutation 0)' an epharmonic variant that hali become fixl'd.
Several instances of juvenile blossoming have already been Il.ivt'n. The
followinp, are additional examples :-Ranuncu'lus Lyallii Hook. f. (tht! juvt'-
nilE' has a reniform leaf lind thc adult 8 peltate; reversion 1£'llves occur
.1& a result of hIId nutrition; tht'rE' arE' intermediates between thl' two
typl'S of leaves): P1'ttosporum tmui/ol!um Banks & Sol. (the juvenile
seems to me identi('sl with P. niljrescetl.S Hort.,. thl' plant so much used in
certain parts of New Zealand for hedges; as II hedge-plant the juyenile
form is alone to be tll'en, it being preserved by the constant cuttingt):
('lematis indivisa Willd.: Drooophyllum arboreltm Cockayne: A.gathis aus-
tralis Salisb.: Nothopan,(J!J; Eilgerleyi Harms. (one semi-juvenile form blooms
and is the var. serratwlt! T. Kirk): and AnisotomP fili/olia C'ockayne und
Laing. There are also It number of forest-trees which remam in the shrub
I>tage and flOWl'l' (see Co~kayue, 1908, p. 22).
Each of thl' alJovl' ca,ses would need dedding Oll Its merits 8S to whether
the flowering juvenile might be the beginning of a new line of descent, or
was merely", l't'Vt'l'SiOll. I will only dis('uss the casE' of .-l.»i8oto9lle fiU/olia
Co('lrnyue and Laing.
This is an herh Wlth the Il'aves in all ereet rosette Itnd Ii long tap-root
whi('h grows UPOII StOllY debris where there is a stl'ppe ('limate in the moun-
tains of Nelson. Marlbol'ough, .Uld C3nterbuI'Y. The lpaves are grassy,
some 20 ('Ill. 101lg, ternately divided into segments whil'h lue filiform if
the plant gl'OWl! in thE' open, but 3 mm. broad, or blOader, wIlen growing
in the shadl'. Buth forll1tl produce flowers. Seedlings raised from thl' fih-
form XE'rophytic form had ul'oad segments (see fig. 38, pl. 12, in Cockayne,
1900. and also pp. 295-97). The broad leaves are certainly beneficial for
pl'omotin~ rapid growth ill a. dry station, nor will the seedling be exposed
to as l'igorous lIuI'l'oundi1l#ls as the adult, protected 8S it will be by the
stont's. Its form is therefore epharmonic. The broad-leaved adult of
the shade is th.·ll u. Howl'ring juvenile, whi('h mayor may not be" fixed."
but, if md, it would be an example of ontogenetic evolution, the arrival
of til!' new speeies dating from the first timE' the juvenile plant reproduced
its lik£' {rom sl·ed.
MallY of these ltt'tl'roblastic !:Ipl'cies put fOl'th when adult typiral juvenile
or tlemi-juvenile shoots, as the (,l1.l:Ie may be. ~uch ma.y often 1)1' traced to
1\ specilll stimulus. Thus, IItRms of Pllylloc/nrTus al'P'inu8 Hook. f. when
prostrate 011 wt't lIoil may boar abundance of true leaves, but those in a
drier position have phylloclades only: Discaria toumaiou Raoul cropped
by rabbits produ('l's leafy shoottl only;; and RnnuntJul'UB LY(Jllii Hook. f.
I(roWll in dry soil undl'r unfavourable conditions mar de,·elop a certain
number of reniform seedling leavE'S.
The position of the reversion shoots UpOll the plant differs ill different
specil's, Very often they are confined to near the base, in which case they
may be merely developed resting buds. Pseudopana:JJ crMsilolium C. Koch
and Weinmannia raoem08(J L. f. when {'ut to the ground regenerate from
the stump by means of juvenile shoots, Pitto~1Um tenuiloUum Banks &;
Sol" as a hedge-plant, remains permanently jUYenile through frequent
YI. HYBRIDIZATION.
Handly anything is known as to the o('(:urrence of wild hybrids ill New
Zealand. But field observations 011 this head are, in allY ('uSP, merely
l:Iuggestive, lind, at most. pave the way for experiment.
Long ago hybrids were raised in (.'ultivation by Mr. Anderson Henry
IIolld others in Great Britain from some of the large-leaved lowland species
of VnoHica. What I take to be hybrids-one especially from V. pitM-
looidE's Hook. f.-have originated spontaneously in the eemi-wild collection
of indigenous plants in the Christchurch Domain. Mr. D. L. PoppelweU has
sent mo a. hybrid from his garden which he considers V. salid/olia x V. de-
cumbt'ns. It is somewhat of the salici/olia. type, but with small glossy leaves i
I have not seen the flowers. Recently Mr. A. Lindsay, of .Edinburgh,
has raised one or two hybrids of which the parents art' known. The
COCKAYNE.-Ecological Studies in E~'ol'Ution. 31
most important of these is V. Hectori Hook. f. (a .. whipcord "eronica")
x V. pimeleoides Hook. f. (a small glaucous-leaved straggling rock-plant with
blue flowers), and the result is a plant said to be identical with or
near to V. epacridea Hook. f. If this is true, it opens up much suspiciolL
as to tho validity of many species of the genus ill New Zealand, and, at
any rate, in the case of variation in general. as some of the species art'
gynodioecious,* hybridism may be the simple explanation.
Mr. McIntyre, who had cha.rge of the famous collection of New Zealand
plants of the late Mr. H. J. Matthews, raised a good many hybrid forms
of Oelmisia, all of which appeared to have the so-called C. t'PrbaBci/oliat as
one of the parents. I have seen a Oelmisia on Jack's Pass which was most
likely a hybrid between O. spectabilis and O. coriacea. Also, C. tHollis
Cockayne is possibly of hybrid origin, with O. speclabilis as u parent. In
short, hybridization may account for some of the \'ariatioll in Oelmisia.
Acaena, again, is a very variable genus, which suggests h)·bridization.
Buchanan was the first to call attention to this matter, and he described
a supposed hybrid between A. Sanyuisorbrt.e Vahl. and the introduced
A. ()vma A. Ounn. (1871, p. 208). Kirk reduced this to yar. ambigua of
A. avina, notwithstanding that the inflorescence is altogether different from
that of that species. Bitter (1911, pp. 297-321) describes fifteen hybrid
forms of Aooena, illustra.ted by figures of leaves, ill which yarieties of
A.. Sanguisorbae, A. micropkylla, and A. u1nhra arc pa.rents, one or the other.
These forms have originated spontaneously in the Bremen and other Con-
tinental botanica.l gardens. Bitter is convinced they are true hybrids, and
that the only question that can be raised is itS to the pllorelltagl' that he
suggests for them. A lull account is given of each form.
I have seen, judging from the capsule, what appear to hl' wild
hybrids between Phormium tenaa: Forst. and P. Cookianum Le Jolia.
A good denl of the variation in P. tenm; may be due to hybrid ele-
mentary species, for that it is madl' up of many such entities seems very
probable.;
Me'h'cope Mantellit' Buchanan is supposed by some to he a hybrid
between M. simplex A. Cunn. and M. ternata Forst. (see Kirk, 1889, p. U8).
I have proved that it comes true from seed, and ill the absence of experi-
ml'ntal evidl'nce it is quite as reasonable to suppose it is an ell'mentary
species connecting M. sitnplez and M. ternata. All thr~ haye ternate
juvenile leaves; M. temata remains at this stngl' but with much larger adult
ll'aves, M. Jll]fln,tpllii has both simple and tenlatl' leaves ill the adult, and
M. 8'ltnplerr is it dh'aricating shrub v,-hell ",dult \\;th simpll' Il.'1wes.
at! to tht' lik('lihood of I'xtremely lI'naIl'" vcl.riatlOns being presel'ved IIr tlU'
l'ontrnry, It must 11(' undcrstood that tIlt., " struggle" is not only betwl't'u
thl' individuals of tlll' ('ompeting sprC'i('s, hut also betwl'C'1l these and their
C'l1vironmcnt. This \\IIM c1illtinctly stntrd by Darwin, who rC'{ers to the
'1h'ul!,gil' for life ll~ainAt the <1ronp,ht 011 the ('dge of a dE's('rt (1899, p. 46),
]n mUllY lIIAt.mC·I'M thil! tlh'uggl<' with outer l'irc'U1l1stnncl's IS the morE' im-
porhmt; it is nlao tht' 11I'<'iding fal'tor liS to what pIn nt-form ('Oil P;llill II
Jooting in thC' first instllnl'<',
The iormntions tllCn18t'lvcs offt'T vm:iOllS ('Ondltions Ill'col-dmg ItI! th<'y
.Ill' "opcn " or " clos('d," for in thE' fOl'1Ut'1' thert' is apparently room for
new-comers, whert'a8 in the latter it is almost impossihle for a IIp~lJics
from without to gain admittance. This fact is of major importance,
fOI', amonl"rst other mlltters, it has a stl'ong bearing 011 the muoh-d<'hatt'd
question l'egal'ding former Id.lld C'ollntl<'tions with distant islands 3S op-
posed to bird ('81't'1I1ge, &c" across '0idet nreas of ocean. The case ot
Nc'w ZE'aialld as 8 whole is of ~reat intt'l'est in this regal'd, espel'ially
liS many misst8tl'nll'lltl!t havc ('rept into evolutionary writings regarding
the spread of th(' intl'udw'C'd plants Itnd their rapid .. rep7aCI'tIU'llt" of
thl' indigenous fiord. I will Statl' hl'lPfiy what I llC.'IH'Vl' to hC' th" tl'ue
stllte of ttf£mrs.
There hllvc lIl'ell rt'C'orded fol' New Zealaud up to till' P1'('Sl?llt &oml' 555
species of lUt1'odlll'l:'d plantll, but less than 1M can bl' c'onsidl:'red l'OJnmOIl.
whilst oth('l'S arc lo('al, ral'l', 01' (,V(,ll uot tl'1lly established .IS Wild pllln11-1.
Lny at first sight appear better suited to the soil and rlimllh' th<tn are tht'
indigenous sp('cies, and oyer mu('h of thc laud they giw thC' clul.rl1.('(eristi('
stamp to the vegetation; but thi8 i8 only the ca8P where draining, C'u7twation,
ooMtant burning oj fore8t, scrub, and tU,8800X', and the (/mzi"'!l oj n lIIultitude
of dome8tic anima78 "atV' lIIadr absnlulRly 9Ie'IO edap}lIc rotltlitiO'll1l lllhirh a p-
proximafe tn tnnse oj Rump", and whl're it is no wonder that th(' RUI'ol"'Ull
• DUI'illg tht, diM'II"'IOII fnlJllwing thl' l't'dolling ot tlll~ pa.pel' till' ~XPI,(,""'lIl11 .. ('x-
h'C.IJlll'ly 'mall .. wa" crilioilll.'(l lUI not giviult n fBir l't'I'l'{"'<'ntaholl of till' viow~ (If I)nrwill
lind hi~ fllllo\'II'r~, I>al'Wiu, howov~r, wrihoq (ltI89}-1'. 45, .. Val'iatioll", how('wr hlil{ht ";
11. Gil, .. allY Ildv.minp,tl, howI'vcr slight"; p. :10, "cxtl't'm('ly Kliqht modifil'llotioll"";
p, fill, .. individllaJ. llificl'Illl(,I"M, too oJigbt to Ill' IIPI'l't'Oiailod by u~." WC'oonnllll IIlltK
thl' ('I\8C mill'!.' htrulI.~ly Htill (11110, I" 2,i): "Fol' thl' CIUllHtiull iN lIot ml.'roly wll('tiwl'
filli,lll'd IIli11Iltn1ioll' haw fI(')I'Otilln va hI(', but ~hl'the'l th.(' lI"'t IIC'uil1nillgH CIt the..""
&lId whl.'th(,1 till' hJllall, I might ahlloh1 My minimal. irl(,1'I.'IIIt·ulh whit'h havl' II~I 111'11'11111
thllhll bep;illllingif t(l the' }lllril,(·t adal'ttttioll h.i.VI· alHtl hu,d HoI(l(,tillU valul'.'· WUU"'('I',
011 t)U' oth('rhand (18811.1'1" 1211,127). rlaimhtho.t tlulllI~h ))cLrwin u'I('(1 thl' IIII!'!1 .....Iight "
and "hMall IIIIIO\lllt," lht'HC 1('rIWi u.n' "hardly jl1~tililod," ~ill()(' tht' vell'la.bility IIf
many: impOl-tal1t HIIl'l:'il... J~ of C'Onlliclora.bll' amonnt, and may \'ery "ftl:oll 11(' prl)p!'rly
dOl><!1'l bod 8011 larg<'.
t 'rhe CaRe of KrcLbtoa. i1n[lOl'ta.nt a~ it ollU.'rwiH" 1', ..nl'm~ til mE' to h•• VII
but littlo bl'aril1lC on thill queruon, ~jl1Ot' tho dii!1:an('E' fl'om ihc' mailllo.nrl if! too
trifling,
~ Wa.llaoe (11189, pp. 28, 29) refors to Trifolium reptl18 (txh'lminatmg Phormiu.1II
tenQx,' excellent pasture det'ltroy«!'d in th.re-e years by H'II'JIOChom's rflllimkt, whil.'h OIIoU
oven drh"t' out whit<> oloY!!'r; amI S07lCll'lH oZtmc~u, gt'Owing all OVl'r thl' country up to
all elevation of U,OOO ft. Kirk (1896. p. 18) not only attribllteR thl' .. liihpIlJ.('('mont ..
ot P1&ormium to gr_ a.nd olovl'l'H, but aJ.'IO MllrillCI18 'IutUlntUH, IIml ovon Pterfdillm
CBCuie7ltum (bra.oItnn fom). Fu\'ther 011 (p. 19) he sta.tes that Acipl'1lUa Oole.1IROi iq
grsclua.lly rep1acod by Relf'lIOwn paHtura.ge plautR. Bowl'vor, hI! a.L.o oaJllI at.tention
Lo the effect of grazing a.nd trampling by cattlo and ho1'lleh as a.icling th(' pia 11tH
ill their work, which, of COU1'RE', i'l a very difforont mattE'r from tbl' l'fk'ot of planh,
alonE'.
('O('KAYNK,-Bcol(J[1I('((l 8tllfhO III E l'olutlOli
invadeI ca.n replace thtl aborigina1." On the OthCI haud, although this
foreign host IS pr('sent ill its millions, and notwithstandillg abundY.llt winds
,\ud laud-hirds, t the indiYP'TIoufI 'VNJdation IS still viryin and the introduced
plants altoyethn absffli where gmzin(1 cmitnals have "0
access Md wker(' {ill'S
ha'l)(' ?1P'VeJ1' brt'n, On certain subalpine herh-nelds the indigenous form of
the dandelion (Tarnxacum otfic&nale Wigg,) is abundant, and yet the in-
troduced form, with its rcadily wind - borne fruit., has not gained a foot-
hold, nor evcn the abWldant HUpochot'fUJ rltdwata L" though it may be m
thousands on thl' neighbouring tussock pasture, less than OIll' mile away,
On Auckland Island introduced plants occur only ill thl:' neighbour-
hood of the depots for casta,vuys, but 011 Enderby Island, where there are
('attIe, they arc muth more widely spread. Even whele the ram forest
has belm felled or burnt, and eattle, &e .. are kept away, jt is gra.duall~'
replaced by indlp.'llOUS trl:'(,S and sltrult&-t'.e., ill 10c'IIlitics where the rain-
fall is sufficient.
Some of thl:' Illdigenous specics lire quite as aggressiw. Of eveD more
so, than any of the introduced. In primeyal New Zealand each would have
itb plal'f' in the association to which it belonged-there would bl' no aggrc&-
sion; but when the balance of nature wa.s upset by the fire or cultiva-
tion of Maori or Europeml, then the plants best equipped for oc-cupying
the new ground become dominant, their .. adaptations ,. for that purpose
fOl"iuitously present. Till' miles 011 miles of L,''}Jt08pt'rtllllfll 8Copari'UfrI and
Pte,"idi'Um e8culmuutn were absent in primitiw Nl'w Z~alo.nd. So, too, tht>
pa\ltures of DaIYIJJI.onul sem.iannularill R. Br.t in MArlborough. and the many
acres of CkrY8obactron Hookni Colenso (Lilia('.) in the lower mOlUltain
ftlgion of Canterbury. Oelmisia sp('r:tabt"118 Hook. f., an f1pparently highly
&pecialized herb :£or alpine fell-field or tussock-steppe conditions, is now
on thf' increase in many montane pcl.rts of the Ashburton-Rskaia mountains
and valleys, owing to its heing able to withstd.ud fire, the buds being
protected by a dose investment of wet dl:'cuyed lea.f-sheaths.
NOl' are all the mtl'Odu('l'd spec-it·s IIggressiYe, hy any means. Soma
('au barely hold thpir own j otherli a.re limited to certain edaphic condI-
tions. Thus, Glullci'Ut1l /la'Vufll ('ra.ntz ot'('urs. as yet, only on the coast
of Welhngton, chiefly ill th(' ncighbourhood of Cook Strait. It is con-
fint>d to J(J"U.vl'lly or stony shorcs. and appears unable to grow on the clay
hillside And yet where the latter itl. ill olle place near Lyall Bay, ('overed
with gravd there i& u large eolon~' of the plant, whenc~ none have found
thC'ir way 011 to the adjacent hillsid(·. Lupit&1J8 aroort'1J8, now so common
011 New Zt'IIJaud dUllCS, appears Ullttble to spread beyond the sl..ndy ground.
The oft(,ll-quoted storjes (see footnote, p. 32) of white clover (Trifolium
r('pt'nB L.) being able to wIpe out Phormi'Um tena.J;, of Sali.:& babylonioo over-
eomi.ng the watercress (Radicula Na&t'l.lrtium-aquaticllm). of IlypochoerilJ
* Now Zl."al<1olld may be roughly diVided mto three ",n:I1'>-VIZ., the c.ulth'o.ted, thl'
IId.,turl.' la.IlWl, and tho primitive. It is only in the pasture lands that a h'8l strugglt-
betWNlll tho introduced a.ud the indigenous pl&nts bI te.king placu, and even there the con-
tl!bt ilo very unequa.l, through the gra.zing, bunting, and seed-sowing fa.oto1'l:!. Many
:pal5l:ures, hOll'ove-r, are altogether new aq_ia.tionR, all in the case of forest being felled,
then burned, and the ground sown with ~-. &c.., even lx.fore the ashee of the tJ:to.eo,
are cooled, &0 that at onoo thoro i~ a forelgn pasture brought into existt"uCl' and subject
to an entUely new Bl't of ConditiODH from that wbil.'h 1tO'''eI"IIM the fol'el't. ThiH is certainly
not biological •• replacement."
t Introduced, not native, birds.
t The sp~oiN may be lJ. pilo8a, but 1 bav' no lip 'Clmona for irk-'difio .tiOJl..
~Tra.n..
34 T1"an~nctlonl!.
* ~ht, in a carefull.v considered paper (1911). brings forward a good deal 01 very
suggestive evidence as to the probabilit.r__01 a wetter clhria.te on the ea.st than the pre.
sent one following the steppe climate. The most important fact adduced is the for:mer
presence 01 extensive forests where steppe aloue now exists, since such forest. could OJl].y
be eltabliahed during .. period with many rainy days, and no other apla.nation 8OelII8
to fit the ease.
2*-Tr&D1.
7' r{1" ~l/r"fllI f.
suitabilitv fill' til(' 'Itation. ,\t anv ratt'. tht' l'hILII(,o fOl' nlltlll'.t1 ",election t~)
effect anYthing here is v('ry 1'f'lllOtl:, Ilithouv,h till' ('omp(·titiol1 ill POWl·I'ful.
The number of tru(' rO('k-pll\ntll in N('w Z('lIll1l1c1 ill l'ornplIl'lItivc'ly SIIIIIII i
but, 011 tIll' oth('I' hmul, 1\ grl'l1t 1lI1111~' xf'J'()pitytC'II, IIl1d ('VI'II mC'lIopllytt'tI, 1m'
('nool1nwl'l·d nil ch'~' 1'()I'kll. hilt tllC' laltl·r ar(' c'l'hlll'lIl()nic'nll~' lIlodific·d dmillf!,
th('iJ' illdividual dOVl'loP"lI'lIt.
IlJvon hygrollhyt.oll Illay gllill ,L foot,in"" as IIh'Plltiy IIC'I'1l in tllll t'Ulltl of
Hy1nl'nophylllllll 11II1IIi/i(lllfH. Thl' nWKt Hh'ikill", lind t'l'ul~' Ilmllzin~ C':I8(, is
th.o.t of the kidlley-fl'I'n (7'l'iclwlllam'l1 Tf!'I6i/m'lIU' !i'm'lIt. f.) lind 1l!l1ll("M-
phyUwm IItmgllinol('I&tum KYo'., whiC'h grow in tnt' lull blau 0/ tnl' sun upon
tho laVo. o{ Rllngitoto [sland, Am·klallu lIm·hoUl·. Thl' frondA of botll,
as I saw them on 1\ hot summer's day, werC' dry Itnd ('urlcd IIp tlO atl to
appear dead. hut Mr. Cheeseman informs me thti.t in wintr1' tht' kid.nllY-
fern covers t,lw I'od.. s with itll trn.lIAhl('(·llt fronds, ltnd thHt. thosi' of summer
are not de~d at all. ft SI'(,11111 l'vi<ic'lIt thttt in this ('asc' thl' protoplasm o{
these Il'TI s must bl.'havI· similarly toO t hilt or many li('hC'lls, I\nd this will bo
an epho.rlllOlli(·adaptntioll. TIll' qu('stion I~rilll's, do('s s\lC'h It POWN lit· latent
in these 1111'118 as normlll rllin-forC'At plunts, ferns whil'll ('annot tolcrau· a.
dryin~ wind or II hot 8un IUld ciJoy atlllotlph('re; lind, if so, how c'lIn it }Iavo
possibly 1'01110 ahout t Pmuuhl,v thl' porous fO('k c'ontuius II ",oocl dl'HI
of watel', :\1\(1 th(l I~il' iA IIsw~ll'y !lot dry. Although L do not, think that
any modifiratioJl through th(.' struggle' {oJ' oxitlwnc'c' t41k<,s ph~c'(' IIl110ngtit
tol·k-phl.llta, rot thill ('nsc IIhOWfl t11l\1, one ('annot t('11 !Jut t.lUlt 1111' llIOAt
unlikely spoc'i('s might sottlo in (·c·rt.lin tltlltiol1s, and so innugul'uh' It n('w
line of descent, no IDntwr how th(' evolution btl bL'Ought ahout.
In olos('d formations tho struggl(' for existollcC' h('tweell indh"iduals is
very keen. As I write, in my gIl rd('J1 , in 1\ bcd ('I'Owdl.'d with inc1igl'llOUS
plants, two rapidly growing a.nd fu.r-spl'l.'a.dill~ Clmtllll.m Islund h('rhll Imvo
encountered, Imd one (Pmtia, a,ren.a.ria. Hook, f.) is rapidly rllph\cin~ the
other (Ootlila. JJlUflleri 1'. KiI'k) , II happC'ning 'Iuite ill nC'C'ordnnc(' with thc'
fact that tILe fornwl' plmlt is 011C' of tho most widely spl'l'lId of till' Chl\thalll
Island pilluts. H.c'dueled ta its uitillllltt' f:wtc)I's. thl' strul!J!,l(· is chi~f1y Olll'
for nutriment in its willl·st 8111\8(1, lUi (tlcmWllis hllll showll (1905, p. 286) j
there is littlo Iwtw\l nllstru('tioll (1£ OJ1(l pltlnt hy anotlw1', though tlwy fU11I'-
tion illdirllutly hy 11l1tt.iug off light, using up Ilutritiv(' snIts, &c·. In somo
CQfIM thCl grN\tel' pRrt of t,hn Rtrllgglc> tl\kl'll plaUt· nmollgst t,hc yuun", phmtol'l,
I\ll(l it is on theil' l\dapl,l\tiolUl, whidl IMy tliiTlIr mu('h {rom 111()III' II( t,ho
adult, t.lmt tho 1"'1tA,hlishlllllnt of t.hll h,tt.ol· dopotl(lK. Thill iii IIPC'C'iI\U,v c'vidl'n('
in t.hOflC' hett'roblllKtiu 1:I11lloieli 1l)!'<'I\dy dl'nlt wit h whi<'h have· c'('ulogi('lllly
different, fOl'ms in thoil' clifft'l'nnt IIt~'gC'II, In 1\ £or('tlt thl' c'ontiiticHltl for
tho soodling "nel su,pling ('r('('s 1\1'(' v('ry diff('rent fr()m thotll' to whi('h the
adults Ino oxpos('d, A fuv()ul'"blc' v'llril\tion which might prl'Ht'I'V(! a S(I(1(I-
ling in the struggle' with ita onvinmment would possibly hk\v(' littlo to() do
with tho imperative domands of the Ildult. Smnll oUtwa.l-d modifications
of a very few individW\ls could hardly be preserved in the dense growth
of saplings* in an upland forest of Notkn/fJ{}U8 oliOortioiar8 Oerst. The
chief requisite of sUC'C'OBR here is rapidity of f!;rc.wth,t II physiologica.l
• The saplingll may grow II() oloFoly that olle aa.n hardly fOloo a. paHHagO 1hl'II1l8h
them.
t The 0IlI:III deborlbed in my little book, "New Zo&Ja.nd Plants and their Swry,"
of a. spooiotJ of 1I'IfIJQ1JJ1JW.8 overooming tho eminently aggrcAAivo .Lept08fJ87'111um.llC01/(IrWm,
through its more rapid growth, bntll gc-rminating at tho Nomo tim.., iH ilUltruotiv(1 in
this regard
COOUYNB.-IJeologieal Studie8 in ElJolution. 37
charu{'teristic that, b.owover much intensified, oould bring about no specific
differenccs unless corr~la.ted with structural change. In point of fact,
thl! deciding factor in the struggll' amongst a close-growing mass of
t.hese tree seedlings iii probably ago. Could all commenco on exactly the
sllm~ footing, then the determining factor would be the llituation with
regard to t,ho food-supply and tho illumination. and 110 slight beneficial
lfiudifi('ation would count in compa.rison.
As for th(' adult forest-trees, ea('11 haFl, as 1\ l'ull.', its own rooting-plaC'e,
and it!:! death dl.'pf'nds chicfly upon its age, partly upon some disease
or other, and but little upon the superior adaptations of its neigh-
bour. Its growth-form, rertainly, does have something to do with it!:!
IOllj(evity, a.s wher,' spreading branches faVOUl' tho presence of abundant
epiphytes. whose wl':ght may lead to damap,e and permit the attack of
fun~i.
A mixed rain forest, I\pal'i: from modifications due to the nature of the
topography, might be expected to offer constant conditions extending
over a I'onsidorable period. But this is not so; topographically similar
pam of a fOl'cst may show dissimilar undergrowth, the result of conditions
which, similar at first. become dissimilar as the vcgl'tation develops. Thus
in the Waipoua Kauri Forest, of which I made a special study, a state
of change rulod. In one part there was little undergrowth, and in another
part, such in abundanc~. This latter, in time. will, through survival of the
fittt.'tIt, ('hango into forest with little undergrowth. ThoFie are two climaxes,
and are ~xprcssions of the light mctor, the dens~ undergrowth denoting
the maximum and the final open forest with tho close roof tho minimum
of illumination. Between thcs(' two climaxes there are ma.ny transitions.
Bring in moro light still and so inrrease the xerophily, tho hygrophytes will
1(0 to the wall, until, with excess of light, a transition forest and finally
a .LeptospB'rmum heath will hI.' f'stablished (Cookayne, 1908, p. SO). From
the above it follows that, even were Datuml seleotion at work amongst the
yotlng plants of allY species, owing to thE' varying change of conditions
brought about by these plants themselves there would be an insuffioient
1eU¥th of time for any more suitable va.riety to arise, or, if such selection
were very rapid, difforent typos would be solectod within a quite limited
area. The helievl.'r ill tho efficacy of opharmonir variation woUld say that
forest-t,rces have arisen from shrubs, or 'Vice IJ81'sa, owing to the stimulus
of cdaphic, climati(', and other factors, and that selection operated byelimin-
ating those individuals which did not respond epho.rmonicaUy at various
stages of the pl.llonts' development. And the special evidenoe put forth
would be that :many speries possess an unfixed epharmonic tree form and
shrub form, while it is known that stature and other features can be modi.fi.ed
through changes in nutrition. This, after all, is only Darwinian sel('ctioll
plus an assigned ('ause for rapid and sometimes favourable modificu.tioll"';
but it is far from bei~ neo-Darwinian selection.
... It is pJa.in from the very uture of tho caRe that perfect harmony 0ILIl novel' 00
_bliahed 'be1.wecm the growth.forma a.nd the ha.bita.t, since eba.nge, progressivt' or
retro~ve. is & feature of all formatiow!, and growth.forms once epb.&rinonic will
persist long after their epharmonio relation is wP&keiled or destroyed.
t G. Lgalli' has l&ri8r. thinner, a.nd much less hairy loaves t&.a.n the eastern O. ,.ibi-
folia. '!'hey have also cTrip.tips, whioh are frequently 6trongly developod. The juvfl1i1c'
forma aM aom.ewhat aixuila.r in the two trees.
t This term. " hoath " 1 ha.ve used in my writUurs in default of & better. wellmowhlg
the formation is not truly analo~us, except after 1i1'8. with the heaths of Europc.>.
the settlers. when fun 8!'Own, it 18 known &8 "manuka .. or " tea·tref> scrub." At tb."
B,
stage it ill rather foreRt than bf'&th.
VOCKAYNM.-b'cologlcal Studies In b'l'oilltllJlI. 89
full-sized trees of WeitmUlnnia racemOBa L. f. Sophora tetraptera J. Mill. is
restricted in Chatham Island to the forest on limestone near the shore of
the Te Whanga Lagoon, though elsewhere in New Zealand it can grow
abundantly on rock similar to that of the rest of Chatham Island. The
'\ccompanying trees are the same ill the limestone forest as in forest of the
Island generally, but it is evident the volcanic rock of the remainder of
the island favours the other trees. which do not allow Sop'huta to become
.'stablished. Or it may be that Sophora is a comparatively recent arrival. *
The distribution of certain species shows that epharmony is by no means
110 l'omplete between plant and habitat in some cases as one might expect;
or. in other words, that a plant can live ill a position for whioh it is not
perfectly fitted. Thus, Mr. R. G. Robinson, Superintending Nurse-ryman
tor the South Island, informs me that the dominant tree of the Tapanui
Forest, Nothotagus Mtmziesii Oerst., cannot be grown in the adjacent State
DUl'S6ry, although N. /wea Ocrst., a comparatively rare plant in that
locality, can be grown with extreme ease; and yet I have seen N. Menzieaii
It,oowing quite well 011 the flanks of Ruapehu as an isolated tree in the open.t
The slow growth of many indigenous trees as compared with introduced
species is another case in point. On Antipodes Island the plant-associa-
tions are oot distinguished by their different Horistic componeniB so muoh
dS by the relative abundanoe of the different species. This word abund- tt
Jnce " shows that aU are not equally suited for eaoh station, but that if a
plant settles down on ground not. spooially fitted for its requirements it may
he able to hold itS place. the struggle for existenoe notwithstanding. So.
too. with various stations on the Auokland Islands. A highly specialized
species may thrive under conditions that might be deemed impossible.
Such a case is the already mentioned hygrophytio almost a.quatic Trioho-
lIla'11B8 renit0fm6 on the sun-baked rocks of Rangitoto. Here are a few
more pxs.mples: Orassula flwschata Forst., a coastal halophytic herb, is
one of the pioneer plants in the heavily manured ground just abandoned
l>y penguins on the Snares Island. OoZcibaHll,hw mvsc0ide8 Hook. f., a.n
herbaceous dense cushion plant growing normally on ooastal rooks, is an-
other early-comer on the above manured ground, but as conditions become
favourable .for less manure-tolerating plants both are replaoed, tussock
moor or Olearia forest being the climax association. Met1'OBiriHros ~
&1., a root-climbing woody forest-liane, grows in some places on rocks
close 00 the seo.. Griselinia luciiJa Forst. f., so far as I am aware always
aithl'r an epiphyte or a rock-plant, can be oultivated with ease as an
.ordinary gllrden-shrub.
The presence of olosely related species side by side in the same assooia-
tion has a strong boaring on the mutation question, for it is reasonable
to suppose with Leavitt (1907, pp. 210-12) that if natural selection, or
even epharmony, is responsible for species-making, only one type will be
present. .As Leavitt writes, .. Mutation breaks the species, and moment-
arily at least must give a polytypic aspect to the group within a specifio
• H. lL Travers (1869) was of opinion that this tree was a very reoent anivaJ,
espeoialJ.Y all an old :resident, Mr. Hunt, did not know it, and as he found a seed on the
!d1ore of Pitt Island. I have given my reasons for be e~' it aD a.noient OOllatituent
of the 8.ora. (1902, pp. 270-71), and h&ve seen no reaBOll to my opinion.
t The case may not be as strong as it appea!8. since the . are ahade-loviDJ.
whereas those of N. ftucq. oa.n tolerate far stmnger light. Th&1'e is al80 .. fiDe tree m
the dry Chri8tchuroh Do:rnam, wheze the olima.te is muCh mon unsuitable for indigeDoUi
.forest-plants than Tapanui.
4.0 'l'1'(lII~at:/IO'tlh •
ar(.'a" ('00. aa., p. 211). 1 uannot go fully into t.his important matter,
but th(' following are rather striking examples. Many would not (Jonsider
sOme of thE'sc plants .. specIeS," they art' so close; but so long 8S they ate
distinct C'nt.itics ",hi('h rC'pl'Oc} 11('C' t.h('mselvl.'s .. tru(''' they ml'...t the ell&'
1111 woll 01' [)('t.i('r.
Dracollh!lllutn lI(!npari'l.llll Hook. I.. Ilnd !~llothol' SPl'C'!('h c'()Jlsiderud by
(...t}}CCS(,llll1ol\ II form of this Hpc('ic'b (J909, p. 420) bui by Kirk a var .. of
n. Urvillel1ll&ulII, gl'Ow III the /:ll'ruh of Campbell Ishmds. ('l'lmill'ia /)(!'f''fIACosa.
Hook. f. lind O. rampbelleflais Chapmllll, Il very raro plant. grow sido iJ~
sid(' in Auckland and CampLell Islands. Ootula Traillii T. Kil:k, O. pul-
ehella T. Kirk, and C. (obscura T. Kirk) ? grow together on l'oastal moor
ncar Foveal1X Strllit. Two" species" of Acama grow side by side on
dun('s in Southland: the on(.' has more or less erect branches and long-
peduncled {iowelt.-it may Lo a var. of A. ,nt'crophylla Hook. f.; the other
is pressed lJ10st tightly to the gl'Ound, and has almost se88il(' flowers-it IS
A. mWrophylla va.r. pattciglochidial.a Bitter. Both forms kl.'cp their dis-
tinctive ('haJ'lLcters for yeurs when grown in garden-soil; intermediate
(orms occur amongst the wild plants which may bc variants, mutants. or
hybrids. Vatu/a airaJ.a Hook. f. and C. Dmdy", Cockayne lip. ined. occur
on the lIame shingle-slip. Soveral absolutely dist.ill(,t forms of VerOt'lica
bua:i/olia Bellth. grow on the same subnlpine herb-field (see Plate II
fig. 1). Rubus pl1ll'W8 Buell. and R. Barkeri Coekayne are in nt'IU
proximity in thl' neighbourhood of Lake Bnmncl', W(.'stland. NotllO/agw
fusca Oerst. and N. apiC'Ulata Cod.cayno grow in tOmpHny III thE' fOlll:lt ..
at Day'l! Bay (We-Ilington) and Kaikoul'a (Mal'lbol'OUgh). Alltflia li'l&l'af'I"
Hook. f. and A. 8ubulata Cheesem. grow side by sidl' on mountain·moorN 111
Auckla.nd and Stewart Islands. Raoulia aU8tralis Hook. f. and R. lutesCffll>
Cockayne grow side- by side on l'iver-b(.'ds of th(.' Sout.h Island OlRarw
Oolmsoi Hook. f. nnd O. Tra/iUn' T. Kirk grow mixed together in coastal
scrub in Stewart Island. 0fJ88inia albida Cocka VIlI', O. l'(IIIw&lU(f'8ti Hook. t..
C. luWida Hook. f., and othel: closely t('lated intcrm(>d.iate forms grow mixed
on Mount Fyffe, Seaward Kuikoura Mountains. Two tlistIDct. {OI'DlII of
(Yf.Ul8i",ia Vauvilliwsii grow just uhov(' the' forest-line ill Auckland Island
(SCI' Cockayne 1909A, p. 216). (IntIlLa lnmata Hook. r., U. propinqua Hook. t.,
ILnd O. plun&osa Hook. f. grow side by sidl' 011 the- sho]'E' of Auckland .lud
Camph(.'ll Islands. Olearia ilici/olia Hook. f. und O. mol/fa Uocko.Yll(· grow
tog(·thcl· ill subalpiul' fores1 of Westland. R(.'lut(·d Bpi/(}hia grow lIidc' by
sidl' ill nmny pla('cs; SOIllI' I kuow C'ODlC trm' from H(·(·d. POll 10111111(1
Hook. f. and P. TeflhWllltiu9lIJ grow ('Iose wp,ctlter ill Am'klund [Hl:mll.
Otl'llisi(1 lIessiUf/,ora Hook. r. IUld (1. QJ1'gentfa T. Kirk grow lIidl' by si<11.l
on ('ortllin alpin(.' moo)'s of the lIouthE"rn botani('ul provinc'(" Nolllllpa'fl.a.'C
simplea' Bellm. and N. panntm VockaYllc "1'1' I'ompanioll plun1.s in the tot('lSt
of l:itewart Island and W(.'stland. (1armich.al'lia Mm&rn; Hook. f. and a
rl'lat.cd but more robust species not yet. described* gro," sidl' by side on
steppe and. riv('r-bed of the Canterbury Plain and easterll Routhern Alps.
Oopr08t'1la Petrlei Oheesem. has two Corms, one with claret-coloured drUPC'II,
and the other with faintly blue drupes; they grow side by sid(.' on montane
steppe in the South Island. OOpr08tnfJ Oolensai Hook. f. and O. Bam,k8'ii.
Petri(.' OC('ur side by side in many forests. RafltfMl,CUw.s Lyallii Hook. f. and
... What I t.akt' to be thih pla.nt received tho herbarium !tam(' of O. lH6miUs from
D. 'Petrie XDA1I.y years ago. It has aJs4 been in cultiva.tion Along with (Y. Motll'Of' Book. f.
in tJuo (JhriRtehur('h Domain for a long peoriod.
VOCK.\YNJJl.-Ecotog~cal 8tud?6S m ElJolution 41
a. plant I considered R. Traversii, but which Cheeseman IS of opinion IS
either a hybrid· on a new species, grow together on the Snowcup Moun-
tains, Canterbury. Ranunculus BuchOlM'Tli Hook. f. and the closely related
R. Matthewsii Cheesem. grow in company on certain alpine herb- or fell-
fields of western Otago. Anisotome pili/era Cockayne and Laing and its
var. pirvnaJ,ifi,rJ/um T. Kirk grow in company on peat-covered rocks, &c.,
in the Southern Alps. Leptosperm'Um ericoides A. Rich. and L. 7i?&eatum
Cockayne grow together on northern dunes. Oonaria tmg'UBtissima Hook. {'J
O. thymi/olia Hunt. & Bonpl., and O. 'f"UBoi/oUa L. grow in proximity on
Westland riwr-bcds. Aoiphy11a Oolensoi Hook. f. var. conspic'Ua T. Kirk
and the val'. mazw-na grow near one another on certain herb-fields or in
scrub 011 the Southern Alps. Two forms of Ourisia sesBi7liflora Hook. f.,
the one densely villous and with large flowers, t the other d. smaller plant
in all its pam, the leaves darker green and less hairy cl.nd the flowers
fewer and smaller, occur on the same herb-field in the Southern Alpll.
PittospO'I"Um riuidum Hook. f. and P. rJ.ivarioatum Cockayn.· (see Plate I)
occur in the same forest-area on the volcanic plateau. /3opkOla micro-
phy11a Ait. and S. pr08tr'ata Buchanan grow side by side in the bed
of the River Waimaka.riri at the lower gorge. Doubtless a number of
other examples could be found. The coupled plants are in all cases so
closely related that they are considered by most New Zealand botanists
either varieties of one spscies, the type and a nriety, or forms not worthy
of or that have not yet received a name. They are quite sufficient in
number to show that it is fWt 'U7'6UB'Ual lor closely relafRd 11,prellitary p/mnt
mtities to eziBt side by siiJe lor COf&siderabZe periods.
The oocurrence of distinct races of the same species at dIfferent points
of its area of distribution is known in a few cases. As Leavitt say&, such
cases do not look like the work of mutation, nor can they he readily corre-
lated with epharmony. ·The following Ilore two stl'iking examples: Rubus
GUBtralis Forst. f. is a common plant both in forests a.nd the open through-
out the North, South, and Stewart Islands. In the northern part of the
North Island it has, a.& a rule, mueh narrower leaves than in the &.Outhem
part of its range-so much so that typical plants from the two areas
have a very different appearance. The primary seedling-leaves seem to bl'
identical in both forms: these are ovate or ovate-lanceolate, and coarsely
toothed; they are soon succeeded by narrow leaves, much resembling
those 01 R. POll'VUS Bllchana.n, even as to their yellowish or slightly brownish
marking. Seedlings growing in the forest-shade, and only 2'5-5 uDl. taU,
bear these narrow juvenile leaves, thus showing the form to b(' inherited,
and not merely an epharmonic sun form. Sinco heredity is undoubted,
the northern form demands a name. StypheZia, laMiAulata (Forst_ f.), a
heath-like small or tall shrub, extends from the North Cape to Canterhury
and Westland. The adult form varies but little throughollt its range,
but the juvenile of the Auckland district has altogether broader leaves
than that of the south. An example of a more local character is that of
the mountain-herb OeZmisia coriacea Hook. f., which from Mount Mau-
ngatua and other mountains in that part of Otago can be distinguished
at a glance 88 a cultivated plaut from other specimens collected on the
*ThB ooourronce of this p.la.nt on WaJkor's Pass far tram R• .iUonroi Boo!... f.
removes the suspioion of a. hybrid origin, whioh Cheeaemam adopted, p.lol"tly at my OW'll
Bugge'ltion in tho first instance.
t rro lim plant I gave the MS. naDlI! of O. spZeMldtJ some years '\(lU'
42 'l'ra'l/,SftCf10IlB.
actual dividing·rang~. The lowland form of thl' plant growing ne~r tlle
6ea-cli:tis a.t ChaTleston, w<!st Nelson, is also distin(·t in appell,rancc.
2. 18olatiO'l'l.
'rhis spcuial forlll of distrihutiOll ill l'OllSidcl'l!d by !!lome to hl' III tht·
~rea.tClst evolutiona.ry importIUlCl'. TIll' New Z('I~hmd biologicl~1 urclL ofic'r"
many ideal localities for geographical isolation, difieriJlg in d('Il,l'('(:', and it
is intC'rt'8ting to soo 8S to how far they u.:ft'ord ('xampl('s of rclu.u,d IIPCOit·!>
which a.ppear to have either diverged recentl!l from II stE'm form. or Ollt:
of them to be the actual parent plant.
(a.) The Kermacleo IBlafIGs.
The total number 01 species of vascular plantS is 114, of which tWl'hre
II.re endemic.* These latter, one cxcepted, are closely related to, and iu
IIOme instances almost identi('lIl with, Now Zeo,land, Polynesian, or Norfolk
Island plants.
(b.) The Three King' l,lCIfIIU.
There is strong geologica.l evidence that a.t no distant date th~ islands
werl' united to the North Island.t The total number of species of vasoular
plants is 143, of whieh five a.re endemic; with these A ledJryon e:rcelsunl.
Gaertn. vaT. graMtll Cheesem. may be included. Cop'08ma mo.crocarpa
Llb-eesem., one ot the five, is related to (I. grtmiii/olia Hook. f" and 1ll0r4'
distsntly to O. ,obusta, both of which are present on the island. Pittospo1li1ll
FtWrcMtilM Cheesem. is allied by P. clCJ8sijolium A. Cunn. and P. UllIbpllntulII
Banks & Sol. Veronica insularia Cheesem. is related to V. diosmae/ollfl
R. Cunn., a. species of the neighbouring mainland. and Parof"opkis Smithil
Cheesem. to P. opaca Brit. & Rend .• while the fern Davallia Tasmani Field
is not allied to any New Zealand species. None of the endemic pl!l.nh,.
then, except the A1ect'7lon, a.re particularly dose to their mainland allies..
(0.) Tile North Oape.
This high promontory was undoubtedly quitE' J'ooently 11.11 island. The1l'
are three endemic plant&-IIalorrhagis oartilaginea Chcesom. (a Ilear relativl:'
of H. erecta Behind.), GHl.iostflllla ligtlstr·ijoli'll.m A. Cunn. va,l'. crMlllWI
OheeAem.,* and OaBB1'nia amocna OheesllDl. (probably relnted to (1. l'llu-
tJilliersii Hook. f., but which lattur is not (olllld nl'al'cr than tilt' '"oil'Nui(
pla.teau).
(u.) islands lying tl) tilP ltJnhtu'Urd of .tllcklalld.
Veronica .BoUonst·i Co('kaYlle. n spol'ios closuly l'ell,ted 10 r. IJltUJrUUTll
Hook. E., is cndumic on the' Poor Knights Islands. Pittollporwm int6l'-
I'II6flium T. Kirk, intermediate between P. teftui/oliu.m l~nks & Sol. and
P. ~ T. Kirk, is found ollly Oil Kawau Isltmd; only OIlt' phmt
has been found, and this bas been d('.stroyed.§
(e.) 1.'116 OTUltliam /lllaniJB.
The tota.luumber of species plus named varieties is 286, of which thirty-
one are endemic. The genera M'IIosotidium and Oo:eelZa are endeJ1lic a.nd
monotypic. The following is a. list of the endemic plallts; those related
more or letlS closely to New Zealand species are marked with an asterisk:
Adiantum affine WiUd. var. chatMmicum Field (Filic.). *PO(], chathamica
Petrie, Festuca Oozii Hack. (Gram.), *Oar6Z appr6B,a R. Br. var. ,ectoideB
Kiikenth., *Phormium te1'&tlZ Forst. var. with broad thin drooping leaves
(Liliac.), Geranium T,ooer,ii Hook. f. and var. elegans Cookayne (Geran.),
*unum monogynum Forst. f. var. o1wtkamicwm Cockayne (Linac.). *Plagi-
",mnw betulinus A. Cunn. var. ckathamicw Cockayne (Malvac.), AciphyUa
Tr(Jf)(fl'rii Hook. f., oouZZa Dieffenbachii Cheesem. (Umbel.), *Oorolda
InGC'fOCMpa T. Kirk (Cornac.), *Styphelia robusta (Hook. f.), *Dracophyllum
arbo'feum Cockayne, *D. paludo8um Cockayne (Epacrid.), *Suttrmia Oozii
I~ockayne (Myrsinac.), *Gentirma okotJw,mica Cheesem. (Gentian.), Veronica
DiefJenbaohii Benth., V. Barm Cookayne, V. DO'I"I'ien-Smithii Cockayne,
V. ohatkamica Buch., *V. gigantea Cockayne (Scroph.), *Ooprosma okat-
hamica Cockayne (Rubiac.), *Olearia semidentata Dcne., "0. ohatham.ioa
1'. Kirk, O. T,aversii Hook. f., *OotuZo, Muelleri T. Kirk, C. Featherstonii
F. Muell., *Stmecio radioZatw F. Muell., *S. H'l.lmtii F. Muell •• "Sanehus
granilifolius T. Kirk (Compos.).
The nineteen .. species" marked with an asterisk are closely related
to forms found elsewhere in New Zealand, while sixteen of these are very
close indeed. Veronica gigrmtea would certainly be considered a variety
af V. ,alicifoZi,a Forst. 'Were it not for its distinct iUtJenile form, which stiU
perrists up to a BttJtu,e of at least 80 om., and its arboreal habit. It is the
I)nly true forest-veronica, and it may be that the juvenile form is a direct
adaptation to forest-undergrowth conditions.
(1.) Btewa.rt Ialand.
A number of species have, as yet, been collected only on Stewart Island,
but in the face of the fact that year by year shows more of the plants
thought to be endemic fairly common on the mainland, &c., it is quite
possible that the island contains no endemic species.
(g.) The Sv.bo.7Itarotic IBlanda of New Zealand.
'l'here are 195 species and named varieties, of which fifty-one aro
Ilndemir., nineteen of these being closely related to New Zealand species.
No list is given here, as these endemic species are treated of by Cheese-
man with considerable detail (1909, 'Pp. 463-66). With regard to special
endemism in the various groups, the Auckland Islands have six speoies,
the Campbells four, Antipodes Island four, Macquarie Island three, and the
Snares two.
(h.) Isolation o~ tke Main Ialarula
Endemism is not confined to isolated islands, but the various floral
districts contain their peculiar species and forms. The most striking
examples are western Nelson and western Otago, with thirty-three and
thirty-eight endemic species respectively. The northern part of Auckland
(thirteen species). Marlborough (fourteen species), and other localities show
a distinct local endemism. It is obvious, then, that a strong endemism
can exist apart from such a barrier as a wide stretch of ooean. But :figurea
.such as the above uro not :final; further investigations may decrease or
oven increase them. Also, it is certain that not aU the species included
have originated in the .. isolated" areas; some of the most distinct have
probably been much more widely spread, and are ., relics" merely.
The continuity of distribution of species of the New Zealand fiora varies
from those with a fairly continnous distribution to those which occur in
only a few localities far distant from one another. Notable examples of
extreme discontinuity are: DamAotMa fJfttarctica Hook. f., common in
'flI'O'flSflCtiOtll •
Auc.k:la.nd ILnd Campbell Islauds, hut eOllfull'd (list-when' to II. tew rocky POlllt"
ILnd small islands in the fur north of tlw North [glaud; Urtica (Justt'td?$
Hook. t, common in Chathu.m, Antipodes, nnd Auc'khmd Islands, but in N('w
Z(.'aland pl'Opl'l' occurring only on Dog and COlltl'l' Islands, Fov('lJ,ux Htrall j
/)roset'a p!I(/maea D.C.• only re('orded from 1l01U' KaitailL ill tItl' l'xtrl'ml'
north and the Jlluii Hill in thll extl'l'lllC' south. I'ittosporum o(JOlJrdatum
Raoul, ()('c'UI'S 8parillAlv noal' Kuitaia, and Aklloroa, 13lmks PllIlimmla j Pla'll'
allikuR eymosu/J T. KiJ'k, only I'ecordl,d from J)unpdiu. Lytteltoll, SOIliC'
of the Marlborough Souuds, and Ktt.itaia; S'IIIJ,mwl. cilafJlUmica Moz, com·
mon in tho (Ihlltham Islands, and fotnld ill two 10('ulitips in Hh'wart Island;
l.epyro(Jia Traversii F. Muell., common ill Chatham Islllml, and fotnld ill
c'ert&in bogs of the.> Wnikato and Itt OllU locality nellr Ko.itaill: StY'Pke1ia
Rickei Lahill., ('ommon ill Chatham Islalld. clnd found elsewh(.'rc only neal
the North Cape, Melic/pus macrophylluH A. CunD., l'ommon in ('(.'rt&m
Auckland forests, but absent oisewhor(', oxcept one 10(,lllit,y I1l'a1' DUlll:'din.
Other !.'xamples uf dilIcontll1UOUb diHtl'ibution, HlOUgl1 mol'C' COlUlC(.'wd
than tho above, iu('ludc RloPOC~((riH Rphaoelata R. Br., DraoopltyUum lati-
folium A. Cw.m., (!lelllatis «foUata Bu('h., Quinliflia oouti/olm T. :Kirk,
Velmisia Traversil IIook. f., l'RPUlio1)fJ7'&(J(J' fet'o:J.) T. Kirk, Oarmio1w.elia
!I"uilis J. B. Al'mstg., ()oprosmll rubro Potl'J(." Vet'on;ca speciOS4 R. l1ullll.,
&('. Wen' thero merely 0111' Of two (lILSC'S thl' disc'ontilltlOllS distriilutio1l
might bo attributed to Chl).l1('O, but as thore Ul(' Humorous ('I.\St.'8, and ~b
these gradually merge into examples of greater and Arl'atel' continuity.
it is p:oobo.ble that the species in most cases werl' at Olll:' t,imll mOft' widely
gpread, and that in the extreme cas(.'s as abov!.' w(' are face to fae'l' with
the phenomenon of a speoi<.'s llI\turll.lly on the' V(.'rp,e of ('xtiDction,
'r.ne New Zeslalld bpccies. Wlth but few exccptions, IeproJuce them-
selves readily and rapidly from seed. um be easily grown from cuttings,
nnd are not restrIcted to any special soil. Some respond qwckly to chang(>
01 !.'nvironment. The genus oc{'urs III 1I11 parts of the New Zealand region,
except Antipodes and Macquarie I&lands. It has representatives in almost
every plaut formation, hut there is ouly Oll!.' true forest species (V. gigamea,
of (''hatham Islllnd). Au ll11alysis of distribution shows that seventeen
species arc {'oastal, thirte!.'ll do not Ilbct'nd beyond 300 m. altitude, ten to
between 300 m. and 900 m .• thirt(,!.'ll to 900 m. and less than 1,200 m.,
and forty-three to that altitudc lind up,vards, whIle fifty-two of the
ninety-six may bl' considered strictl~· mountain species.
Regarding th(>u' growth-fol'Ill.8, pClhaps blX species mi~ht be consideredl
herbaceous; the l't'mainder cl.t'l' all more or Il.'ss woody, thl.' great majority
being shrubs. Beyond New Zealand there is one shrubby Veronica in
Fuegia and th(' Falkland Islands, r. elliptica Forst. t, Identical with or
closely related to one or other of that beries of forms known as V. elliptica
III New Zeal.loUd, and 1'. formosa R. Br. and V. dem,/olia F. Muell. of
Tasmania alld south-east Australit\ respe{·tively.
Leaving tho hl'rhacoous cushion plants, formerly referred to the genm.
Pygmaea, 011 om' !:lid!.' for the prl.'sent, the remainder of the herbaceous
lIud sufirutl{'Obl' wro11lcas (Divisiou EUlJf!ronica J. B. Armstg.) are dis-
tinWlishod by theu' dtdy1l'&o'US capsul(>. But the shrubby V. loganioiiles
J. B. Armstg. halo Il. similar capsule. This plant resembles in many
respects a juv£'llile form of th" whip!.'ord section of Division I, Hooe. There
18 anothel' cphllrmonically snnilal' plant. V. CaBSOOOide8 HorL, which has a
capsul(> of the IIebe type, and which represents a fixed juvenile form of a
whipcord Yeroniou, such as that fixed or semi-fixed form of V. t.et'l'agqna
Hook., which OCCUTb occasionally on the volcanic plateau (see Plate V.
fig. 2). Still mort' IS the relation to whipcord veronicas shown in the
toothed leaves of reyersion shoots. With a broadening of leaf, a not un-
('OmmOIl occurrence, thel'l' is a close II.pproac'h to V. bt.£zi/olia, Benth. In •
considorlllg the phylogeny of the species of Ymmwa the change from herb
to shrub would be I.'pharmonic, as may now be seen in the series of forms
from just sufirutwos(" to almost shrubs. III such manner V. logamoideR
might Misl', ,md, tho form of C'o.psule cho.n..,';'nl': by mutation, there would
he 1r. cQ8sinwidl's, which 011 the OllCl hlmel could develop by war of V. "buzi-
/(llw illto th!.' nJcsophyti(' spc('ies, or througll pressing of leaves to the SteDl,
11nd n l'(>l1iaiu amount of rwuctioll, into the xel'Ophytic whIpcord forms.
Of l'OUl'bl', I do not lDu1ghll' thesl' art' the actual 11ncI.'stral species, but it
do!.'s !lot sC(,lll. ILhsul'd toO tak!.' them as approximative to such, Som-e-
further details mny shod a httle light on the matter.
The shrubb~' wrollico.s full into three opharmonic classes, using Cheese-
man's synopsis. The first would include n'Om }'. 8pecioBa R. Ounn. to r.
pimeleoiile8 Rook. f.. the second from V. Gil1i68iana T. Kirk to V. WIVI.'fIqra
T. Kirk, aud the third fl'Om Y. macrantha Hook. f. to r. Raov.lii Hook. f.
ThC' first class shows a leaf gradually decreasing in size, and varying from
the willow form, broader 01' narrower as the case may be, to the small morc
or less oblong or ovate leaf of so many of the subalpine species-that is,
there is a. reduction of leaf-surface in lI.{'cordance with inerease of xerophytic
conditions. Where lowland species Ol'CUpy xerophytic stations large leaves
a.re thickened in texture, as in V. DieDf!fIbn£hii Benth., r. spetJ/OstJ R. Cann.,
and V. macroura Hook. f., all plants of coastal rocks; or reduced and
thick(>ued. ILS in r. chafhamica Buch., a.nothel' t'oastal-rock plant; or mut'h
46 Tran~(lct10n.1l.
X. CONCLUDING REMARK!:!.
The objec.:t of this paper is to supply material for consideration by
students of evolutioll culled from a field which, although not altogether
neglected, is much less cultivated for the supply of evolutionary pabulum,
especially by English writers, than is the wide domain of zoology, whence
come the bulk of the facts of so many works on evolution.
Whatever of value there may be in this ecological material lies in the
fact that it is drawn from an isolated and virgin vegetation, and one, too,
where the grazing animal played a most insignifiC'ant part compared with
its rOle in the Old World.
The details have not been selected to support any particular theory,
though, of course, as ecological observations are the basis of the papor, the
relation of plant to environment takes the l~ place.
By one celebrated school of biologists the ultimate inheritance of cha-
ractem. evoked by stimuli affecting the body-cells is either considered
impossible or an occurrence so rare as to be negligible, while such evidence
as I have advanced, is looked upon as worthless, or, at best, as quite
manywritcr
altogether his much more convinCing" Origin
lugpatdve material.
48
insuffioient. But another and equally tamous sl'hool bl'hove hucll inht'rit-
ance to b<, ,\ more or Ir811 frequent O('(,UrTt'n('(', hot,mists, as it rull.', bl'in~
more in its favour than are zoologista.
Sp<'llking of thl'Oril'S or evolution generally, there SN'ms 1I,00d r('ason
to rOllsidl'f that such. if not pr(,llmturl.', I~rc ('hil.'fl.y of Va.llll' ..IS a stiml11118
to biologiclll mlcardl. Our ignornnc(' I\S to thl' minute strurturl', tht.'
chl'mistJ'Y, ILlld the physiology of the protoplasm is pr010llllu. N()thlll~
il5 known as yet J'eglLl"ding tho Iwtual 1',\UlII' 01 v,Hiation. All ephllrmollu'
tltimulull could do nothing wore it not that thl' innl'1" l'Ollstitution o( tht.'
plant is "'ready Il.bl<' to respond-i.p., thl' ., mnehillllry" iR thllrl.' ready to
produr,' the possibly rphannonic variation so tlonu itS it ~cts tltl' lll'CI.'ssar\'
tonoh.
The ronstrU('tioll of 1'1ll.borak' theories is not the method. by wlm'h
progrl.'SS ('all be made. Actual experimonts in the gal'<il'll, thr laboratory,
a.nd the field can alone lead to thl' truth. Even in taxonomy, only expl.'l"i-
fllent ran artually decide lUI to stabll.' nnu hl'feditnry forms. But OLRI'I'YII-
tiona from nature o,rc also demanued, and here croloAY coml'S in, with tho
attempt to makl' USl' of the wild-pJ.a.nt world, where thert' a.re spl:'dl's in
the making, as II. sourcc of obsel"Vntion. Thl' duty of thl' c('ologist is tiLl'
eollocting uf facts ;n 11S a('curato n manner rs possihlr. TIll' KtllUY of
cpharmollY in its manifold phl.scs is urp,ently requin·d. Itl:! vigorollK PI'(,tI(,·
cution should yield a ri('h hn.l"Vcst of ohsC'rvatiollS, to 1)(' l'x.tmilll'd ill tbe
light of exprrilll('ntul evolution.
XL LITERATURE ('I'rJlJD.*
Annstronp; ••J. B. 1881. .t A Bynopsis of thr N('\~ Zl'I\lllnd HPl'I')('h of
VerolUca Linn., with Notes 011 New Sperics." Trl\Jls. N.Z. IIIS1., vnl. 13,
p.344.
Balfour, I. B. 1879. "The Colll'etions from Rodriqut'z- Boto.m·." Phil.
Trans. RoS., vol. 168, p. 302.
Bitter, G. 1911. .. Di!.' Gattullg .1CQ.Pl1a." Stui.tg..lft.
BJaringbcm, L. H107. ~< JI[ut.1tion I'L 'rrl\umatir,ml'." i>lIr·".
Huehmmll, J. 187(). Intmdurtory Rt'nmrktl to .. Lillt of PIIlU1h fount!
in the Nortllcrn DiKtril't of tltl' Pl"OVilll'O of Altl·kIIUIIl." TruIlK. N.Z.
rnst., vol. 2, p. 239.
--- 1871. "011 IIfnm' Now SpI'I'it,tI nlld Variot.il'S or NI'W 7"('ul.llul
PhUllfl." TranK. N.Z. rIlKt., vol. :J, p. 20R.
UuruH, <t p. 1\)11. •• lMaphi,' (Iondii iOIlK in 1'('111, BOilS of Houtill'l'Il
Mil·hignn." Jlot. Gaz., vol. 1i2, p. 101i.
OhCI'SI'l1I1Ul, T. F. lS!l1. .• .]i'uri hl,t Not('s 011 thl' ,]'hrl'I' Kingll IHlandK."
Trnnll. N.Z. Iust., vo]. 23, p. 408.
11106. .. Mununlllf th(, Nl'W Zl.'aland Flol"l\." Wl'IlillAtoll.
- - 1907. "(I.ontrilJUtic)I1s to n [j'uller Knowll'dge of thl' Flora nf
N('w Zoaland." TrSlltl. N.Z. [Ubt., vol. 39, p. 439.
-- --- - 1908. Tbid., No.2. Trans. N.Z. Inst., vol. 40, p. 270.
-- 1909." On th(' SystematiC' Botany of the lslu.nus to the South of
New Zl'Bland." The Subaut.. Islands of N.Z., vol. 2, p. 389.
('hilton, C. 1884.. Thf' Distribution of TClTl'Stial ('ruslact'I.I:' N.Z.
tt
• Works consuitro. but not r('furrt'd to in the- W'xt. "1'(' not inoludt'd, t'xoopt In a r,..w
Jllllta.noell.
COOKAYNE.-Ecologuat S'Vtut/(~B in EIJol,tftolt :1:9
('ockaync, L. 1901. "An Inquiry into the Seedling Forms of New Zea-
land Phancrogams and their Development, Part IV." Trans. N.Z.
Inst., vol. 33, p. 265.
- _ - 1902. .. A Short Account of the Plant Covering of (,hatham
Island." Trans. N.Z. Inst., vol. 34, p. 243.
1904. "A Botanical Excursion during Midwinter to tlte Southern
Islands of New Zealand." Trans. N.Z. Inst., vol. 36, p. 225.
- - - 1907. "NotE' on tht" Behaviour in Cultivation of a Chatham Island
Form of Oopr08n1« propinfJUa." Trans. N.Z. Inst., vol. 39, p. 378.
- - 1907A. "On the Sudden Appearance of a New CharactE'r in an
Individual of LeptoBper'TJtUm 8coparium" New PhytoL, vol. 6, p. 4:3.
1908. "R('port on 0. Botanical Survey of the Waipoua Kauri
Forest." Wellington.
1909. "Report on a Botanical Survey of Stewart Island." WE'I-
hngton.
- - - 1909A. Tho Ecological Botany of the SubantarctIc Islands of
Ncw Zealand." The Subant. Islands of N.Z., vol. 1, p. 182.
- - 1910. .. On a Non-flowermg New Zealand Species of Rubus."
Trms. N.Z. Inst., vol. 42, p. 325.
- - 1911. .. On thE' Peopling by Plants of the Subalpine RIver-bed
of the Rakaia (Southern Alps of New Zcaland)." Tra.ns. Bot. Soc.
Edinb., vol. 24, p. ]04.
Cook, O. F. 1907. "Aspects of KinetiC Evolution." Proc. Wash. Acad.
Sci., vol. 8, p. 197.
Costantin, J. l898. "Les Vegotaux et les Milieux Coslluques." Paris.
Cross, B. D. 1910. "Observations on some New Zel:l.l<Lnd Halophytes."
Trans. N.Z. Inst., vol. 42, p. 545.
Darwin, C. 1899. "The Origin of Speoies.' London. (6th 00..)
- - 1905. ., The Variation of Animals and Plants under Domestica-
tion." (Popular edition, edited by Francis Darwin.)
D.:l.lMn, 11'. 1908. PrMidontial .Address. Rep. Brit. Assoc.
Dendy, A. 1902. "'rhe Ohatham Islands: a Htllth- III .BIolooY." :r.If'm.
and Proc. Manch. Lit. and Phil. Soc., vol. 46, pt. n.
-- . - 1903. "The Nature of Heredity." Rl'p. R.M. A.A.S., vol. 1.
Diels, L. 1906. "Jugcndformen und Bliitenrc·lfe im Pflanzenreieh."
florlin.
Goebel, K. 1889-93. .. Pflallzenbiologisuho Schilderungell." Marburg.
- - 1900-5. "Organography of Plants." Oxford.
- - - - 1908. .. Einleitung in die Experimentelle der P&nzon." LeipZig
and Berlin.
Griffen, E. M. 1908. .. The Development of /:Iomo New Zeamnd Conder
Leaves with Reg&.ro to Transfusion Tissue and to Auaptution to En-
,ironment." Trans. N.Z. !nat., vol. 40, p. 43.
Guppy, H. B. 1907. "Plant-distribution from an Old Standpoillt."
Author's copy of paper read before the VIet. Inst
Hall, H. M. 1910. "Studies in Ornamental Trees and Khruhs." Umv.
of Cal. Pub. in Bot., vol. 4, p. 1.
Haswell, W. A. 1891. .. Recent Biological Theories.' Rep. A.A.A.H ..
vol. 3, p. 173.
fTenslow, G. 1895. .. The Orig;n of Plant ~truutures." London.
- - 1908. "The Heredity of Acquired Characi;(>rs in Plants."
London.
50 T I "flMltt IOUb.
Tms paper is divided into three sections, the first containing the usual
general plant-habitats, the second devoted to the neighbourhood of the
Franz Josef Glacier, and the third to the plants of the Omeroa Saddle.
My list of plants in Dr. J. M. Bell's report on the Franz Josef Glacier
is, as is therein indicated, most incomplete. With the addition of the
species here published and those recorded only for the Omeroa Saddle the
total is increased from 287 to 356, while a fair idea may be gained of the
vascular flora of that part of Westland from the sea-shore to an altitude
of 1,200 m. Doubtless there are still many mountain species not re-
corded, since, so far as the higher land was concerned, I was only able
to visit the fell-field, &c., on the right-hand side of No-go Creek, where
the slopes are very steep and much broken, and the vegetation merely in
patches.
The Omeroa Saddle is situated on a spur which is crossed by the bridle-
track leading from the Franz Josef to the Fox Glacier. Its altitude is
about 330 m. There is a small amount of open boggy ground, but the
bulk of the vegetation is forest. The occurrence of certain subalpine
shrubs is remarkable for so Iowan altitude, but it must be borne in mind
that every Westland lowland river-bed contains a percentage of plants
which are generally to be found only at a higher altitude.
To Mesars. W. Willcox, D. L. Poppelwell, W. Wilson, and C. Foweraker
I am much indebted for various specimens mentioned below. Mr. Poppel-
well further has sent me full lists of his collections on the Garvie, Eyre,
and other mountains, but these are so extensive and important that it is
better that he himself should publish them.
Am'. III. S,ll/it' XO(I'H 1m Ihl' BlJlooll oJ 1111' SI'I'lI.JIt'I· il.flllmlaitlH, ,0'/1,/1 fl
T-illt of Ihl' 8pt'Oif'1I (loll.rctl'd.
B) R. M. I,.UNO, B.S,·.
I Hi'I"/ 111'/011 II" 1'llIlo'lOpMrlll In~flIlljp IJj (illll/prlmfli. 18t NouemlH'r, ISiI 1.1
ROUTK.
IN December-JI\IllULry, UHO-H. w~ arru.uged a tUlmll party· to take pank-
horses and asc~nd the headwaters o( the Waiau IUld the 01arenM Ri\'"ortl.
We left Hanmer, and went I>~. way of Jark's Pass I~n(l Fow]('J"s 11as8 to
the out-st.'ltion on the Ada. We campl'd near thl' foo1, of tht' saddli·, nnd
explor('d thl' surrounding rOlUltrr I>otnnicall~·. 0111' n('xt camp was ill
Glacier Gully. 1\ tllllUll tribuml'Y uf the Waiau, tlomt' nw milefl farth~r to
the east. 'Phenrt' we crossed Maling's Pass to Lnkl' Tellnyson, in tht'
neighbourhood of which several days were Bpent. Bad weather, unfor-
turmtely, pr('vontrd the !\Scont of allY o( thu highlll' ponks. The highl'.st
point nttain~d was probably under 6.()OO ft., on Mount PrinCI'BB. Rome
of the upper !llpinl' plants may, thert,foro, have escaped oUsol'VlLtion. Thl)
return 10 Hllnmel' \\'IIS made via tho Olu1'('n('1' Vallay.
HIHTORI(,AL.
Tho Speruwr MOWltainti form a little-known district of tho Southern
Alps, lying at the headwatel'B of tho Wuiau. ClaI'enee, a.nd the Wairau.
The district was first explorE'ld by Mr. W. T. L. Tmvcrs during the end ()f
February and the beginning of Mlm'h. 1860. An I\roount of this explora-
tion will bo found in the Nelson E:tamNnR1 of the 14th Marrh, 181)(). During
the trip hI' named "thl' Sp<'llsor Mountains in honollr of the poet of that
namt'." The name is 11011" Oftl'lI misspelt ., Hpcnccr." Maling's PU.SII is
so designated iu honour of Mr. 0. Malin~, who "cl(lOmpaniHd Travel'll, und
who had 80nu thl' pUlIS on n proviolls trip with Mr. Domet,t,. Maling'K
PI\BB 11'.a<iH frolll the wM.ol'llhGd of t,ho Oh,renc:o into thlLt of tho Wll.iau.
'l'ributaril'fI (If 1,ho W"iall WI'I'll lllLnlt'U lIy Mr. Travors nftAr his chilclrlln
-(.ho Ada, tl\<, Honry, and tho Anno. Oc/.m'sia TrCWCfrsii wnll originally
tiisC'overc>.d on ti1(' summit of j,he mountain betwet'll tho A<hL nnd t.ho Anile.
()tll<'r novc)til·t! ditlcovert.'d by Trave1'll in the dist1'iot W(II'Q Roou'llOulw
"I'iIJvtM!oliWl, R. lit/aTli, VILI'. TrtWt'rMi, R. Hi'llOlairii, PiU08ptlrtlhn. patulum,
(htapMZium nitiduillm, ImtI Walllcmbtlrgia ClJrtilagiftea. Of those. G'M'p'halimn.
mtidtaZtwn. hl\s not, again been found, unloss, u.s I'ppoars likely (8<'1' subjoinod
list), it is ImlongKt, thtl specimens collected by us.
Since tho time of Tmvers tho district has apparently BOvero.l timos
been visited by collect,ora and botanists. However, there is no published
aooount of its botany, a.nd the only list olspeeies drawn up for it is a short
one appearing at the end of an artido on the asoent of Mount Franklin by
Park (Trans. N.Z. Inst.• vol. 18, p.350). This contains seventy-eight speoies,
idontified b~' Buchn.nan. Of these, about a :I:ifth were not collected by us,
but it has not ueen thought advisable to include them in our lillt. Indl't'd.
it appears to us that several of those there recorded are most unlikely
inhabitants of the district-e.g., DodO'n(J('(I V18008a, Gentiaw, oon0i7ma, G.
S(U1;()8a, Dracophyll'l.l.1n U'I'villeanufIL (typical form), Veronica otWra, RatI'I.I.n-
oulus pVng'l.l.is. Various species. also, which we had t'xpected to ~et were not
found by us. Amongst these may be noted Oelm:isia Traversii, of which,
however, we SllW specimens from Mount Percival, at the back of Hanmer,
and Ram.moulus Itgallii, which we did not se~ at all, though we were
assured that it grows in the district. It is quite clcar that neither of these
species are common in the Spenser Mountains.
There are several reports on the geology of the district. References
to it will be found in Haast's " Report of a Topographical and Geographical
Exploration of the Western Districts of the Nelson Province," 1861. He
visi.ted the Buller and Grey Valley in 1860, and saw the Spenser Mountains
from their western sides. He speaks of .. the high mountain-('hain, called
by my friend Mr. Trnvers the Spencer [sic] Mountains, whose highest peak,
clad. with etel'nal snow, rose grandly above the low hills in front of it. I
Mmed this mountain, Mount Franklin, in honour of th~ late Sir John
Franklin."
In the ., Reports of Geological Explorations during 1888-89" (Wel-
lington, 1890) there is an article by Mr. A. McKay on the .. Geology
of Marlborough and the Amuri District of Nelson," which describes the
geology of the eastern slopes of thf' Spenser llountains (throughout the
report spelt " Spenl'f'r ").
TOPOGRAPHICAL.
The Spensol' Mountains are some twenty-five miles in length, aud are
generally rcgllordcd as lying between the saddle of the Ada (3,300 ft.) and
.Mount Franklin (7,671 ft.). The peaks are of a nearly uniform height of
7,000 ft., with a.n upward tendcncy towards Mount Franklin. The height
of 10,000 ft. allotted to Mount Franklin by some of the earlier explorers
was an error, doubt,less du(' to its t'xtellsive snowfields and alpine magnifi-
cence. The Waiau. Olo.l'(,11('e. Ilud Wairau all converge upon this peak, and,
indeed, their chillf SOU1'l'CS lie upon it. To tho south arc Mounts Guinevere,
Aeneid (7.050 ft.), Pl'incf'SS (6,973 {t.), Una (7,1).1.0 ft.), Imd Faerie Queene
(7,332 ft.). The Telll1ysonian names are duo to Governor Weld.* In the
valley of the:" Wailm lies L'1.kc Guyon, and in thl:Lt of the Olaronce Lake
Tennyson. Both urI' glacier lakes, due to thl' hanking-up of the waters
by morainil' dcpositA. Indeed, thl' wholo ('ountry gives evidence of
having at 0110 tim<l boo11 heavily glaciated. Tho Ado. Stream runs
through n wide glacial valley, and therl' has been a large terminal
moraine aoross the Waiau about 0. mill' and a half below its junction with
the Ada. The head of the low saddle lied also in a fiat open valley, a.bout
200 yards wide, bAving at its highcst portion a Sphagnum bog. This valley
shows no terracing. Opposite its mouth there are a number of parallel
lines extending up to !l.bout 800 ft. on the left ba.nk of the Waiau. These
arc perhaps lines of ~lacial pressure. Glacier Gully has doubtless at one
time carried a sel'ondllr~' fl;lal'ier, but now it ('an scarcely be regarded as
true to name. It opens out at its heud into 110 large cirque on the ftanks of
Mount Una, and the bed of the stream (about 3,300 ft. altit.ude) contains
fragments of melting Nve about 10 ft. thick. Judging by l~ppC'arances.
this neve would scarcely last through t.he summer. The low('st portion
was detached from tho rost, and WaR :.Lbout 100 yu.Tds long and 25 yards
wide. It was bisoctod by tho strea.m. A little clear icc was visibll' at thl'
foot of the neve. .A pronounced hanging valley on tho J'ight HI t.lw stream
and i,he remnants or a lateral moraine shOWl'd tbat at 011(' timl' u. glacier
.){ respectable dimensions had filled the creek-bed. Tho valley is an open
une, about 100 yards wide.
Thl' country becomes progressively drier as we go oastward. from the
Waiau to the Wairall Valley. There are few shingle-slips to be seen OJl
MOIDlt Faerie Queen. as looked at from the Ada Valley. Those on Mount
Una, as viewed from Glacier Valley. arc a little more extensivo, but in the
neighbourhood of Lake Tennyson they become more numerous and O('('up:v
a larger area. The Wairau Valley, from a saddle abovl:' Lake Tennyson,
appeared as dry a.s the region in the neighbourhood of Mount Arrowsmitll.
Doubtless the westerly rains pass over the saddll' into the fl'rtil(' .Ada VI~lloy
and Stanley Vale, but are llnable to pelletrate to thl' l'ollntry ut i,ll1' head-
waters of the Wairau.
The greatl'r denudl\tioll in thl' Waiau Valley prl'vl'nts thc' Iwcumll!atioll
(If shingle-slips there.
VEGETATION.
In the ausene<> of Dlctcol'ologi('al statistics, it is, of COUI'80, impossible
to show directly how climatic conditions are affoct4ng the v6glltation, and
indirect evidence only is available. Thc conditions in the d.istri('t, however,
resemble those that prevail in the Mount .Arrowsmith region, as the plant
formations are very similar, and a large number of species are ('ommon to
both distl'icts. Indeed, the general description given of the plant fonna-
tions in thE" Arrowsmith district* would apply to this with but few modifi-
cations. Rock, river-fan. river-bed. tussock steppe, bog, la.k<:,. forest, fell·
field, and shingle-slip pl'<.'8f'nt similar features and similar plant·ussociations
ill both districts. Certo.in suba.ssociations of the .Arrowsmith district wele,
however. not notieed in the Spenser Mountainll. Dwarf O(JNlltiolmeZias were
ohserved only in the Waiau Riv(lr bed near Hanm(lr, and thore only a few
plllnta of an unidentifiE'd spC'cics. ThE' lU'rompanying Bpedes of the Moullt
Arrowsmith wstri(lt- (I.fl. V(J7'onica pinlt61eoiileB Vilr. minor and MfJl'hlM-
iJfrM epkNlrm.·de8-wllrc not, observed in tIll' more northern arca, nor did
W(\ se(' in the SpC'.tlflcr MOlmtains lilly such forest as tho subalpine totara
forest of thn Opper Rakaia Valley .• Indeed, forest-trees of tmy kind, with
the exception 01 spooirs of N otkofagus, N otlttYpamJfD. Gaga, IIond PiU08'POf"Utn..
were completely abBl·nt from the Spenser Mountains. 'rho complete a.bsence
of I'ny of the species forming the usual coastal forests of New Zealand is
perhaps the most remarkable feature of the district. The subaJpine scrub
is also poorly represented both in quantity and number of species. River
steppe. fell-field, shingle-slip, and rock occupy nine-tenths of the district_
The N otkofagus forest was found only in the river-valleys, and deoreasllCi
in quantity from west to east. Only a few acres are to be seen in the
Upper Olarenco Valley. close to the sjdes of Lake Tennyson. The upper
portion of the .Ada Valley, however, contains considerable quantities of
the forest, and has contained more; but some has been destroyed by fire
and some out out for timber. A fuller description of this forest and of
the Sphagnwm bog on the Ada Saddle are given, as they differ ('onsider-
ably in composition from the similar associations observed in thf' Arrow-
smith district.
PHYSIOGNOMIC CHANGES.
The district has been in occupation by runholders almost sincE' ita
discovery, and as a result many changes have been effeoted in the general
composition of its vegetation. On the river-flats of the Ada and Clarence
Valleys English pasture grasses have been sown, and flourish luxuriantly.
displacing to a large extent the native plant covering. The lower portion
of thE' Ada Valley contains beautiful pasturage of cocksfoot and white
clover, with here and there a considerable admix.ture of Yorkshire fog.
Occasional patches of Acaena miarophylla, A. SMlguisorbae, Asperula perpu-
silla, OrefYffI9j'l"1'l/f's antUoola, Ootula dioica, O. squalidtJ, and specimens of
StaokhO'UB'ia mWnw. Rammculus Joliosus, &c., ocour in the midst of thc
pasture. In the stonier portions it is crossed by lines and thickets of
DiBCOIf"i,a semb, which rises to a height of 15 ft. to 20 ft., intermingled with
occasional specimens of Ooprosma propinqua and Veronica C'Ulpressoides.
This pasture in the Ada Valley passes at its upper margin into Yotl!o/agw
forest. The original tussock steppe and the forest-area has been much
altered by burning. Severe burns have evidently taken place from time
to time, and much of the southern beech* is second growth, with the stumps
of the older and heavier trees still standing above it. Sorrel is rapidly
gaining gl'Ound in many places, and is even invading the shingle-slips.
Above the bush is Dantlwwia steppe, which has a.Iso in some places
been subjected to severe burning. The fell-fields, too, have suffered occa-
sionally from this cause, and new shingle-slips have sometimes formed
where the old vegetation has been burnt out. It is difficult, however, to
say whether consolidation from shingle-slip to fell-fields is not taking place
at an equal or greater rate in neighbouring 10calitiE's.
• I am using. a.t Dr. CoCkayne'1I suggestion, the term .. southem beech .. (NoIlIofagu)
to distinguish our fo1'8lltll from tho beeCfi (Fllgu.r) fol.'ellta of the Northem He-millphere.
Tf'orttrartWnI .
into the forest thc mtroduced grasst's disappear, tho lo)"('st-fiool" hct'onlt's
damper, and moss appears on it. Th(' husli ill still fairly OpOll; distan('('s
of 10ft. i.o 12ft. often occur bI'tw(lI'n Tlt'l!!,hbouring trOt's. Horn III'C' putc'hclfI
covered with a C'lIlp<'t of moss. wi1h YOlmg h('<,C'h<,s growiJU.\ 1hrongh ,
IJlt'('l~"""'1! IX'lllIa marU/I1, ('Oprll~III" Hlm/(lt~~a, .111<1 C)('C'ItSiOlllll plllnts of
C. v1trI'R('('fIII? now app<'nT, wi11L h('1'c' lind Ol<'r(' 1\ YOllll!!, plnnt of N(lt1l.o-
P(f,fI,(J,X arborl'lItn. Through nll tllill poniol! of till' fOl'PHl (':~111(' haY!' hO('l1
rUlUliuj.!,.
In paflSiuj.( mto th<' portions Clf th(' fon'st whi('h bio('k Imvt' ilot
destJ'oY('d, th(' undNgrowtll b('c'onl(,s d('ns('l', hut still oonllis1s ()f b<'ech(·s
in all stag('s of dev<,lopm(·nt. Oorylsa,lth('s tril()ba, .AIll'flocMlu8 Ijracllill,
and O/~ilogl(lftis corfl,ufa arc now to be found. AI. we ILBcl'nd hom the
Tiver-ftat. which is WE'll coV<'rcd with soil, ihl.' torl.'st now hl'comllll
stonier; the trees bocom(' largt·r, out mBlly of tIl<' lurj.!,('r Ofl('fI
(1 ft. to 2 ft. in diamctE'r) haw fallell from lIonll' ulldch')'milll'd l'lmll('.
About ,I quarter of It milc' fronl th(· lllflJ'gin II hund of NofJwj(Jytls
Mmziesfi is found, while Oc'('usionai plllllts of ~(j('fl,('Cf() II('llir//(I/t/('S tl.IlpSnT
on the mossy :6.001', Vmmicll 1Jl'f'ni('(ls(£ llCconws 0101'(' ulmlltll~llt, alld II f('w
sppcimclls 0'1 ('O'PT08flla lifiariifCllia ILppco.r. Ht'rl' the t'dgl' oj ,~ llusll-
"reek is IJ'ing('d with MflellletWl'C'kill WGlllMis, Al"Owillg illto Ion!!; ()V('rhallging
flPra.ys, giving th(' plant II v('r~' difil'reut ILppt'lmlllC(' from its riv(·r-bro
form. NOfhojat.JU8 jf.l.8(:a, in ('lumps, is found higll('r up 1.110 riwr-fl.nt; but,
neithE'r it nor N. Menziesi, ultug(·ther r<,ph~l'l' th(' N. oliDorliniliRs, wluc·h
prohably forms the grea.ter hulk of th(' forest, on to Ittl uppl'r nungill. TJli"
at least, wtlofl the cas(' in Olaoi('1' Valley, wh('re it l'ltss('s up int!) 0011a
Lyallii.. but thp upp<."r limit of tbl' forest was not examined in the .Ada
Valley. This may hi' placed at 800 ft. to 1,000 ft. above tho bed of the
v~ney, and the sides arc so st('('p that they arc often sw('pt by I~valan('ht's
of stones, which carry away th(' beech-trees. Their phlce is taken by
plants of the shingle-fan. I\nd thE' beeches grow into this vel!,otlltioll Irlllll
thplsidcs. and pl'obably in ('out'lle of timl.' will 1'('O('('upy t.}\(' whole arl·U.
'rll(' uPller porti()J1 of j,h(' Adu Saddll' ill a Hnt 0P('u VILU('y about :100
yllrds wido, with English }l1.atllTILgt· (oocksroot, Yorksllirc' fog, ('lClwr, I:I<lrr('J,
lllllKk, &c.) und BOutl\<."Dl be('c,lI (Nutho/agUII olitfo'l'tioitil's) forI'st (If II Himilar
tY}I(' to t,hllt, desc'rilwd in th(' Ada Vall<,y lIomc' BOO ft. b<'low, AriHt<Jlc,{ill
/n/'1008(1, Azof('lia trijolio{atfl, and AC(Xt'fU6 Sat&g1lill(lf'ilar ()c'c'ur 011 tho forc'st-
tI(lor, 1~lonA witJl man.,- of fh(' plants "ITOILtly ml·nti()l1od.
'rh(' 1\(':1<1 of t,h(' IItLcldl(' is ()('cupi('d hy a 8pllagwufII hOA ahout ~I()
yards squaro, which divide'S thc' ('"stem aud w('stel'l1 WI\wrsh('ds. A slnlLll
pond a.ppears in the C'elltTl' of tb(' bog. TIl(> t'clgel:l ItT(' fringed with
Dracwphgllum 'UfI,';'{lon/tll, P()rl~arpu8 'lVivali8, PhyllocladUII alpitws, a.nd
DfIhf'Yditunn Ridwillii. The bog is evidontly risillp', as in many places thp
DracophyUut'll is being buried. ThE' Sphagnum is dotted with cushions of
Oreobolfl8 pectinatw Illid O. strictus, and clumpo 0:( (!elmiBia longilolia var.
alpina. Round thE' edges ar(' Ourisia maoropkyUa, OI'I'l1i81", oori.aclta and
O. 8pectabiUs, HelicMy8um beUitlioide8, MitYroseris ForstDri, 8enecio lagopus,
8~a tt.itens, and Pratia cmgulal.a. Other spE'cics growing in some
quantity on tho hog art' OalfJd.etm·o bi.loUa, Rost~'ooia gracilis, B~
(JflNliflg'lw.mii, 8ohoeM68 p_flqrus, 00lft'a) rteUulatfJ, O. GOAJt!i<'1wudirmt',
('a,.~.;,fI,(' hetef'otp1lylla, For8ft!f'Q BiilfltilTfi, and DrOllt"l'o, arct1Wi.
'_AINt..- -Botany of the Spenser .lfoltlttaim.
I appt'nd sOIn!.' notes 011 forms of special interest, and Q li&t of species
gathered. •
FLORISTIC' NOTES.
1. Gunnera densiflora (!) Hook. f.
There occurs in the forest of southern beech on the western side of
Lake Tennyson and on the margin of the lake a species of (]u",,,,era, which
is probably the imperfectly described G. ilen8iflora Hook f. t The plant
grows in C'onsiderable abundance on a sloping bank at the water's edge.
It agrees fairly well in character with the description of G. deft8i{lora.
The following is a fuller description:-
GUfllTl,Cfa sp., with short succulent rhizome, herbaceous, creeping, root-
ing at the nodes, with rather stout villous stolons, 7 cm. to 10 cm. long.
Leaves clustered at the nodes. Petioles stout, with rather short silky
ho.loirs, somewhat :flattened, 4 cm. to 6 cm. long. Petioles stout, with rather
short silky hairs, somewhat :flattened, 4: em. to 6 em. long, straight or
recurvC'd. Blades cordate, reniform to orbicular, 2 em. to' 3 em. wide,
glabrous or with a. few white or brown hairs on the margin and midrib,
sharply dentate to dentate-sinuate, auricles bent upwards towards the
upper surfuce of the leaf, though not appressed to it, veins distinot,
paImate. Flowers monoecious, in simple spikes, which are usually uni-
sexual, but occasionally a. few female :flowers are found at the base of
the male spike. Male spike 4: em. to 7 em. leng, ascending from the axil
of the leaf, the peduncle usually shorter than the fertile portion, and
covered with a soft woolly pubescence. Flowers shortly pedicella.te, pro-
vided with a minute linear acute bract, 2 mm. to 3 mm. long, arising
from the base of the pedicel, and 2 minute linear sepals smaller than
the bract. Petals 2, linear-spathulate, larger than the bracts, hooded with
blackish scarious tips, S mm. to 5 mm. long. Stamens 2 or S, :filaments
shorter than the anthers, anthers ellipsoid, 2-colled. Female :flowers sessile,
densely crowded with bract, as in the male, calyx-lobes 2, deltoid, apiculate,
pl'tals wanting, fruiting-spike elongating to S em. to 5 em., and covered with
a \ .llous brownish pubescence. Fertile drupes 2 mm. to 3 mm. long, rather
spa.rse, pyriform, seBSile or shortly pedit'ellate, C'rowned hy the persistent
calyx-lobes.
Mr. Cheeseman has drawn my a.ttention to the {act that the plant is
ptlrhaps identical with Tasmanian G. cordi/alia Hook f. (Benth., Fl. Austral.,
2491; M,'Uiganiat ooriUloUa Hook. f. in Ie. Pla.nt., t. 299). This is more
fully dl'scribed by Schindler in his monograph on OW1/hl,era in the Pflan-
zenreich. The description ill the " Icones Plantarum" is very imperfect, and
differs from that given by Schindler in sevt'ral important points. Schindler,
also, has not seen the mature drupes, and his account of them in this key
to the species is not consistent with that given in the specific description.
Doubtless the identity or otherwise of the two species cannot be determined
until a definitt> description of the fruit of the Tasmanian plant is obtained.
H the description of the Tasmanian species in the P:fl.anMnreich is to be
relied on, the drupe is ovoid und costate. The drupe in my plants is
pyriform, rounded, and without (·ostae. My descrjption was drawn up
on the spot from fresh specimens, of which there was abundance.
• M1 thanb are due to Dr. Oookayne and :MesirrN. Cheeseman and Petrie for much
kmd aHIIlBtano(', freely given. in the ldentifioa.tion of thp HpeOil'llo
t HAIldbook N.Z. Flora. p. 68.
3-Tr.mo.
'136
piDlLlloe teruately divided, upper ternately lobed, the whole leaf deltoid-
ovatiC in outline. Petiole as long or longer than the blade, terete, strait,
with short broad mmnbranous sheath, 6 mm. broad at the base. LoWOl
petiolules 10 mm. to 15 mm. long, upper shorter, ultimate divisions broadly
cuneate, flabellate, dentate, teeth subacute not piliferous. Flowering-
stems, sovel'al, compound, much exceeding the leaves. Peduncles with
1 or 2 bracts, the lower, if present, one-third of the distance from the base,
usually with 3 linear lobes and a "broad clasping base, upper bract set
midway on the poduncle and smaller. Umbels compound, primary rays
2--3, with a simple linear bract at the bas\! of each ray. Secondary rays
5-7, somewhat unequal. pedicels rathel longer than the fruit, 1'5 mm. to.
2'5 mm. Fruit surrounded with an involucre of small linear acute bracts.
Head heterogamous, the central florets generally male. the ouwr her-
maphrodite. Calyx of I') lobes, teeth dc'ltoid. acute, minute. Petals white.
Styles subulato. equl1,lling tho ovary in length. slightly rc('ul'ved. fl'uit ellip-
t:loid, carpels with 5 equal ridges.
011 the upper riwr steppe, Mo\ult Princt'ss, above Lak(· TC'llnysuJI :
nitit ude, about 3,000 ft.
A specimen is dC'posited in the Cnnterbury Museum.
4. Myosotis Laingii Oheeseman.
A hithC'l-to-uudC'lIcribl'cl spE'ci('s from Lake Tennysoll. now Milled br
t.'hccscmall.
5. Haastia pulvinaris VIlT. minor Laing (val'. 110\'.).
In omnibus partihus minor quam typllS: rami unaculll toliis 6 mm.
nsq11c' ad 15 nun. IlLt,i. Pappi capilli nOli supra incrassati, magis autem
sC'..abricii, pene nmbriati.
Two distinct. forms of this spccietl .'ppear 011 Mount Princess (alt .•
5,OOOft. to 6.000ft.). The smaller fornl is apparently distinguished by the
pappus ha.irs being scabrid, almost fimbriate at the tips, and not thickened
I1.S in the typical fornl. The tomentum of this variety in the specimem
we got is much whiter tllan that in the normal variety.
A specimen is d('positc'(l ill the Cantc.rbUl'Y Museum.
6. Clemisia petiolata Hook. Val'. membranacea Kirk.
'[ wo {(lrms of this vltril'ty wero ohsC'rved, 0110 with the lea! nearly
glabrous on hoth surfaCC's, and tho other with margins covered with & ful-
VOUIl PUbCBl'On(·t'. The I('IWl'lI in tho glabrous form are often cordate at
the base, and 1m' hl'Ol\del' und shorter than ill t,he form with marginal
pubesccllc('.
FAll-fiold, Glacior Gully; about 4,000 ft.
7. Gnaphalium nitidulum Hook. f.
Specimens of a species of OMphaliu"" were collected on the upper river
steppe of the Clarence Valley, at an altitude between 8,000 ft. and 4,000 ft.
It appeared to me to agree fairly well with G. mtidulum in its characters, a
plant that has not been collected sinco first obtained by Travers :fifty yf'ars
ago. I sent a specimen of it to Mr. Cheeseman, who thus l'eports upon it : -
.. No. 1760. GMpktitium sp.-The female florets are many times more
numerous than the hermaphrodite, and the pappus hairs are very numerous,
scabrid at tho bue, and the aohenes u.re faintly downy. These chamotera
place tho plant in the: genus ''ih41'llalium. The lea.ves a.1'O about i in. long,
8'"
6t! l.'ra",,,actIOla!
·The Kbw l1.uthoritiCII now (March, 1912) I't'JII'Iri tho specImen u,rwa.l'Ilpd to be-
identioa,l with the typI. of Q. "uitltiNm.
I.AINfL- Botnn11 of the 8peft,er llountl7l""
Cyperaceae.
l!:laeochaft8 Vwnninqham&1 Boeek.
OarpM alpiM H. HI.
8ckol'/ll'U8 paUciff,m"U8 Rook. {.
OreobolUR pectiMtUR Hook. f.
strict'll8 Berggr.
UflCinia tmeinata (L. 1.) Kiikenth.
jusco-vaqinata Kiikenth.
rubra Boott.
lepfostaohya Raoul.
C'arn stellttlata Good.
Gaudichaudiana Kunth.
temaria Forst. f.
Raoulii Boott.
Petriei Cheest>m.
luoida Boott.
t"Bfacea Sol. A ycrv aberrant totm, or new (D. Petri!')'
m
.')irwlairii 0. B. Cl.a.rke. Immature.
Roatkovla gracilis Hook. f.
LlZUla camppstris D.C.
JU'1UJU8 f!ntl(l('-'U'ltmdiae Hook. t.
Naiadaceae.
PoUWllugetoo 01wt>lIf!1t11anii A. Bl'un.
Liliaceae.
AslRl,a montana (T. Kirk) CockaynE'.
Phtmniu,,~ (Joo":itmllm IJe .Tolis.
('hrysobactron lll)o/(('ri 001.
Orchidaceae.
Thl'lY'tlltm lonIJI/oli(l Forst. t.
Mictotill u'lli/olia (Forst. f.) Reichrnb.
PrMopklllll.Wtl Golensoi Hook. f.
PttrUBtylis Bcmksii R. Br.
Valadt'nia LlIaUii 11ook. f.
hi/olia Hook. f.
ChiZoglouis comuta Hook. f.
Adenockilu8 (fl'a(yjUs Hook. f.
CorgBanlk"B triloba Hook. f.
Gastrodia OuMinqhamii Hook. t.
Fagaceae.
Nothojagus oUtJortwides (Hook. f.) Oerst.
/USCCl (Hook. f.) Oerst.
MfflZie8ii (Hook. f.) Oerst.
70 /1( "' ((flOl/l
Urtlcaceae
Thllt" /(lI.'a POIT
Lorantl'aceae
Ellftllln.lll£ tttrnp(IIII(' (F01~1 I) Ellul
{ttll'1I7,! (Hoo!' t) Engl
Santalaceae
Exat'ul pu, RI(IUl~llfl lIoo!.
Portulacaceae
(llaytoYlw IIIu,tlula"C(I Huo). I
1.11ontw lon.tana L
Caryophyllaceae
BtellaulJ Ilouqkn Hook I
Oolohallf/lus Bdlald'l."." Fpnzl \ II fllpl'IIII' l' KJlk
aotCUlat II> Hook f
Bc.lRr«lfltllu\ IliflOlIl~ Hook f
RanuJ1l.ulace-ae
('kIm!'" au\/lah, T Kuk
Ran.ufu ,!III'I .,'SUjfUb Hook i
Monro, Book t
MonrO? Hook f Vell tkntutu, T. K'll
Bmclmt1/& (l) Hook t IdE'lltJilfl hun not epriam In thE' ,\b"l'l1l'l> of
flowers clIld fllnt
..p S('orc.dy poBSlbl(l to IdpntJiy In the Ih.,(nl.t til flOWI'I .. dond lrult
/urtUB Bank.. &; Sol
lappaoeU6 bnuth
JO/WU8 T Kirk.
l'tWimt8 Bonks &; !:!ol
CketN'MOOI'6 T KuL.. TIll'> I'> pu}taps ouly do h)t!l!Iphytl( lUI iii of
R loko6U8 It IS evelY\\ IU.'I< ,a1lUllcl.m1 thlc)Ul!,llont HII' OI'1t 1I 1 ID
pool'1 ,mei '1low-nmnmg 'Wdt€>\
Cruclfelae.
{'arda'NlI!( hftuopkut1n (FoUI1 t) I:ichult.1 Vdl lIIuHllltlia 1-1<111111/.
ht'krnphljlln (Fo1ll1 r) Fkhllltz val UN/flora Hook. t
deprl'~'Q I fook f
JastlQ10lu Hook t
Ent/\II Chc eaeul
Notothlabp'l rosulatum Hook. 1
o.U8tu,l(' Hook £
Droseraceae.
DrObt'ra ard,,1'/, Hook.
Crassulaceae.
Cras8,dtl S11'bmatltJ &hultz.
Plttosporaceae.
PtIiW8JHHllm pntululII Hook t.
Lu" -Boto'fllj of the YpenMI 1l01lllfaml> 7l
Rosaceae.
111lbll6 IlIHt I alu. EO! st j
,thlll1delICY/,dt ~ A Cunn Vdr (%latU8 r KIrk
"lbpaWPi'latlt\ Cotuvn('
P0'fu(lotfltn Suuth
Pote'llt./IO 07lserttIQ L Vcl.l mneHnoule~ (Raoul) T Ku k
4((11/&((' bar&f/Uuolbai' Vahl Ven p1losa T Kn1.
mCI fms Hook f
IM71111b Hook f ViI.) [ont/neapa B)ttof>I
1II,clophylia Hook 1
/ltlblO Buch
l"l~CM\(l(,TI\ (7) Hook 1 non Vahl
Legumlnosae.
f)l/IlIIlthapl,a \ubu/ata T. KIrk
Geramaceae.
(.,l'lanIUm 11II(,lophVll'Ulil Hool.. {
1/10111' L
Oxahdaceae.
(h al'l> colt~uJulala L
fflflQl'llaftlo((' FOlSi
Conanaceae.
t (/1 lalla ,u80l/olta L
thl/t'llt/olta Hum!> & Roup
Ultqust/,8stma Hook f
Stackhouslaceae.
Rhamnaceae
DI \('m 10 toumntou Raoul
Elaeocarpaceae.
Amtotplw IW(lcoso Hook
Malvaceae.
f7tMJfl Ilbi/ulto (F Muell.) «)C,kavn<.
LIfOl'" (Hook f) RaJrf'T
Vloiaceae.
Twla (.lunumgMtlul Hook f
H !JfIlPlulintlu Ta rll'fltato 'R Br Vcl.l G"'lush/olta Benth.
Thymelaeaceae.
Punelea rtaueisli Hook f.
Lyall", Hook f
sl"f1CeO-tI&llosa (forma) Hook t
StItet't (7) T KIrk
Dtapl'ie8 1)1110'10 Cheeseru val mulfA.flora Cheesem
72 7''1'a'~B(U: f I tJllX
Onagraceat'.
Epilobwm pllllidif/orum HoI.
BilltlrdinianulII APT.
ill'''~'lItll 1401.
p"IH''I1/I A. ltil·h.
flumumioulII. IluUKllk."
pit" 11ft! Pl'l rit'.
tl'1liUi1J6S 1100k. 1.
alHitltJitU'lI A. Cunn.
li'M&al'oit/ps Hoole. f.
ll'UmmularijoZilum R. (Junn.
macropus HooI.:.
graci.lipt.'11 (t) T. Kll'k.
cras8W'tI Hook. f.
vemwoSUIII m ChCC8f'1Il.
micropll!Jllum A. Ric,l,.
glabelht7t1 Forst.
fI,(J'I)(lt'-Zl'lamliae Huusllk.
i'his was c'olIcctcd by (lhl'('Bellum ILt Lako
'P'!I(JfI.()8tachYIIIII Hlmllsk.
TC'unyson. hut not c()II('CtC'd by liS.
Halorrhagaceae.
lJalfnrhOl.Jis ikprt'.88a W111p.
(I'~Mf'I'a rtemifl,ora (') Hook. f.t
dffltata T. Kirk.
Araliaceae.
NotkopWflt(Jtl' aroorewm (Forst. f.) Sl'om.
Umbelliferae.
Hydrocotylt.· t&OOaNe/Miliae D. fl.
(J8iatira L.
A"lchiuiletn,II llol(,flhii (Hook. I.) Domiu.
paUidUfIt (T. Kh'k) Domin.
tf'i/Qliolahml (Hook. f.) Oomiu.
nitefl., (pI'i,ri(') Domin.
Ort'omYN'ltil:J anttw(JUr End!. vaLl'. Oo/,"",s(n (Hook. f.) 'r. Kirk.
andi(''(Ila, End1. var. f'OIIfWS" (llook. r.) T. Kirk.
A O'iIphylia (lown/Illi Hook. J.
8quarf'n8(J Fot:At.
MOMOi Hook. I.
AMllotomt' llaa8lii (F. Muoll.) (Joclka.y"C BlId Laing.
fiU!olia (Hook. f.) CoCkU.YII6 1\11d Luing.
~(J (Hook. f.) Cocka.yn(· and LaiJlg.
fWomatica Hook. f.
imbrioata (Hook. f.) LI.ing.
&ysii (T. Kirk) Laing VIU'. t#!tmllBomt.mitlm (Luilll().
piUjtll'(J (Hook. f.) Cocka.ync and Laing.
AflgeUca Gih&gitUum (Forst. f.) Rook. f.
var.;
tkoipitms Hook. f.
• He"' W!Od to include the Now ZoaJa.nd formll of tho lJubantal'tltit' /C. fXItI.terti/olilWl.
t Yide "Floristio NotoR."
~ A wry distinct ah.ingle.s1ip form (01' new 8~), of whioh, howover, only OD.O
Kpeoim(l1l was obtained. It iA therefOl'O not further deaoribecl at preRenl.
r~ArN(1.-Botanll of the SpenRer JirmntaVnI1. 73
Cornaceae.
Ooro'kia OotoneaBter l:taoul.
GriBelima lit/oralia Raoul.
Ericaceae.
Gaultheria antipoalJ ForKL. f.
rupestris R. Br.
Epacridaceae.
pentaclwndrll pumila (Fol'st. f.) R. Br.
Btypll£U,a aoeros(J Sol.
Oolensoi (Hook. f.) Dinls.
/lJ8eWulata Forst. f.
FraseN (A. Cunn.) F. MuoU.
Oracophyllum rosmarini/olium (Forst. f.) R. Br.
uniflorum Hook. f.
Gentianaceae.
GentilJna corymbi/era T. Kirk.
patula (T. Kirk) (Jueescm.
bellidi/olia Hook. I.
dWisa (T. Kirk) Ohooscm. var. 'fM{Jflifica T. Kirk.
Borraginaceae.
My080tis au&tTalis R. Br.
Forster? Lohm.
Travers';'i Hook. f.
laPta Chcosom.
macrantka Hook. f.
T~i Chellsem.
Labiatae.
Mentha O'mMtagMmii (A. Cunn.) Benth.
Scrophularinaceae.
MfaUI radioa7&8 (Hook. f.) Cheosom.
Veronioa subtJlpintl Cock.'lyne.
dWer!lm~ mChu1l8cm.
saliDi/olia FOlK1. r.
tll!'l"lMcl»la Hook. f. var. oanterbtmemis Armstr.
buwi/olia Brnth.
buwi/nUa Bllnth. var. patem Oheoscm.
eup1csHoiar.a nook. f.
epaeridl'a llonk. f.
macrantka Hook. f.
Raoulii Hook. f.
puWinaris Hook. f. & Benth.
lifIi/olia Ilook. f.
catarractae Forst. f. var. ltmooolata Hook. f.
Lyallii Hook. f.
Bitlwilli. Hook.
deoumbem Armstr.
pingui/olia Hook. f.
leiophylla Cheesem.
Gilliesiana T. Kirk.
lycopOtlioitk..a Hook. f.
Haaseii Hook. f.
74 Transactions.
Petrit'.
Lentibulariaceae.
lltriculaNa ml1Xl('-:iwlalldlUc' Hook. f.
Plantaginaceae.
Plantago .Raoul", Decne.
BrO'W'l&ii Rapin.
8pathUlata Hook. t.
lMigera Hook. f.
Rubiaceae.
(to-prm,rna 8e'I'f'Ulata HooJ....
rhamnoides A. Cunn.
paroiflm(!, Hook. £.
ramtlloaa Petrie.
'Direscens m
Petrie.
bruMea (T. Kirk) Coc·kayn(·.
propinqua A. Cunn.
linarii/olia Hook. t.
repeM Hook. f.
Petrie;. Cheetlem.
aalium tenuicaule A.. CWln.
,unbrosum Sol.
Campanulaceae.
Pratia angulata (Forst. i.) Hook. f.
macrodon Hook. f.
Lobelia Roughi;' Hook. f.
forahlntbt'Fl/ia cartilagi'flR(( Hook. l.
,.
Stylidiaceae.
l'hgllaohllR clatJigel'a .F. MuC'11.
Oolensoi Bcrggr.
/t'orstt'rtl BitlwilTii Hook. l.
Compositae.
TAyenClphclfo, pff.wlcUu Hook. t.
Barke71 T. Kirk.
BrOl:hyc0m6 pintlflla Hook. r.
Thomsoni T. Kirk Ysr. mrmbranl/oha (?) 'r. Kirk.
Sinclairii Hook. f.
Olearu& oymbi/olia (Hook. f.) (1h_eesem.
~/oli(, Hook. f.
'fIirgata Hook. f. Forma with ri~id bl'auches and dpiny hl'lllchiett'.
Ot'lmwa Walkeri rr. Kirk.
latmilis Buch.
discolor Hook. t.
lfacana Hook. f.
i,i('ataQ. Hook. f. var. petiolata 1'. Kirk. *
",tMata Hook. f. wr. membranaet'll T. Kirk.
• Vide .. Floristic Notf'~.·'
',AlNG.-Botany of flit SpenBer .1lOllllfn;'nll. 76
AR'r. IV. -Nfltf'1I (19' the Plan'/ Oooo"''''fl 01 Oudfi,HiI iF,lllffd (ifill tilt {lvI/qed
IsltmrlH.
By D. I•. POPPII11.WIilI.L.
C. PLANT-ASSOC'IATIONR.
The vegetation may be fitly dealt with under the srveral headings of
(1) Dunes, (2) Cliff", (3) Fol't'st. I
(1.) Dunes.
The shore cl.t Sllalerb' Bay consists of a sandy 1I('ach about half a mile
long, terminating in rocky abutments, and having a row of dunes at the
ba.ck. These dunes a.re only a.bout 5 chains deep, and rise at their eastern
extremity to I~ hl'ight of about 100 ft. They are fairly sheltered by the
background of hIlls, but Itrl! exposed to the north and north-west winds.
Most of thr dunps arp fix('d, but in parts the sand is still unstable, and
towards the ('alit tlw dunes show evidl'll('e, in their greater height and loose
appearance, of thl' I'ifert of t.he northerly gales. The fore dune. as is usual.
is covered with the ('ommon sand-binding ScWpus frO'tlJd,08U8. Behind this,
however, Ii much mOl'C complex vegetation is found. 'rho principal plants
of the association UTe Poa tXJespttoBa, vdth on abundance of Linum mono·
gymtml, Ooprnsma, acerosn, and Pitnelea LJallii .. while dotted throughout
these arc 001"'os11la Oolensoi, Halorrhagis erecta, E9.IIph,orbia glauca, SoiRpuB
notlosus, Senecio lalltUS, and Anisoilwl(, i'/ltmnedia, with stunted forms of
Myrst'I'/R Urvillei. Coprosma ~erosa forms ill many places an aIni.ost COll'·
tinuous mat ruuning flat over tIle sllolld, and Pimelea Lyai/M, a.lso takes
on a. l:Iimilar habii, ('xcept it is dimlJing through some other plant, when
in places it rt'adu!s u heigbt of I~bout 4. ft. In damper situa.tions patcher
of IHeroohloo redolC'flS and IIydrocot1l1c nooae-zealantUae make their appear-
ance, while hl'r(' and th(']'(,' Acarna BfI'l'Ig'UiBorbae, G(!JI'am4um BetJstli'/l()ll'llrIl, or
M'lJeklefthPc1.-in colnplnn (·lCc.>p ove1' tht' sUl·facp. Tarna:acum o{fioit6ale var.
glabratum lind Erecktites prf'll(Jntlwidt>8 arc also fOllnd. with ol'casiona!
plants of Verollivn (,l/i,1)tica, C'aly8legt'a Solilanella, J.Altllnn'a alpfM, Epilo-
biuf'll 'Y&erlt'rioidf's, E. itlftCl'ul1I, GnaphaliAun tuteo-albuftt, alld G. iap(mitJu?n.
In parts therl' u.rc many plants of C'raspeiliA umflora var. robusta, with
the ]l<\tur;~liZl'cl FoonicIllu1l1 officinal!'. Where the dunes ara absolutely
"tabla the plant coyering changes somewhat, and the followi1l8 typical
assooiation is fouud: Pteridium aquilinutn ill patches, Poa tXJespitoBa,
&i,rpUB nod08U8, Pllormium Oookianum. A.etrel1.a SanyuiBorbae, Halorrhagi~
('recta, Veronica elliptica, with dWlI.rl specimens of .tl.riBtotelia ,.acemosa,
Di.cksonia sqUa.T'1'OBa. A.spidium I.H'stit'llm, Senecio rotundi/olius, Astelia
1wtW8(l, ]Mnaria cape'Mi8. A.splmium lucidum. Ora.spetlia umflol'a, and
MgrBim.e U'I'oillei Further hack, next thl.' edge of the bush, Lepta-
llpertnutrl sooparitllm is found, with here and there sUlalI pa.tches of Lagmw·
ph,ora 'J'U1IKla, G1J.nnera arenaNa. SOfU!lltlll littoraliB. and A.pitmt 1"'Ostral.ufII.
and nea.rrr tIl(' fI}lOre Rumea: 'lI'qiPrtIiH. Peg/tlca littoralis is also fairl~·
7H ]'1'.7 /1 ~Il r f ItJ " ••
plentifuL M their highobt point, where the dUlll'-plants merge into the
forest, a heath is found, ill which the principal plants arc EOf)lOHia caprp.91S.
LeptOBpermum scopar'illm, Mueh1enbeel'ia oomplexa, !Jycopodi/um, 'lJotuln"/t',
G®Ztheria amipodl.lln vaT. erecta, Aristotelia raoem08a, Oarpodetus serratus,
stunted Wf'il'lf?lfln,,,'11 rlloomosrt., DraoO'jikll11wm 10llflt·/oliu?n. and Ptl"l"l·rli'l.l'n'
aquilinum.
(2.) OlifJs.
The Ilssol'il~tioll in these situations difiers 11 good deal according to tlll'
varying situation, the principal factor in the change being, appo.r('ntly,
wind. Thus, on the exposed points, where the wind. has most efi(,(:·t, Lhc'
principal plant is Olearia angusti/olia, which is so plentiful in placos us 10
form an almost pure association. Hitherto Olearia mr,gustifolia hat! bl'en
reported only from BOuth of Paterson Inlet, on the east coast of StC'w,Lri
Island, and from the north and south ends of Mi\son's Bay, on the ~('SL(,l'lI
coast. The only other localities where it has been observed, apart {rom th('
Stewart Island habitats mentioned, are the base of Bluff Hill and Puy&c·gur
Point. Not only is it the chief plant of the coastal rliffR of Codfish Isltmd,
but it is equally a.bundant on the seaward base of tho Rugg~dy Mountu,llUI.
It forms almost the solo plant covering of the Rugged Islands, whorl' thl'
whole clifi-sid('s for hundreds 0:( feet are one close mat of stunted wcatll('J'-
beaten plants whose handsome grcy-green rosette-like foliage and rounded
form stamp the physiognomy of the coast-line in a most marked ru.a.nner.
Dr. Cockayne, in his Stewart Island report, dl'&W8 attention to the differ-
ence in the size of the leaves on differont plants of this spl'cies, noting
two forms of leaf, one abont i in. to 1 in. in diameter and the o1her only
about l in. wide. This same peCUliarity was noted by me on bushes grow-
ing side by side, and seemed to me to be constant throughout all the
leaves of the particular plants, so as almost to suggest varietal distinction.
.Although Olearia angUBti/olia is the chief plant on cliffs, its prooominance
is confined to the water's edge, and even there in places it is much mixed
\\ ith 86'MOio rotuMi/oZius. Spenking generally, Senecio rotunai/olius in-
oreases as a greater height is reached, when Olearia OOl6'flBO~ croeps into
the a.ssociation. The threE' plants named {orm the basis of tho .. Scneoio-
Olcaria" Ilssociation so exllausti vely dealt with in Dr. Cockayne'l:I rl'port
above mentioned. Growing throughout this association 'Will bo Ioulld
numerous specimens of Vnonica clliptica, with horo and th('r(' pL'lnts of
Phormium Ooo/Cian'Um, Ani80tume illtf'f'lnedia, Dracop/,ultuln longifulitlm, and
. the shore-fol'Ilfl Lomaria t/;wra and Asplenium lucitlutn. OClOO8ionu.} speci-
mens 0:( Nothop~ OoleMoi push their green heads through the close-
growing scrub. Ou thc rocks 11.1, the foot of the clifis the plants llOted
were Ora8sula mosohata, Selfiom radicam, Apium prostratum, My080tiB
albiflora, Soi''P'US notlosua, and Gentiana SOtZ08a. Where the pea.t was drier
GnapMUum trineroe Bnd Aspidium fJB8titwm were also observed.
At the wostem side of the bay, where the cliffs are more sheltered, a
much richer flora was seen. Here, as before Olearia angustifolia and
StIMOio rofllJln,di,/olius predominated, but Olearia Ookmoi also appcared in
increased numbers, until, as the top of the steep faces was reached, it
took the place of the first-named. species in the lower formation. or
SmAller plants, the principal were Tetragonia trigyna, AniBolomr. inter-
f7I8dia (plentiful), Gemiana 8rJaX)8(J, Ora8sula mosonata, Poa Astoni and
P. Oo'leNoi (on the bare pointe), AspZemum luc:ULum, A. OOtuBatum, MlMlef1I-
bryantMmtSm a'UStnlle, PM /DZiosa, £amaria mwa, Phrmn.;'wfII OoolMftUm.
POPI·EI.V. ~Ll •. -Pll£l~t Cnl'el'mg, Oodfill/, illam' IIlId /I"!lfI,d IRtCII~h 7~
the semb, the growth of lovely plant forms which groets the visioll is
delightful. Here the coast-ferns Aaplenium It.It:idum and Lomaria dura
grow to perfection, while straggling plants of Pkormiwm Oookianwm are
found seemingly somewhat out of their station, which is usually on the
r.oastal (lliffs in this region. Great ooloniea of Btilbocarpa LyallH make
their appearanco, in places over 3 ft. tall, and with their large renifonn
leaves suggesting more a tropical growth than a subantarctic one. These
latter plants are extremely abundant, the pat6hes in places extending
over areas almost a quarter of an a(lre in extent. As we ascend the hill
the association changes. AaplefliulI& obtusatum of large dimensions put.ll
in an nppearance, along with Lornaria discolor, Hypolepia f.efrIId/olia, ABtelia
tlervosa, Polyponium BiUardicri, NotliopooaJ: Oolensoi, M!psi'llf Uroillet.
OoprOBma luoida, and DioksO'flia squarrosa. Here there is evidenco of bird-
life. The burrows of the mutton-bird (Puffi;tlus griseus) ramily in all direc-
tions through the peaty soil, and no doubt affect the plant-association by
draining and aerating the soil, while the traffic of the birds themselves
must destroy much of the usual undergrowth of the forest. What appeared
to be a direct efteet of the nests of this petrel was noticeable in one place
where a mass of the fern Lomr.vria dura was growing on a heap of humus
whi6h was literally honeycombed with burrows. The plants were much larger
than usual, and each had a distinct caudel: about 12 in. high. At a height
of about 250 ft. the forest became more mixed and the undergrowth thicker.
Nertera i!.ic1&tmaKae/olia grew on the logs, OoprOBmtJ foetidissima became
common, and Aspidtwm tJUtitwm was added to the former aasoci.o.tion. At
300 ft. the first Me/,r08iiJ,erOB Zuoida was encountered and DraoopkyU!Jm
became fuirly plentiful. NotkopUlf&(1ll; Edg61'le'IJi also appeared, and M!pBiIIe
Urvillei, DiokBooi,q. squarrOBa, and Notkopcmu,z Oollm.8oi became plentiful.
Tho forest-:8.oor now became cevered with Lomrmia CIJ'[I6'II8i8 nnd Lomana
discolor, the former especially attaining large dimensions, whilo the spooi-
mens of Asplenium obtusatum booomo larger also. As the top of tho ridge
waH roached Pittosporwm Ooltmsoi var. /asc1oulaium appeared, with a few
plants of ~ aquiUmim nnd Rubus austra.lis. From the top a fine
view was obtained to the south-west. Looking over the forest-top, one
could see the wind-swept appentnnce to the south-west, the principal plants
being stunted MeflrosiiJ.ero8 luoitltJ, Olearia. Oolensoi, BtIA8IJi,o rottmditolWs,
Draoop1ly'llwm lOtlgi/oliwm, with ocoasional specimens of M'IJ"Me UrvillIli.
Proceeding along the ridge, WeitlmlJMia raoemosa became more plentiful,
and ABtelia increased on the forest-:8.0or, along with Lomrma oapemis,
Aspidium ~, Aaplenium bulbi/uwm, and A. obtusotum, the general
depth or the :8.oor-covering being about 4 ft. In hollows where the
ground was damper a strong growth of fom-trees (Dioksonia 8qucm'OBa),
with No!1wpUlf&(1ll; simpls:», B'MfflerG digitata, and a broad-leaved species of
O~ tmlaria., was seen, while the logs became covered with Lv.zt.wi,aga
margillala, and :6.1my ferns. Here also a few examples of Podaoarpus
I~ were seen, and the tangled atems of the RlWpogonum BOIJtIIlIms
blocked the way, and marked the edge of the forest proper.
•• TAlI " lUmti-KIDIICIM" l'oruI.
Lying to the back of the sandhills, an extensive area of forest of this
class is visible. Its outer fringe touches the sand-dunes, and it extends
to the highest parts of the ialand, although tho trees become stunted on
the heights, and rooks show through the low scrub on the very summit
The lowland bush consists prinoipally of GriseWMa 'littoralis, M~
'fRANS. N.Z. IN~'!:, VOL. XLIV.
PUTE: IX.
D. OONCLUSION.
(1.) PTERIDOPHYT.\.
Hymenophyllaceae.
HymnlOph!lllulII dPlIIll!IIUJII (Forst. t.) Sw. On logs III damp fOll'lt
plf'ntIful.
tlzlotatUll1 (Forst. f.) Sv.-. On logs III damp forest; plentiful.
sanguinolPn.tum (Forst. f.) Sw. On logs m damp forest; plt'ntiful.
tunbrirlgense (L.) Sm. On loll,s in damp forest; plentiful
Cyatheaceae.
Dt.ckso'tllU squafTosu (Forst. f.) S". Abundant III fOI'I:!&t.
HemiteUa Bmithii (Hook. f.) Hook. Forest; not pll'ntitul.
A.lsophiln COlpll,oi Hook. f. Forl''1t: rare.
Polypodiaceae.
PolypodfulII Billardierl R . .81'. Plentiful III forest.
/lUllfral" Mett. Logs in damp forest.
Pterirliu)11 aquill1/.uI7I Kuhn. Heath and stable dunes.
Pteris inelsa Thunb. Damp forest.
A.spidium vestitum Swartz. Fai:rl~· abundant in forest.
A'plemum bulbi/eruln Forst. f. Abundant in fOTest.
fllWCiilum Forst. f. Abundant in forest.
luoidum Forst. f. Coastal scrub; plentiful.
LJmaria alpina Spreng. Dunes; rare.
dura 1\Ioore. Coastal scrub: abundant.
/fmreulatta Spreng. Forest; abundant .
.cllpplIsis WIlld. Forest: abundant.
dUlculor Willd. Forest; ahundant.
Hypolprpis felllli/alia (Forst. f.) Bernh. Forest: abundant.
Lycopodiaceae.
LyCllpfldwill t'Oluulle .Forst. f. Stony heath; plentIful.
(~.) ::)PERllOPHYTA.
Taxaceae.
Pndooorpu8 ll(ulii T. Kirk. In forest; commOll.
jerrugineull Don. In forest; fairly common.
DtJC'fI{dilt'fll ('ullff'ssimmt Sol. III forest; abundant.
Gramineae.
Hieroohloe ,,&ioltlts (Forst. f.) R. Br. DaDlp dune-so
Poa /OUOBa Hook. f. Coastal clifis •
•4.Btoni Pt'trie. Coastal cliffs.
C~spit08a Forst. f. Dunes; abundant.
c.'olensoi(n Hook. f. Coastal elifis; rare.
FelltuM It'ttflralil! LabiU. Dune-s: f:lirl~· ple-ntiful.
POPPBI.WELL.-Pla/~t COI'erlllfl, Codfill/I /IIWnd IJntl Rugged Island.. 8S
Cyperaceae.
Scif-ptUI nCH1.osus (R. Dr.) Rottb. Dunes; plentiful
/"~8U8 Banks & Sol. Dunes; plentiful.
Uncmia pedicelloUr Kiikcnth. Dl\mp forest.
Oar~ lemaria Forst. f. Wet ground; common.
trifion Cny. Damp ~tro\md; open forp8t.
Liliacea.e.
Rhtpogoou1/l Ilcandem }.'ore.t. Plentiful m forest.
Luzwriaga marginata (Banks & Sol.) Benth. & Hook. 1. L06s lD Iorest.
Alltelia fIeI"Vosa Bunks & Ro1. Dunes, sht'ltered rocks, iorc·st.
Phormium OooHa'1ltVtn Lp Jolis. Coastal rocks and scruh
Polygonacea.e.
Rlmltz negleotus Kirk. Stony beach.
Mu('1I7.en.lJeckfa cmnplea'n (A. C11nn.) Meissn. Dunetl, damp forest, hpatil.
Aizoacea.e.
MesembrycmthetllU111 austrn[r Sol. Coastal lOclal; rart'.
Tetragonia trilJlItln Bunks & Sol. ('oastal cliffs; ral'<'.
Crassulaceae.
Vms8ula moscnata Forsi. f. Coa.stal rocks.
Saxifragaceae.
Oarpodetu8 serratus Forst. Forest; plentiful.
Pittosporacea.e.
PiU08pOJW,& ten.ui/olilflltl Banks &; Sol. In fOl'<'st; ran.
C'olensoi var. j(J8CiCfl7atfll'll (?) Hook. f. In for<.'st; rare.
Cunoniaceae.
"Wt'inmamua facemosa L.!. Abundant in forest.
Rosaceae.
RtIbus australis Forst. f. In damp wrest.
sc1wnMeZioides A. eunu. In damp forest.
Acaetla Sangtti8MOOe Vahl. Plentiful on stable dunes.
Geraniaceae.
Geranit.vm Be8silifiMutn Uav. Dunes; abundant.
Linacea.e.
Lmum ,lltmOIJplum Forst. f. Dunes; abundant.
Euphorbia.ceae.
Buphorbia glauca Forst. {. Dunes; ahundant.
84
Elaeocarpaceae.
A",stotl'ltU raU1IImta (A. Cunn.) Hook. f. Plentiful: tstahlc dUDt'f! and tOI'OIIt-
M!_!e.
Thymelaeaceae.
Ptf'llRka Lyall;! Hook. f. Dunes; abundant.
Myrtaceae.
Leplospermum scopanuli/ Forst. Edge of forest; plentIful.
Mel.r08iileros luoida (Forst. f.) A. Rich. Abundant In for(,Rt.
kypetioifolia A. Cunn. Rare in forest.
Myrttlll pedtl!1lCulata Hook. f. Damp forest. rarti.
Onagraceae.
EpilobiulII nertenoide8 A. Ounn. Dunes; faIrly plentiful.
;ut&06lJm Sol. Dunes; rare.
linnaeoidell Hook. f. Dunes; rare.
Fm·hBia l!ZOariicata L. f. Edge of forest; 1'8rt'.
Halorrhagaceae.
Halorrhagill ereota (Murr.) &hindler. Dunes; plt'nhful.
~ftef'a flrl'1tnria Cheeseman. Dunes.
Araliaceae.
Stilbocarpa Lyallii J. B. Armstrong. Coastal &crub; almlHlant.
StltAopana:& simplez Forst. f. In forest; rare.
Edgerleyi (Hook. f.) Seem. In forest; plentiful.
C'oleMni (Hook. f.) Seem. In forest; plentiful.
,schsfftera digitafa Forst. Damp forest.
Ps£uoopanax crassifolium (Sol.) (I. Koch. Pl('ntiful in forest.
Umbelliferae.
Hydrocllt"it IW!YJI!-se'tdandiae D. C. Damp dunl'&.
Apium prllstratum Lab. Coa&tal rocks and dunl's.
AnmMtt( i"tl'I'ml'flia Hook. f. Ooastal rocks; plentiful.
Comaceae.
tflotselmUl ',Uoralis Raoul. Forest; not plentiful.
Ericaceae.
Gaullhena antipoda Forst. f. var. erecta Ohel'sm. Forest; oomparatively
rare.
Epacridaceae.
o't1Jikelta Sol. In forest; fuirly plentiful.
act!1'08t1
Dracophyl1um longifolium (Forst. f.) R. Br. Coastalscrnb; abundant.
Myrsinaceae.
Myrsinc Uf't,illtl (A. D. C.) Mez. Dunes and forest, coastal scl'Ub.
l~mttmia ditlQrtca'a (A. Cunn.) Hook. f. Damp forest.
Primulaceae.
SaMOlw, 1epffl~ Forst. nf. p1'OCtlmberis, R. Knuth. Da.mp rocky situu-
tiona noor short·.
POPPI!IMvHLr•• -Planf ()Ollulng, (.'odjr.JI 1.1)md ami R1tgged 1.1400.. In
Gentianaceae.
GtflttutW SaXON(( FOl'Kt. l. (lolLstal rocks: plentiful.
Convolvulaceae.
Va111steglQ Sn/tf(11l('11a (I,.) R. Hr. Dunes; lah'.
Boraginaceae.
Mgosot'UI alhiflnra (T. KII'k) Ch(>t'Rt'm. Rod{l, ncsr sea.
Scrophularinaceae.
I'mJntca BaZ,eijoUa F01'St. t. In damp iort:st.
t'llipti~a Forst. t. Plentiful on ('oallhtl l'Ol'ks.
Rubiaceae.
l'Q'jJrQllm.a iucula F01'St. f. III iorest; rare.
arooiata Checsem. Damp forest.
foetidis,,'ma Forst. Abundant in forest.
ihotmnoideB A. Cunn. Plentiful in damp plaC<'s.
aeet'OSG& A. Uunn. Abundant on dWles.
propinqua A. Cunn. Damp forest.
{lolensoi Hook. f. Plentiful in damp lorest.
N('T(erQ deprt'88o, Banks & 801. On logs in forest.
tlirn()ntlrat'/olia (A. (illlll.) Hook. f. 011 logs ill forl'st.
Goodeniaceae.
8rllit!l"a I'ad,Ctmll (.lay. Dlunp places on rOBstal wt'kt..
Compositae.
LagenopktJla pumikl (Forst. f.) Cheesem. Stablo dUlles.
Brae1lyoollle T1AomBorvii T. Kirk. Stable dWles.
O/('aria angllsti/oUa Hook. f. Coastal cllifs; abundallt.
Oolemoi Hook. f. Coastal scrub; abWldant.
iJnaphaliUtn. triner'OO Forst. f. Dunt's; plentiful.
[ltteo-album L. Dunes; plentiful.
japonicum Thunb. Dunes; plentiful.
(}ra8pedia unitf,ora Forst. f. var. robusta Hook. f. DUlles; abundant
Erechtite8 prtmafftboideB (A. Rich.) D. C. DUDf's; fairly plentiful.
Senecio la'Utt18 Forst. f. Dunes; rare.
rotundiloZiIus Hook. {. Coastal scrub; abundant. In forest; raJ't!.
Tarazacum glabratum (Forst. f.) Cookayne. Dunes; airly plentiful.
BO'flchm 7ittora1is (Kirk) COCkayllf'. Dunes.
ART. V.-IA.st of Ltc'heM (md FUll,,' collected 11/ fhe Kt'rmlltf,'O 1/lIQl~d,
1908.
ttl.
By W. R. B. OLIVER.
[RllOd ftfore 1M A.uclrlaflll In!diflltt, 28th Novemb8l", 1911 J
THROUGH the kindn('ss of Mr. T. F. Cheeseman, F.LJ:!., \vho forwarded
the lichens and fungi which I collected on Sunday Island to the Director
of the Rew Botanical Gardens, and subsequently {urnished. me with the
names of the species represented, I am able to publish the following list.
The fungi were identified by Mr. George Massee, and the lichens by Mr.
A.. D. Cotton. I know of no record of fungi from the Kermadecs, but in
the" Handbook of the New Zealand Flora" (1864-66) Sir J. D. Hooker
has recorded two lichens-Oladonia /urcata and Leridea intermi:l:ta-collected
by Mr. J. Milne, who visited the group in H.M.S. "Herald" in 1854.
Neither of these was collected by me.
Lichens and the fungus Scoriaa spongiosa form a conspicuous featurp
of the forest on Sunday Island-not by the number of species represented,
which are few, but by the abundance of individuals. In the UppOI' wet
forest almost every stem' of palms and. trees is covered with mosses and
lichens. Of the latter, those especially common are the foliaceous Sticta
uariabilis and Lepwgiwm cyanescens, and the crustaceous Baeomyces
pertem,uis and Physcia speciOBa. In exposed places, as on ridges and cliffs,
where more light penetrates and the wind is more desiccating in its
effect, the tree-stems support chieft.y the foliaceous Stiaa aurata and the
drooping US'MfJ barbata. On rocks along the sea-coast Xaflthoria parietima
and Physcia pulveruZenta are frequent. All the upper branches and twigs
of the pohutukawa, especially in the lower dry forest, where they are the
dominant trees, are completely clothed cith SCO'I'ias spongiosa, which.
showers its black spores copiously on the {Ol('st below.
LIOHENS.
Baeomyces pertenuis Stirb.
Forest j on stems of nikau-palms and trees.
Cladonia capitella Ba b.
Forest; among moBSe8 on horizontal branches of trees.
Cladonia Florkeana Fr.
On logs in open.
Cladonia aggregate. Eschw.
Forest j on damp ground.
Leptogium cyanescens Kbr.
Forest; on nilmu-palm stems, in damp situations. Thallus soft and
moist, like an algo., and. in dry weather shrivels at the edges.
Sticta variabilis Ach.
li'OfflSt; abundant on tree-stems.
01.1' J<..R.-LII·/UIIR alltl f',m!J1 roll,.,·tul III Kfllflftlitl Isla",l. 8'7
FmmI.
Arcyria punicea PerA.
Trichia fa11ax Pers.
Forest-floor; on underside of dead leaves of nikau-palm& (Rhopalo-
I>tglUJ Baueri).
Scorias spongiosa Fr.
Forest: on pohutulru"'a-trec& (Jieb'()8iikr08 lJill08a). Thlb tw.tgUb com-
pletely covers the upper brunch('s of the pohutulmw3-tl'el:'s with a soot~·
black moss-like growth to a depth of i in. to i in. It continually sheds
its black spores, so that the 1t'3ves of all trees Ilnd !lhrubs below :trl'
('overed with a black dust. Locally it is called .. pohutukawa !loot,"
which well expressc.>s its appearance." and habit of ('odtinj;! j'vprY1:hing with
1\ layer of black.
character may be variable even within the limits of the same specil's j
therefore we must not assume that if a character separates natural genera
in one instance it will also do the same in another. There is no scientific
reason why secondary sexual characters should not be used to definE'
genera in those cases where they are found to indicate natural genera in
accordance with the above-mentioned principles; tufts of hail' (probably
scent-producing) in the male sex are, however, found in practice to be of
specific value only-at any rate, as a general rule. Hampson oddly and
inconsistently refuses to use any sexual characters for defining genera.
whilst invariably employing these same characters, even the specific tufts
of hairs, for forming sections of genera; whereas these should in any case
be limited on exactly the same principles as genera, being of smaller value
but precisely the same nature.
I will gi~e one or two specific instances of the unsatisfactory nature
of Hampson's results, to illustrate my meaning. Hampson makes a new
genus En'opygor1Rs for two European species and the Hawaiian euclidias
Meyr. This could only be explained geographically by supposing that
at some former period a straggler of the genus from Europe had reached
the Hawaiian Islands, which is unlikely, but, of courso, possible. Bul;
euclidias is an insect of striking appeara.nce, and two other Hawaiian
species, oompBiaB Meyr. and wipluldopa Meyr., are structura.lly and super-
ficially so close to it that it is impossible to doubt they are closely related.
These are placed about seventy pages 0:11 in the genus Hyssia, which con-
tains about fifteen North American, European, and New Zealand species.
and a separate origin from another straggler is required for them. The
difierence stated is that HysBia has the thorax clothed with sca.les mixed
with hair, the abdomen with dorsal crest on first segment j Eriopygodes,
the thorax clothed with hair only, abdomen without aresta. But euclidias
(of which I have a long series) certainly has Ii small abdominal crest. and.
the difierence in clothing of thorax is imperceptihle. I conclude that
eucUdias must be transferred to Hyssia.. But the only distinction be-
tween Hys81'a and the cosmopolitan genus Oirphis. with 140 species, is
that Hyssia has the thorax clothed chiefly with hair-like scales, and
Oirphis almost entirely with hair. This is a distinction ,vithout a
difference, and, in effect, I am quite unable to distinguish the species.
assigned to these two genera by this or any other structural character.
though they are separated by two hundred pages, and plsced in widely
remote branchetl of the phylogenetio tree. I am therefore obliged to unite
them, which makes the Hawaiian species a local group representative of
a cosmopolitan genus. and puts quite a difierent face on the matter. But
on examinjng '&ri,opyga, with 100 species (chiefly American. some European),
only stated to difier from the above by absence of abdominal crest, I find
that some at any rate (e.g., the European Tu?ca L.) certainly possess Ii
small crest (no doubt the character is often difficult of observation, beca~
the base of abdomen is clothed with rough hairs, aud the thoracic hairs tend
to conceal it also, but when present it is formed by scales of Ii different
character and difierent colour), and must be referred to Hys8$a also. I
am not well supplied with the American species. but the genus at lea.at
requires cleansing. And BoroWt, with forty species, mostly A.frican and
Australian, is only stated to difier from Eriopyga exactly 80S H1J88ia. does
from (Arrpkis, a distinction found to be inappreciable, for the supposed
difference in form of wing (more oblique termen) cannot be seriously rt"-
garded as a generir oharacter, and therefore this a]so needs reconsideratioJl.
90 TrOHso('t,ons •
OABAD.B.lNINA.
I adhere to my ,iew that the namo N od.ua, carrying with it the group-
Dames NodN.iiJ.ae and Noct.v.ina, is inapplicable in this connection, and it
haa now heen abandoned by most authorities; but Hampson proposes to
MXYRIOK -RI!,I"~'OII of fIll .\'/"11' Zealnl'u l (!SI.IIl!·ininll. 91
118t' it in a sense in which it haH ntlvor betln used by an~' 011e, a result (If Ius
principle which can only induce confusion.
The OaraiJrimina are a highly developed modern group of iIlLmt'lllItl
extent, Lut, with the exception of the Melanohrid group of the Carad'fl-
nidae, they are represented in New Zealand only by a very few scattered
stragglers, and some very extensive families and subfamilies are not repre-
HE'nted at all. There can be little doubt that these stragglers dre the
outcome of accidental wind-borne immigration over a wide expanso of
sea, which accounts for their scantineBB. If Ne,,- Zealand ever had easy
communication with any land, such land did not at that time contain any
of these poorly represented groups; but, as these groups art' of relatively
recent origin, such communication may have existed in earlier tunes. Now,
as the Melanchrid group pOBBeSBeS no sort of advantage that \\ould explain
their easier introduction, and as this group is, on the whole, quite .\S well
developed in New Zealand as in any other region, I eonsider it good e...idence
that an easy communication with some land did once exist, clond that thc
Melanchrid group then existed ill the land in question and made their
way into New Zealand. It does not follow that the Melanchrid group is.
older than any other group of the Caradri",ina, because any or all of the
other groups may have coexisted at the same time in other regiens cut off
from New Zealand and the land in question by wide seas. This raises the
mteresting problem of determining where the land ill questIon Wdl:I, and
Ii proper comprehension of the classification and geographit·al distrihution
of the Melanchrid group would enable us to solvE' it with tolerable cer-
tainty. We do not yet possess this comprehension, Lut offer the following
considerations. The only possible lands seem to be four-viz., .Australia,
the Pacific islands, South America, and the Antarctic Oontinent. Australia
may be excluded; the Melanchrid fauna is pretty ,,-ell known, and llU:Lkes
no near approximation to that of New Zealand. The South Pacific islands
are certainly incompletely known, but there is no evidence that what exists
of them at the present day pOBBesses any special Melanchrid fauna such as
might be expected on this. assumption. The Antarctic Oontinent naturally
poBSesses no existing fauna, and, althougll it may haye served as a rout"
of communication, there is nothing to show that it ever had one of an
aboriginal type. We are therefore reduced to look to South America.
and the few species known from Chile, Patagonia, and the Falkland Isles.
(probably only a small fraction of those existing) are of a cha.racter which.
in my opinion, agrees well with the New Zealand types, and probably
indicates real affinity. I suppose, therefore, that the Melanchrid fauna
entered New Zealand from South America, probably by way of the antarctic
land, where it may have undergone some modification during a perhaps
prolonged passage, at a date so far remote that considerable speoific and
IIOme generic development has taken place since. With it doubtleBB came
XOIIIikO'l'noe, Notoreas, SeZidoBema, Orambw. Diptyc'hopnora, Scoparia. and
Bor1rJulIuBenia, the largest and most characteristic genera of the New Zeals.nd
lepidopterous fauna. Probably the original source of this fauna was the
temperate regions of the Northern Hemisphere, and it travelled to South
America by the great mountain-chain of the Rocky Mountains and Andell.
At the time when this fauna left North A..m.erica probably the Indian region,
which has been the principal source of lepidopterous evolution, was isolated.
amd extensive developments may have· been going on there; but, as the
C",aiJlf.maae as a whole must have originated in some one region, it cer-
tainly seems that the Melanchrid group must have bf'6D. spe&k:in~ generally.
th,· earliest bra.nl·h of the famIly, and I propose to regard it as such on
this ground, since the IItructuraJ chamcters are such as t.o give no help
either for or against the theory.
The generic characters gi\'en below are, for simplicity, drawn to apply
to ~ew Zenland specit'fI only.
1. ARCTIADAE.
Vein 1:1 of hindwin~ anastomosing with upper margin of cell from baHe
to near middle .
.A. large cosmopolitan family, which is barely represented, whilst the
allied Syntomid. ~olid. and Lithosiad groups are entirely absent.
1. Metacrias Meyr.
Jlttactias MeYl·., Proc. Linn. Soc. N.S.W., 1886, 749: type, erickrl18tl
Yeyr.
Tongue obsolete. .Antennae in ~ bipectinated to apex. Palpi short,
hairy, concealed in long hairs of head. Thorax and femora densely hairy
beneath. .Anterior tibiae with apical cla.w, posterior tibiae without median
spurs. Forewings with 7 and 8 out of 9, 10 sometimes connected with
9 above 7. Hindwings with S, 4. 5 nearly approximated, 6 and 7 connate
or short-&talked. 8 anasromosIDg to .. of cell. Wings in !j! rudimentary
or absent.
This interesting endemic genus is of doubtful affinity, but appears to be
nearest to Ocnowna. which is a genus of about a dozen species located
round the shores of the Mediterranean: Hampson also assigns to it one
species from Peru.
I. M. Hutto," Bud.. Cist. Ent., :l, 487 j Meyr., Proc. Linn. Soc. N.S. W.,
1886, 750; Huds.. N.Z. Moths. 5. pI. 4. 6: Ramps .. Cat., 3. 468.
LA ke W'akatipu.
2. M. mcllrYBn lIeF" Pro!'. Linn. Soc. N.S.W., 1886. 74:9: Nuds .. N.Z.
Moths, 4, pl. 4, is: Hamps .. Oat., 3, 469.
Mount Arthur; 4:.000 ft. Larva on Senecio.
3. J1. 8trategica Huds., Entom., 1889. 53; ib .. ~.Z. MothR. 4, pI. 4:, 4:;
Hamps. Cl1t .. 3. 468.
Richardson Ra.nge j S,OOO ft.
2. Utetheisa Hubn.
Utetlwilla Eubn.. Verz.. 168 (1823) j type, omatrUr Lilm. Deiopeia
Steph., Ill. Brit. "Ent. Haust., 2, 92 (1829) ; type, puZolzella, Linn.
Head smooth. Tongue developed. Antennae in d ciliated. with longer
setae at joints. Palpi moderate, ascending, with loosel~' appt'essed scales.
Thorax smooth beneath. Posterior tibiae with all spurs \'cry short. Fore-
wings with 7 and 8 out of 9, 10 connected with 9. HilIdwings with S, 4,
!.S rather approximated. 6 and 7 connate or short-stalked. 8 anastomosing
to middle of cell.
A small cosmopolitan genus.
4. U. pulohelltJ Linn., Syst. Nat., 1, OM (1758); Meyr.. Tra.ns. N.Z. Inst.,
22, 217; Ruds .. N.Z. Moths, 3, pl. 4, 3.
Wellington district. .A. recent immigrant, doubtfully established ;
occurs throughout Europe, Asia, Africa, Australia, and South Paciflc
islands. Larva on J'liyOBOti,s, grasses, &C.
M.JlYRICK.-Revi8ion of tM lYe1/) Zealand Caradrininllo. !I~
2. HYPSIDAE.
rein 13 of hindwingll connected with cell by bar near bllde.
A rather small family, chiefly tropical. The following gellus ~
tormerly placed in the A'I'ctiadae. the approximation of vein 8 being SCI
"rOSE' that it appears to be anastomosis.
:1. Nyctemera Hilbn.
lactioi",~a Cram. Leptc-
Nyctemera Hiibn .. Verz.. 178 (1823) j type,
BoiRd.. Voy. Ash. 5. 197 (1832) ; typl". annulata Boisd.
BOtIItJ
3. CARADRINIDAE.
Vein 8 of hindwings shortly anastomosing with cell near base. thtlnce
diveTging; I) obsolete or imperfect, rising from middll' of tnt.nsverse vein.
A.n extremely large family, of which, as explained above, only one sub-
family is adequately represented in New Zealand.
Subfam. 1. AGROTIDJilS.
Eyes glabrous j tibiae spinose.
4. Heliothis Ochs.
Heliotkis Ochs., Schml'tt. Eur., 4:, 91 (1816); type, tli,pBfMJB(J Llnl1.
OkloriiJ.eu West",".. Jard. Nat. Libr.. 32. 198 (1841) j type,l1lr68CtIl8
Fab.
Face with rounded prominence. Antennae in ~ ciliated. Thorax and
:iI>bdomen without crest. Interior tibiae with apical inner and outer clsws.
A rather small cosmopolitan genus, of which some species range very
widely; one of these has reached New Zealand. There are about a. dozen
other generic synonymB. which it seems needless to quote: some anl
.explained in the p:rt'limina~' remarks.
.6. H. aNnigera Hiibn., Samml. Eur. ~hmett., 370; Meyr.• Trans. N.Z.
Inst.. 19, 34: Huda., N.Z. Moths, 32, pl. 5, 4:0, 41: conferta. Walk.,
Cat.. 9. 690.
North and South Islands; a l'osmopolitan insect. I.arva. poly-
phal!ous, on seeds and flowers.
94 TransactIOn •.
5. Euxoa Hiibn.
Eu.eoa Hulm .. \Terz., 209 (1823); type, df'cora Hiibll.
Face ,vith small truncate-conical prominence Wlth ralBed lllll. Antellllae
In If bipectinat.ed, towards apex simple. Thorax w'ith rathel sprClnding
anttrior and posterior crests. Abdomen without crests.
An extensiye cosmopolitan genus. There are auout twenty generic
synonyms.
i. E. radza'M Guan., Not.1:., 1, 261; In'Uttaa W!l.lk.. Oat., 10, 348; 008'-
,.,otata, ib., II'S, 1686; turbulenta, ib., S2, 70S; iwitmcta, ib., 32, 70S;
scapularis Feld.. Reis. Nov.• pI. 110, IS.
Dunedin. Common in Australia; also from Friendly Islands and
~orfolk Island.
~. E. adlldratio1,i8 Guen., Ent. Mo. :Mag., 5, 38; Hudb., N.Z. Moths, 31,
pl. 5. 37: sericea Butl., Cist. Ent., 2, 490 j Huds., N.Z. Moths, 31.
pl. 5, 38: incoMpicua Butl., Cist. Ent., 2, 545.
Christchurch district.
!to E. ceropac1wiik8 Gllf'n., Ent. Mo. Mag., 5, 39; Huds., N.Z. Moths,
32. pI. 6, 1.
Rakaia.
6. Agrotis Oehs.
Aqlotl8 Ochs., SchD1ett. Eur., 4, 66 (1816); type, upsilon Rott.
Lycopnotia Riibn., Verz., 215 (1827) j type, strigula Thunb.
F8.(.'e \\ithout prominence. Antennae in cr bipectinated, towards apex
.imple. Thorax with anterior and posterior crests. Abdomen without
cream. Anterior tibiae short, thickened, not longer than first joint of
tarsi. A rather limited but generally distributed genus. Hampson
separated Agrotis and Lyoopkotia by the "rather flattened" abdomen
of the former. but it is quite impossible to distinguish them praC'tically
hy this indefinite test.
10. A. gpsllml. Rott., Naturf., 9, 141; Meyr., Trans. N.Z. 11ll1t., 19, 32;
Ruds .. X.Z. Moths, 30. pI. 5, 35, 36: 8uf}usa Hiihn., Samml. Em.
~hmett.. 134.
Xorth and South Islands: a cosmopolitan insect. La.TVa poly-
phagous.
11 .•4. iM(lminata Huds., N.Z. Moths, 31, pl. 5, 39.
Wt'llingtou. Christchurch.
7. Grapbipbora Oehp..
GraphiphMa Orhs., Schmett. Eur., 4, 68 (1816); type, ob8cUr(1
Brahm.
Face without prominence. Antennae in a
ciliated. Thomx with an-
terior and posterior crests. Abdomen ,vithout crests. Anterior tibiae
moderate, longer than :first joint of tarsi.
A large genus, of universal distribution. Hampson includes this genus
in .Agrotw, hut I think the sepamtion is natural and practicable. This
is the group to which the name of NOc!lua ,vas formerly applied, but it has.
now been genemHy discarded. There are numerous ~eneric synonyms.
MEYRIOK.-Retluion of the New Zealand Cal'adrinina. 95
1:&. o. ('()'milia Wall .. ('at .. 10, 404: ltntnUntB, 1b •. 10, 4-30 Huds .. N.Z.
Moths. 7. pI. 5, 29: quadrata Walk., Colt .. 11, 745: mnocua, ib.,
15. liLO. '·Icipr()ca. tb .. 32, 672: brtviu80ula, tb •. 33. 716. ,'Of,,·
IIIW&tcata, ib., 38. 716: (J(J(>tlt&rl Feld .. Rl'lS. Nov., pl. 109. 6.
North and South Islanru.. Common III Australia, and reacb.iJ.1.§£
N('\\" Ht>hl'ldt>lI. L.lrY8 on Frlica.
:5ubfam.:3. POLIADE'I.
E~'l'b glahrous. hut overhung uv lonfl cilia from mar¢.ns; tibiae not
"'PIllOA(',
8. Austrarnathes HampEl.
AU8tramatne8 Ramp!!., Cat .. 6. 4:92 (1906); type, 11urpurea Butl.
Face without prominence, Terminal joint of palpi rather long.
Antennae in rS ciliated. Thorax with divided anterior and spreading
posterior ('rests. Abdomt'n without crests. An endemic genus of some-
what doubtful affinity; it is not very distinct, but the palpi are rather
chara.cteristic.
13. A.. pwrpurea Butl.. Clst. Ent., 2, 490; Huds., N.Z. Moths, 8, pI. 5, 32 :
Oef'amOikB Meyr .• TrllDs. N.Z. Inst .. HI. 31.
Wt'llin¢on, Dunt'dm.
9. Hypnotype Ha.mp!!.
Hypnot.qpe Ramps., Cat., 6. 411 (1906); type, placetlB Walk.
Face without prominence. Antennae in rS ciliated. Thorax with
d.nterior anAles ridged and projecting. and with anterior and posterior
crests. Abdomen without t'rests.
This Ilcnus is founded on a single South American species. I can only
refer the following species to it with considerable doubt, as I have not a
specimen for examination, and Hampson, unfortunately, had not seen
a. specimen either, but his ('onjectural reference of it to Sympistt'll is, I
think, undouhtedly woon!!..
14. H. pe880ta Merr. Trans. N.Z. Inst .. 19, 29; Hud!!.. N.Z. Moths, 6,
pI. 5, 26.
W('lJjn~ton. OhristchUTl·h district.
Rubfam. 3. MELANC'uRIDEs.
EYNI hairy: tibia.e:' not spinose.
26. L. sf'mwittata Walk., Cat., 32, 628; Huds., N.Z. Moths. 13, pI. 4,
21, 22.
North and South Islands.
27. L . .mlcana Fer., Trans. N.Z. lust., 12. 267. pI. 9, 3; Huds., N.Z. Moths.
13. pl. 4, 19, 20.
Akaroa, Dunedin.
28. I.... stulta Philp., Trans. N.Z. lnst .. 37. 330. p1. 20. 1.
lnvercargill district.
H. Physettca lIevr.
Physt!l.ICfJ Mt'vr.. Trolne.. N.Z. rn~t.. HI, ;) (1887)tvp . . , cfU'ruu.a
Guen.
Face without pronunPllCt'. AnteWla.e In a ciliated. Pelolpl III a WIth
tenninsl joint greatly dIlated, with orifice on outer SIde (mstead ot apex).
Thorax clothed with hair, without crests. Ahdom('n with 'lmnll crest on
ba~l se~ment.
Probably an endemiC dpvplopment of A~tta.
4.1. P. caerulea Guen .• Ent. Mo. Mag., 5.38. Hudb.. N.Z. Mothb, tI, pI. 4,7.
\\ ellin~ton, Blenheim, Rakaia.
45. P. compOBita Guen., Noct., 2, 114:; Ruds., N.Z. Moths, 22, pl. 5, 8, 9 :.
lJW&ngii Westw., Proc. Ent. Soc., 2, 55, pI. 20, 1: aAJef'8a Wa.lk.
Cat., 9, 113: maori Feld., Reis. Nov., pI. 109, 24: peracuta Morr.~
Bull. Buff. Soc. Nat. SOl., 2, 114: dentt'gela ButI., Cist. Ent., 2, 542.
North and South Islanda; common also in Australia. Larva on
grasses. I see no reason to revive Westwood's forgotton name
m face of the established use, still less under Hampson's unrecog-
nizable amended form evingi.
46. P. arotis Meyr., Trans. N.Z. Inst., 19, 11; Ruds .• N.Z. Moths. 12
pI. 4, 18: aulaciaB Meyr., Trans. N.Z. Inst., 19. 11.
Wellington, South Island.
4-7. P. atriBt'l'iqa Walk., Cat., 33, 756; Huds., N.Z. Moths, 10, pI. 4, 12'
ant!poda Fcld., Reis. Nov., pl. 109, 23.
North a.nd South Islands.
48. P. plopm Walk.. Cll.t.. 9, Ill: Huda. N.Z. Moths, 11. pl. 4, 13.
Blenheim. Mount Arthur (3,BOO £t.), Mount Butt.
;;0 ••1ItI. JllctuZa WhItt'. T.ld. N,''A Zea.l.. pI. 1 3; Huds .. N.Z. Moths. l!l,
p!' 1. 3i.
L!l.ke Coleridge.
S1. M. rnodoplellra 1\11.';\"1 .. Tra.us. N.Z. Inst .. Hl. 19, Rude., N.Z. Motlul,
19. pI. 4, 38.
Auckland, XdoplCI. Wellington. Hampson oddly umtes this and
the preceding as sexes, which is certainly incorrect, as I have males
of both. Thcy are also not only distlllct and apparently constant
in colouring, but differ somewhat in the form of the spots, occur in
different Islands, and my type of pictula has the tt-gulae distinctly
ridged, and would therefore be placed by him in a different genus
from rhodopleura, in which there seeDlS to be no ndg(', however,
on this last point I lay no stress myself.
52. JI. nquisltrr PhIlp., Trans. N.Z. lnst., 3D, 246, pI. 32, 2.
lnvercargill.
5;). Jl. plena Wa.lk., Cat.. H3. 744:; Huds., N.Z. Moths. 17, pI. 4, 32:
spnagJ!(,(/ Feld.. Reis. No"., pi. 109, 17: l'ir1i1i~ Butl., Cist. Ent ..
2.547.
Welhllp,ton, South Island. Larva on grasses and low plantb.
53A. Jl. pall(a PhIlp., Trans. ~.Z. lnst., -1:2, 54:4.
Wairarapa. Invercargill.
54 .•l:l. octans Huds .. N.Z. 1\Ioths, 25, pI. 5, 1.
Invercargill.
55. ~ll. grfJllldima Philp .• Trans. N.Z. Inst., 35, 246, pI. 3:3, 1.
Invercargill.
56. JI. ileoorata Philp., Tra.ns. N.Z. Inst .. 3i, 329. pI. 20, 2.
Inverc3rgill.
5i. J1. maya Huds., N.Z. Moths. Ii. pI. 4, 31.
Mount Arthur (3,500 ft.). l\Iclocetown.
58. 31. zanthogram1lla Me~·r .. Tranb. N.Z. Inst .. 44, 117.
Wellington.
59.•11. illsigllis Walk.. Cat .. 33, 724; Hud:... N.Z. MJthil, 16, pI. 4.. 29. 3U;
Hampe., Cat. 5, 368, pI. 88, 2(): turbida Walk., Ca.t., 33, 754:
skelloni ButI.. Cist. Ent., 2, 54:7: polllcnroo Mpyr., Trans. N.Z.
Inst., 19. 16.
North and !:!outh Islclonds. Larya polyphagoUli on low plants.
fill. J1. mtltans Walk., Cat., 11, 602; Huds., N.Z. Moths. 18, pI. 4, 34:-36 :
Hamps .. Cat .. 5, 369. pI. 38, 21: lirrniJusca "Walk.. Cat., 11, 603:
spurcata, ib.. 11. 631: veJ:ata. tb., 33, 755: anqllsta Feld., Reis.
Nov.. pI. 109, 18: acceptri.1), i'b., pI. 109. 19: debilis ButI., Proc.
Zool. Soc. Lond., 1877, 385, pI. 42, 6.
North and South Islands. LeuTa polyphagoUl:l on low plants.
en. Jr. bromias Meyr., Trans. Ent. Soc. Lond .. 1902, 273; Hamps .. Cat.,
5, 370, pl. 88, 22.
Chatham Islands.
62. M. tlstistriga Walk., Cat .. 11. 630: Huds .. X.Z. l1oths. 26. pI. 5, 20:
Hampe., Cat., 5, 377. pl. 88, 29: ligniBecta Walk., Cat., 11, 631.
North aud South Islands. Lar,"l1 on Lonioera.
63. M. paracausta Mey].'.. Trans. N.Z. lust., 19. 16; Huds .. N.Z. Moths,
15, pl. 4:. 28.
Mount Arthur. Castle Hill, Invercargill.
UEYnICK.-RtVI81011 of fh~ Xw' Zealand U,\r'l.dt"illllL,l.. 101
81. M. let·is Phllp.. Trans. N.Z. Inst .. 37, 330, pI. 20. 4.
Invercargill.
82. M. litkias Me",r.. Trans. N.Z. lnst .. 19. 17; Huds .. :S.Z. lloths. 17
pI. 4. 33. .
Castle Hill.
83. M. hcmW80ia Meyr.. Trans. N.Z. lnst., 19. 21: Huds .. N.Z. Moths
21. pI. 5, 7: Hamps., Cat., 5, 378. pI. 88, 30.
Wellington, Blenheim. Larva on Pomade1'ri8 erun/oUa.
84. M. wlI'perata Walk., Cat.. 15, 1648; Hamps., Cat.. 5, 385. pI. 89. 6:
1f1cept'IWtJ Walk., Cat .. 15. 1736: decepflu:ra. ,~ .• 1737.
Locality unknown.
85. M. pricmisti8 Meyr., Trans. N.Z. lnst., 19. 27: Hude .. N.Z. Motha,
27, pI. 5, 21; Hamps .• Cat .• 5. 384, pI. 89, 5. .
Wellington. Rakais..
86. M. phricias Meyr., Trans. N.Z. Inst.. 20, 46; Huds .. N.Z. Moths. 27.
pI. 5. 22; Hamps., Cat .• 5. 385, pI. 89, 7.
Manawatu district, BIl'nheim, Christchurch, Lake Coleridge.
Subfam. 4. CARADBINIDES.
Eyes glabrous, without marfdnal cilia; tibiae not spinose.
4. PLUSIAD.AE.
Vein 8 of hindwings shortly anastomosing with cell near base, thence
diverging, 5 well developed •
.Also an extremely large mmHy, but more espeoially ohamoteristio of
tropical regions.
Subfam. 1. HYPBNIDBS.
Hindwings with 5 nearly parallel to 4.
104 l'raIlB(lctIOIlB.
Subfam. 3. PLUSIADES.
Hi11dwings with 3, 4, 5 approximated at hase: tibiae not spiuose.
39. P. ozygramma Hiibn., Zutr., 37. f. 769, 770; trOlll.8fi,za Wlllk., Cat., 12,
884; subchalybaea, to., 33, 833.
Thames River. Widell distributed in Asia, Australia, and Pacific
islands. .
27. Opbideres Boisd.
OpMrLerea Boisd., Faun. Ent. Madag. Lap., 99 (1833); type, fullo'Nica
Linn.
Antennae in ~ ciliated. Palpi long, ascending, second joint thickened
wIth dense appressed scales, terminal joint moderately long, slender, some-
what thickened towards apex, obtuse. Thorax clothed with hair-scales
rather expanded posteriorly. A.bdomen without crests.
A rather small tropical genus, of which some species have a wide range.
100. O. fullonica Linn., Syst. Nat., 1, 812; Meyr., Trans. N.Z. Insli., 19, 37.
ChrIstchurch, one doubtful specimen. Widely distributed in Asia.,
Africa, Australia, and Pacific islands.
28. Dasypodia Guen.
Dasypodia, Guen. Noct., 3, 174 (1852); type, selenopkora Guen.
Antennae in ~ ciliated. Pa.lpi long, ascending, second joint thickened
with dense scales, terminal joint modera.tely long, slender, somewhat
thickened towards apex, obtuse. Thorax clothed with long hairs, with-
OUli crest. Abdomen without crests. Posterior tibiae densely hairy.
An Australian genus; probably of only one species.
1111. D. selenopkora Guen., Noct., 3, 175; Huds., N.Z. Moths, 35, pI. 6, 4.
North Island, Nelson, Christ~hurch. Common in south-cQst A.us-
tralia. Larva on Acacia (~).
29. Rhapsa Walk.
Rkapsa Walk., Cat., 1149 (1865); type, BCotosialis Walk.
Antennae in ~ bipectinated, towards apex simple. Palpi very loDf§.
",bliquely ascending, olothed with rough scales throughout, second joint
above in ~ with tuft of long projecting scales above towards apex,
terminal joint moderate. Th.orax olothed with scales, without crest.
Abdomen without orest. Posterior tibiae with appressed scales. Fore-
wings in ~ beneath with large broad costal fold on anterior half.
A. closely allied sp3cies occurs in south-east Australia, 80 similar that
it might be thought identical, but with the antennae of ~ furnished with
long bristles instead of pectinations, vein 8 of hindwings anastomosing
with cell to beyond middle; the characteristic palpi and costal fold of
the forewings are similar in both species.
102. R. 8OOtoswis Walk., Cat., 34, 1150; Huds., N.Z. Moths, 36, pI. 6. 5. 6 :
Zilaoina Butl., Proe. Zool. Soc. Lond., 1877, 388, pI, 42, 11.
North and South Islands. La.rva on Piper ezcelsum.
In the following indices the numbers refer to those attached to the
genera and species in consecutive order. Names italicised are synonyms.
INDEX OF GENERA.
A.chaea Hiibn. 25 Bitvla Walk. 20
Agrotis Och8. ..' 6 OhGbuatlJ, Walk. IS
Aletia Hlloo. 18 I Okloridea Westw. 4:
Alusia Guen. 18 OirphiB Walk. 13
Ariathisa Walk. 21 Cosmodes Gue'1l. 23
Austr&mathes Hamp8. 8 Dasygaster Ourll,. 19
1(6 TranMt:tloIl8.
INDEX OF G.lIlNE:RA.-contitlued.
DlIosypodJa Gut",. .. 28 Lycopkotia Hiibn. 6
Deiopeia Steph. 2 Melanchra Hiibn. 18
Dipaustica 11.'.1. 15 Metacrias Meyr. ]
Emns Walk. 17 .}[eteraM ButI. 18
Ruoa Hllbn. 5 Niiooris Guen. 21
GTammode8 Guen. 25 Nyctemers Hubrl. S
GrapkOlll.ia Hamps. 16 Ophideres Boisd. .. 27
Gmphiphora OOhs. 6 Ophiusa Ocks. 20
Heliothls Dohs. -! Persectania Hamps . .. 16
Homohadena Grote .. 10 1 Physetica Mert. a
Hypenodes Gum. . . .. 24 Plusia Ochs. 26
Hypnotype Hamps. " 9 Rhapsa Walk. 29
HyBsia Guen. 13 Side"idis Hiibn. 13
Iclm.eutica lUegr. 11 Spodopterll Guel1. 22
Lept080t'lta Boisd. 3 Tmetolopkota Hampe. 16
Leucania OenN. 12 Utetheisa Hl/brI. 2
INDEX OF ~PECIES.
acceptri:& Feld. 60 cucullina (}uen. 40
tu:etiM Feld. 12 debili8 But!. 60
acontistis ~l1eyT. ., 19 deceptura Walk. 84
admil'atiOrus Guen. 8 decora.ta Philp. 56
agomstis MCYT. n defigurata. Walk. 88
alcyone Buds. 68 dentigera But!. !5
alopa. Meyr. 24 diatmeta Buds. 65
tmg'lJ8ta Ft'ld. 60 dioneHuds. 16
annulata Boisd. 5 disjungens lYaZk. 43
anticlina MegT. 95 dotata Walk. i4
amJ.-vpoda Feld. 4; dO'l.lbleday~ Wt:ill:. 5
lSracknias M.~yr. 66 dunedinensis Hampo. 23
armigera H ubn. 6 elegans Don. 93
u.rotis Megr. 46 epiastm MaYT. 42
asterope Buds. 75 erichrysa Meyr. 2
o.tristriga Wall'. 0:1:7 eriosoma Doub!. 98
aulaciaa Meyr. 46 6tOim.gii Westw. 45
a_sa Walk. 45 exquisita Philp. 52
basinotata Walk. 7 e:uularis Mew. 94
bicom.'ltl Guen. 91 em-cmea. Guen. 31
blenheimensis F f!f. 25 /acuis Walk. S;
brf1Jiuscula. \Valk. 12 fulsidica l11eyr. 37
bromiaa Meyr. 61 fortis Butl. .. 15
ca.erult.'a Guen. 41 fullonica Linn. 100
cerQllnooes Merr. 13 graminosa Wal~·. 49
cere.unin.s Meyr. .. 17 grandiosa Philp. 55
ceropachoides GuI'Il. 91 griseipennis Feld. 34
uhaleites Esp. 98 hollandiae Guen. 87
coe1eno HtttllI. 641 homoscia Meyr. as
Walk.
L'OlIlDla 91 huttonii Butl. I
commumCGta Walk. 12 immUftis Walk. 12
composita. Grmt. 45 implno. Walk. 91
compta. Walk. 12 itteeptura Walk. 84
CtlAjeffa Walk. 6 inconspicua But]. 8
rostistrigaJis Stt"plr. 94 inconstans Blltl. 3~
!\IEYRICK.-Rt'vision of tile .V flU Zealallil Caradl·inina. 107
INDEX OF SPEOIEs--continued.
infensa. Walk.,. 66 I p"oteastis Meyr 78
mfuncta Walk. 7 pulchella Lirvn. 4
inMCua Walk. 12 pulcherrima vile. 117
innominata Hudll. 11 purdii Fer . .. 18
insignis Walk. 59 purpurea Butl. IS
iota Rude ... 15 quailrata Walk. 12
iunieolor Guell. 20 radians Gum. 7
leucanioiiles Guen. 87 r601lproca Walk. 12
levis Philp. 81 rhodopleura jJeyr. 51
lignaua Walk. 70 rogationis Guen. 98
ligni/usOG Walk. 60 rubescens Butl. 69
Ugnisecta Walk. 62 sCf1l[Jularis Fl'ld. 7
lilacmIJ But!. 102 scotosialis JValk. 102
lissoxyla Meyr. 22 selenophora Guen. 101
lithias Megr. 82 semivittata Waa·. 26
loreyi Dup. 30 s67'ioea But!. 8
maori Fold. 45 sericea Butl. 89
margarita Hawth. 92 sistens Gum. 38
mauritia Boisd. 92 skelloni But!. 59
maya Hu"",. 57 sminthistis Hamps. 38
melicertE' Drury 96 specifica Guell. 32
merope Huds. 72 sphaqnea F(.'ld. 58
micrastm Megr. 29 spurcata Walk. 60
mitis But!. .. a3 steropastis Meyr. 44
moderata Wan·. 33 stipata Walk. 71
morosa Butl. 80 strategies Huds. 3
munda Walk. i stulta Philp. 28
mutans Wall·. 60 su1Jckalybacn Walk. 99
taervata Guell. 48 suffusa Hiibn. 10
neurae Philp. 23 sulcana Fer. 27
nullifem Walk.
ollhthistis Mf'!IJ'.
. 32 tartarea Butl.
79 temenaula Melli'.
76
35
octans H1~ds. 54 temperata Wall.:. 84
octiaB Huds. 95 thoraci(.'a Wall·. 88
omicron Huds. 73 toroneura Mellr. 21
omoplaca. illegr. 67 tf'MlBfi,za Walk. 99
oxygramma HI/bll. 99 traversii Fer. 96
pachyscia Mfl/r. 36 tlJR'bida Walk. 59
pallida H udll. 90 turb"lRnta Walk. 7
paracausta Me.llr. 63 umbra Ruds. 67
pauca. PMlp. •• 58,A, unies Walk . 20
peZistis Meyr. 80 unipuncta Haw. SI
peract.lta Morr. 45 ustistriga WaU,. 62
pessota Megr. 14 'IJ61'ticillata Guell. 98
phaula. Me!Jr. 28 vezata Walk. 60
phricias Megr. 86 'I1igms Walk. 4:9
pictula White 50 But).
'I1ir68Cl''M 84
plena Walk. uS viridis But!. 58
plusiata Walk. 91 vitiosa But!. 78
poIycIwoa Meyr. u9 'Ditiosa Ruds. 79
prionistis Meyr. 85 xanthogramma JIpyr. 58
propria Wall:. 48 ypsilon Roti. 10
lOt! Tl'flm(lctloJ/~
(4:.) The reference to volume, page, plate, and figure in the catalogue.
.. Fig." means that there is a woodcut in the text: the mark ~
that there is 110 illustration of the species. When the insect
has been recognized since publication, any oLsolete reference to it
is placed in square brackets [ ].
Ann. 1tIag. Nat. Rist. = .c Annals !lond Magazine of Natural Hit!tory."
Hmpsn. = "Catalogue of the Lepidoptera-Phalaenae in the British Museum."
Ruds. = c. Ne'v Zealand Moths and Butterflies," 1898.
Subantarc. b. N.Z. = "Subantarctic Islands of New Zealand."
Trans. N.Z. !nst. = "Transactions of the New Zealand Institute."
Trans. Ent. So('. Lond. = .• Transactions of the Entomologi<:al Society
of London."
Wellington list = ,. Hand-list of NeW" Zealand Lepidoptera. Dominion
Museum. Wellingtoll. 19()!):'
.J[etacrinB hllttollj Butl.• Buds. p. ;;. h-. II 20113. Jletncria8 hlfltolli Butl. (5), vol. 3,
p. 468; fill.
Metacliaa Htrrripuira Rudb.• Hud... p. 4,* 20!l4,. M etaeriaB Btrrrtegica Buds. (3),
iv, 4 vol. 3, p. 468. ....
MllllzcriaB eljrl,ry~u lIcyr .• Hudh. p. 4.... 201.1.). lJt[atacriQ6 mcl,rYBa Meyr. (1).
h·, G vol. 3, p. 469. ....
Utetke.isa p,ikkelill Linn.. Hud... p. 3. 2088. UtetkeiBa p"[(o1I,lla Linn., vol. 3,
iv. 3 p. 483; fie.
Nycte:mfJTQ unlmlata :8000.. Hulin. p. 2. - Deile:mera a'ntlulu.ta BoihCi.t
iv, 1,2
HeliotlliB armigera Hiibll.. Hudb. p. 112. ;;6. [Ohloridea al'7lligua] Hiibn., now O.
T', 40, 41 cib8tJleta Fab., vol 4. p. 4li; fig.
Evzoa radian, Guen. , 285. Euzoa radian, Guen. (I, from N.Z.),
vol. 4, p. 164. lx, 7.
AUrali8 admiratiOili. GUt'll.• Hud'l. p. 31, I 300. Euzoa admiratiOiliB Guen., '1"01. 4,
v,37
Agroti" .arieta ButL. Ruds. p. 31, V, 38 .• J p. 173: llyn. Btricea Butl. ~.
.dg'ff1ti. cuopachoid€8 GW!'1l., Buds. p. 32, • 301. Euxoa ceTopacllOidu Guen. (0)•
vi. 1 '-01. 4, p. 174. lxi. 7.
.dgroUB tf1Irilon Ratt., Buds. p. 30, v. 646. Agrotill tf1Irilofl Rott •• vol. 4, p. 368 :
35,36 fig.
tJrtli08ia i,nmu7IiB Walk.• Huds. p. 7. ,', 211 702. A(/rotlR c(1l1&ptn Walk., vol. 4, p. 409,
lxx,ltl.
•4groti6 intlOflli7luta Hud.... Ruds. p. 31. 811ii. LyclTJlhot,ia illnOinillata Rudh. (1 j,
v.311 vol 4, p. 515; fig.
Ecto]'alrin a/lplTu WIOlk •• Wellington liSl 1123. Ectopatria aBpera Walk. (3 N.Z.),
vol 4, p. 61'>4. lxxvii, 27.
Ernlln qram,'uOBtl Walk., Rudf". p. 2\). '-, 1128. ETUfla graminosa Walk., '1"01. 5,
24.25 p. 8; fig.
JI,lu.lIcll1a rluJllopleuTD Ml'.\·r., BudJ,. p.I9, 1374. [Polia pictulu White] (3), vol. 5,
1'1", as p.174. 'If. MiB~lia pictttla White.~
JIdandlra pictrda Butl. et Huds.. IIPC (1374.1.) JIiBtlia meyn'cci Hmpsn. inetl.*
White. Buds. p. ]9, iv, 37 (2). fl.
Ltucu/lin gri.eipenui8 Feld., Buds. p. II; IJ26. H1JBsia griBtipumi8 Et'ld., T'ol. 5,
but iv, S. would appear to represent p.278. ,...
L. ,lIOIlernkJ
tThis iN placed by HampllOn in the H1JP,idae, but it is taken here for cODvenience.
is now restricted to cE'ria.in Afriean moths formerly called Otroeda, DOW plaoed
}fycts7IIera.
in thl:' Lgmtz Iitriidae.
t BamP'40D has reoently given the genlU'ic name MiBelia Tr. priority over Polia Tr.
With the imperfect matorial a.t hill dispoll&l whl"D. writing his oata.logtlE' he coDllidered the
North and I:iouth Island for0l8 sexes of White's speoiee. The type in the national col.
1eotion is thl' Bamo inaeot &8 Meyrit'k's ,.kodoplsllrG, so that name sinks. The dcsoription
ill tho ea.t&logue requires correction owing to tht> confusion of the two !IpeOies. ~
Ann. )[ag. Nat. Rist. (8). viii, P. -'21 (1911).
LONGI!TAFF.-.\'omt'ncttltul"( of tlte Lepidoptera of .v.Z. III
HIP8ia iltCOlt8tal18 Butl., Huds. p. 0, 1527. HY6Bia i1lC01l6tana Butl (3), vol. 5,
included under L. grf8Pipe7lni6 p. 279, lxxxv, 23.
LttlCGnia temeflaula :Meyr., Trans. ~.Z. (1627.&.) Huuia temenaliia Meyr. (1\. , .
1nBt. 1907, vol. 39. p. 106
J/elatlMra crllJullina GUl'll •• Hudo. p. 27, 15211.HYh8l'a woo.lli1l0 (hE'n. (4), vol. ,j,
v.23 }l.27P, !xxx, 27.
LeilCG'fIia padlYhcia Meyr., l'ra.ru.. N.Z. 11li28\.) H1J88iapachU8iaM.eyr. (0). ~.
Inst. 1907, yolo 39, p. 106.
Leueanin moderafn W&Ik., Huru.. p. 9, 1529. HU88ia modwata Walk., vol. 5,
! iv, II p. 280; fig.
Hyuia IIminfhi8llh Hmllhn., W(·llington 1530. H!J88ia 8'mi7lthiBfiB Hmpsn. (I),
list ,,01. 5, p. 2S0, lxxxvi, 17.
.Leu.cania 71vJlijera WaJk., Huw.. p. ", 1531. Hy68ia nvJlilu(I Walk. (3), vol••i •
iv, 9 p. 281; fig.
J/elanc1wa plena Walk .• Hudh. p. J7. 1671. M omBOflia ]l/plla W &lk., vol. n.
iy.32 p. 367. ~i
(lfi71A.) J1.orri80nia cM.orodonta Hmpsn.
(I !j!). ~. Description in .\nn. MaIJ.
Nat. Hi~t. (S), viii,~. 423 (lAll).
Jlelatle/tra ill8'iglll8 Walk., Hudli. p. III, 1672. MorriBO'lIia in8'ignl8 WaJk., vol• .>,
iv, 29, 30 p. 36S. lxxxviii, 20.
JlelallMra ''''UtaI18 WaJk., Huds. p. 18. 11173. Jlorril101lio mutan8 Walk, vol. 5,
iv, 34, 35, 36 p. 369, 1::axviii, 21.
jJ elane/11'a CtIewno Huw.... Huds. p. 26. (1673A.) AIorri80nia caelI!f!o Huds. (1 !j!)
iv, 39. [voL 5, p. 612, ill"ot.].
•llelanc1wa beata Howes, TraUb. N.Z. lru.t. (1673B.) Jiorl'iBOflia beata Howes (1). ~•
1906, vol. 311, p . .>11, xliv, 2
JlelancMa levi8 Philpott, Tr&nR. N.Z.
InBt. 1905, vol. 37. p. 329, xx, 4
.JlelanMra pa'l'ru:QIl8W lfeyr., Huw,. p. IS,
,.
(1073('.) JiOl'ri80nia leMa Philpott (2 ~).
t The male of this speoiea ill not known to HampllOn, who t.hinks it likeq to come
nea.r .Alopa.
~ I found this in the British HUBE'Um as M. r~, 80 deaoribed by Hampowu
in ADD. Mag. Nat. Hiat., 19O5, p. m, but he has lllince tIUJIk that nam!'.
112 TralllJartzoml.
3Ielanclira diullIIPl" Ht.tk. Hudlt. p. 21, (168!!D.) .'tlorriaoniu diatmt/u Hud,. (1 ~).
\T,5 'If.
Jlelandlrll do/11m Walk .• Blld~. p. 24, v. 1689• ..Morrisollia dotala Wo.lk. (1), vol. 5,
16 p. 380, lxxxviii, :ll.
JlelafIClIm t·",OIItt But!.. Hud~. p. 20. iv. 1690. MOrri800lu oclitJ.iKf,. l\It>vr .• \'01.•i,
42 p. 380, IXXX\"ili. :l::!. •
•11elallChm Inrtarea But!.. Huds. )I. 21. 1691. JIorri80lIia tar/(IIIII Butl. (3). vol. .i•
"f",6 p. 381, lxxxix, 1.
Jlelanekra omoplaca lIe:\T .• Bucb. p. 2:~. 1692• .J[orri80nia olll'JpT",." lll·\·r. (1).
"1". 13 vol. 5, ll. 382.lx"l:xix, 2.
JlelancMa decorata Philpott. Tre.ns. N.Z. (1693A.) Jiorri8ollin d(cQJ(/tll Philpott
Inst. 1905. "1"01. 37, p. 328, xx. 2 (4). 'If.
Jlela'1lwa li!1'1lana Walk.. Buds. p. 26, 1694. Mom8onia ligllallu Walk .• YO!. 3.
"1".19 p. 383, lxxxix, 3.
•Uelanckra pelillti" }I,,~ r .. Hud.e.. p. 19, 16115• .Morri80nia mor""a But!.. v..1. :i •
V, 3,4 p. 384, lxxxix, 4.
•11t1nnchra prioniBfia lIe.\"l'., Bud,. p. 27, • 1696. JIorri80nia pri01lllftj8 MeYI. (Ii) •
v,21 vol. 5. p. 384. lxxxix, :i.
Leucania tempe/ata Wa.lk.. Huds. p. I). -, 1697. 1l0rriBOflia temperata Walk. (3).
"1"01. 5, p. 38.), .lxxxix, 6.
.J[tlallckrn p}mc.n" lIl·~'r .. Bud.... p. 27. 1698• .l1orri8Oliia phricia8 lIeyr .• "Yol. :i•
"1".22 p. 385, lxxxix, 7.
(1698A.) llorri80llia lonurtatfi, Bow<l&.
Trans. N.Z. !nst. 1911, yol. 43,
p.128; fig.
(1698B.) .l10ITiBOIHa 8equens Howes. 'l'ra.ru..
N.Z. wt. 1912, vol. 44, p. 2<M; fig.
Jl elancltrn 1'01., JlO6ita Uuen., Huw.. p. 22. 1699. Ptraecta'llia n.jllgi WestVi., vol. Ii,
v.tI,!l p. 386; fig.
Leuoonia aroti8 Me~'T •• Huds. p. 12, iv, IS I 1700. Pll18eciallia aulae;a8 Meyr. (2).
~ia i,motaia HoWell, 'fre.ns. N.Z.
vol..5, p. 387, lxxxix, 8. Syn. uroll.
Inst. Ult/!!, vol. 40, p. J34 , lIeyr.; syn. ob8Oleta Boweb; llyn.
i'llnotata Howell.
JIelanckm Bteropaatia Meyr., Hud•. p. 23. 1701. Per8ectGnia 8teropa8h6 Men.. \'01. ii.
"1", 10, 11 p. 388, lxxxix, 9. •
LellOOllia ntrlhlriya "·aUt., Huds. p. 10, 1702. PU8ectlmia atriatriga Walk., vol . .i.
i"f", 12 p. 388; fig.
PAy.dica cae. ulee:: q'lt'U.• Kud.. p. 8, 1786. Physetica caeruleo GUl'll. (3), '\01. 5,
il.. 7 p. 445; fi@:.
1787. PkyBetica vil'ldimiali$ Uuen. (1).
vol. :i, p. 44.}, xci, 2. Ha.m}l!101l
b&yB, •• Hah. (1) U.~..A•• E. Jj'lorid.l.
lDoublcday). 1 ¥ 1.ype. Tho typt·
hal! the abdomen of a ma.lt' of IIOmt·
other Spt'Cies stuck on to it, and will
probably prove to be from Nt"I'
Zealand." Vol.:i, p. 446.
Leuoollia ""cra"'rtl }Ie~'r.• Hud... p. I:!. • 1788. PliYh£tica lIIicra!tr(l. :Mt'~·r. (0).
i"f",l0 \'01. 3, ]'. 446, :xci, 3.
JIe1ancl"a di8junuen" Walk., Hud•. p. 13. 1811i. Grapha7lia. di8ju'1Igen8 WaJ.k., \'01. ;i,
v,~ p. 4611; fig.
Llluool'lia pro'P7i(l Walk.. Hud~. 1). 11. 181tl. TmelolopilOta propria Walk., vo!. .i.
iv, 13 p. 471; tig.
LetUJQ,niIJ unipullcta Haw., HudA. p. 13. 1913. Oirpkib IllliPllIICIa Haw., \'01. .i,
iv, 24 p. 347. ~.
IM1I8l1tica Urallllitl8 lleyr.. Blld... p. l-l. '" 2036. .wuca~da ctrauniaH Me~·r. (3),
iv, 25, 26 vol. 0, p. 590.
!cAr,,:lItica dlolle Buds., Bua... ]'. 14. h'. '" 2037• .Leuca'llia diOlie Budo. (0), vol.•'J,
27 P. 590. 'II.
Leltcll7~itz acrmti8t;8 Meyr., Budd. I" 11, (2037.01..) Leucanr'a accmt1Bti. lIe;:l'T. (3)
iv, 14 [vol. 5, p. 610, ign.ot.).
Leucania t0701111ura lleyr., Tran>. Ent.,
Soc. Lond. 1901.11. G63 '" 20311• .wucania IOFOlieum lIeyr. (J).
LellCUniG IISUral! Philpo1.t, Trallb. X.z., vol. 5, p. ~1, xovi, 1. ~~-n. te~callltl
Jnst. 11105, vol. 37, 330, :xx, :i Mllrall p.wpott.
uKCQRia 1IfI1"ca Walk•• Hudl.. p. 12, iv, 2039. LeucaRia 'Unjcll Wa.lk. {iil, "01. 3,
17 p. li91 ; fig.
LONGSTAFF.-/.YomellcZatlll'e 01 thf Lepinoptol·1!. of S.Z. 118
t Mr. Meyriok (Tmus. Ent. So!l, Lond. 11101, p. 5611) conf08llll8 to ha.ving led MI.
Hudson WIItray.
114 1'ram,lIrflQn· .
At this place in lIr. Budsou's book the group that is usually called the
Geometridae follows. As Mr. L. B. Prout is still engaged in his great revision
of this group for Wytsman's Genera Insectorum" (0£ whIch one part
<t
has already appeared). it does not appear expedient to deal with them here
svstemati(.alIv. but at the samE' time it may be convenient to mention threE'
points:- . .
(1.) XMltnorhoe cineraria Dbld., Huds. p. 6i, pI. viii, figs. 2, 2A.-Person-
ally I have no doubt that the larger form is semi-signata Walk. (pl. viii.
fig. 2A) and the smaller cineraria Dbld. (pI. viii. fig. 2), ,md that these
constitute distinct species.
(2.) Lythria euclidiata Guen.. Buds. p. 68, pI. viii, fig. 35.-My speci-
mens referred by Mr. Bowes to this species agree absolutely with Butler's
type of Af'Ctesthes catapyrrha (in the British Museum), an insect, in my
opinion (and, 1 believe, in Mr. Prout'l! also). quite distinct from the
Australian euclidiata.
l3.) Bestf'a 4wIleraria Walk., Buds. p. 89, pI. x, figs. 1, 2; and Bestra
f/ezata Walk., Huds. p. 90, pI. ix, fig. 37 (very poor .figure). - Walker'b
types are in the British Museum, and I have compared them with his
descriptions. There seems no room for doubt that, bv BOme unfortunatE"
slip. Mr. Hudson has reversed the two spedes. •
(5.) Chrysop}w;nus saiustius Fabr., Huda. p. 117, pI. xii, figs. 18, 19,
20, 21; pI. xiii, figs. 2, 3, 4, 5.-Surely this name should be 8alZust,'u8: the
other spelling is meaningless.
(6.) Cnlrysophanus enysii ButI., Ruds. p. 117, pl. xii, figs. 22, 23, 24.-
The types of enYBii Butler and fS'f'eaayi Bates are both in the British Museum.
They are clearly conspecific, and Bates's name has priority. Mr. Hudson
IS in error in supposing fererJayi to be a form of 8(111ustius.
(7.) Lycaena phoebe Murray, Huds. p. 119, pI. xii, figs. 10, H.-This
IS indistinguishable from Zizera labraaus Godart, which has priority.
(8.) Lycaena o:cleyi Feld., Huds. p. 119, pI. xii, fig. 12.-This was
referred to ZizS'f'a, but has lately been placed in Nf'olllcia Waterhouse and
Tumer.
HYDRIOl\JENWAE.
Chloroclystis lunata n. sll.
a. 18-19 mm. Head. palpl, thorax, aud abdomen dark grcenish-fuscoub.
Palpl Ii. .Antennae biciliated with long fascicles. ciliations 3. Fort'-
wings triangular, costa almost straight, termen slightly bowed, subsinuate
on lower h.a.lf; dark greeniih-/uscoU8; veins marked more or less with
black; lines obscure; some faint thin waved green lines near base;
median band ochreous except beneath costa, anterior edge from -i to =,
waved, hardly curved. posterior from 1 to t. bluntly projecting at middle
and concave on lower half; a thin dentate bluish - green subterminal
line: cilia. ochreous. barred with fuscous. Hindwings fuscous. sprinkled
with ochreous; veins with alternate black and white dots; a thin dentate
bluish-green subterminal line.
2 as ~, but median band almost obsoleto. Ilnd with prominent irre-
gular crescefttic white mark in miililll' 0/ /oT6'IlIing. the limbs du:ected
posteriorly.
Wallauetown. in December and January. .A reddish-bl'OWll mT\'a fOUlld
feeding on Veronica on the 5th February pupated a few days later and·
emerged 80S a ~ moth early in the following December. Unfortunately a
fuller description of the larva. was not secured.
The species differs from most of its allies in its darker ground-colour.
and in the ~ the white crescentic mark is " I!;ood distineti'\"e character.
lUi TrWU,llctIOIiI.
l'RA~IBIJ)AE.
Orocrambus subitus n. i>p.
~. 15-16 mm. Head dark brownish-{uscous. Palpl dark brownish-
fuscous, whitish above. Antcnnae fuscoUII. Thorax dark brownish-fuscous
"ith white lateral IItripc. Abdomen dark browmsh-fuscous. Forewings,
costa straight. apex obtuse, termen slightly oblique. golden bro\vn sprinkled
with white scales, densely il'rorated ",ith white in disc and along dorsum.
white irroration produced as a streak from disc to apex, dorsum narrowly
black from base to i; a thick black central streak from base to ~, apex
obtuse, margined beneath with golden brown; a short black streak above
apex of basal streak, anteriorly tapered; a few black scales before tornUl:l
indicating a black subterminal line: cilia dark golden brown with obscure
darker line. Hindwings dark brownish-fusoous; cilia brown, paler round
tornus.
;!. Forewings with 'IJ.'hite irrofation l'xtending to costa,' blu.ck streakll
as in 13. but margined 'With orange: cilia grey mixed \nth \vhite. Hind-
wings. grey, paler round termen; cilia grey.
Hump Ridge (Okaka); fairly common at 3,500 ft. in December.
~earest to O. tMmiastiB, hut differing from that species in the colour
of the streaks in disc, w'hich are white or ,ellow in thimdastis, black in
subitUR. .
PYRAUSTIDAE.
Scoparia clavata n. hp.
5. 26 mID. Head and thoI'ax white. with a black lateral stripe from
eye to near middle of thorax. Palpi moderate, white above, sides and
beneath dark-brownish. Antennae and abdomen grey. Legs grey, ante-
rior pair sufiused with fuscous. Forewings moderate, posteriorly dilated,
costa almost straight. apex rounded, termen subsinuate, oblique; white,
irrorated with brownish-ochreous. costa narrowly brownish; a thick black
median streak from base of costa to lltlmost t, slightly constricted near
termination, apex rounded; a thick black streak in disc above middle,
ilTegUla.rly sinuate, beginning before i and ending at about! in irregular
dilatation; tJ, subterminal black striga, ilj,wardly oblique aM. rUlated beneath
c:fl8ta and abol)C dorsum,' all streaks margined with brownish-ochreous; a
terminal chain of linear black dots: cilia whitish 'with t,,·o grey lines.
Hindwings shinin~ ,vhite. ochreous round termen: cilia white, ochreous
11tlar apf'x.
Hump Ridge: ill £01'~st, at 3.000 ft .. in Decemb~r; Olle specimen.
Easily distinguished from S. rolueUa. its nearest ally. by the subterminal
black strigae: it is also hroader-wiIl!led than that species.
t.h'fRICK.-j)t!8rrlf!tiulI~ II! -,"tlj' Zealalld Lepidoptera.. 117
CRA~lBmAE.
Orocrambus pervius n. sp.
c1. 25 mm. Head, pa.lpi, and thorax dark fuscous, palpi mixed
beneath with whitish hairs, shoulder with a slight white mark. Antennae
shortly ciliated (1). Abdomen dark grey, apex whitish. Forewings elon-
gate, posteriorly dilated, costa hardly arched, apex obtuse, termen rounded,
somewhat oblique; dark fuscous, irregularly strewn or partially suffused
with ochreous-brown scales; costal edge slenderly whitish on median area ;
a rather DArrow white median longitudinal streak from base to termen,
beyond middle shortly attenuated or interrupted: cilia grey, with a white
bar on median streak. Hindwings dark grey, pectinations ochreous-whitish;
cilia ochreous-whitish, basal third grey. Hindwings beneath largely suffused
with ochreous-whitish.
Lake W'akatipu, 3,600 ft. (Hudson), in February; two specimens.
01....sely allied to catacaustus, which, however, is browner, with a white
shoulder-stripe (not mentioned in my description), and with the median
btripe only seldom showing a tendency to interruption; but the reliable
distinguishing character lies in the form of the forewings, of which in CQta-
(J(JUl)tU8 the termen is not oblique on the upper portion.
PYRAUSTID.AE.
Scoparia triscelis Merr.
This distinct species, origJ.D.J.lly described from Auckland Island (•• Sub-
antarctic Islands of New Zealand," p. 71), has now been found b", Mr.
Hudson at Lake Wakatipu: a very interesting observation. .
Scoparia locularis n. sp.
d. 21 mm. Head ochreous-whitish. Palpi 21, dark fuscous, basal
joint white. Antennal ciliations~. Thorax white .mixed with grey and
dark fuscous. Abdomen grey. Forewings elongate-triangular, narrow at
base. costa posteriorly moderately arched, apex obtuse, termen sinuate.
oblique; lip:ht grey, irregularly .mixed with white, with some scattered
black scales: 1m oblique streak of black suffusion from base of costa; first
line white. rather oblique, slightly sinuate, posteriorly strongly edged with
black suffusion; oTbicular and claviform represented by elongate marks
MEYRIOK.-DeBcrzptionB uf IfelL Zt'fl/allt/ LppidoptOl a 119
blackish basal line on termen of second segment; ciha round tornus and
011 dor&um fusrous. Hindwings rather dark fuscous; cilia grey, base
grev-whitish.
Lake Wakatipu (Hudson), III February. This is an interesting re-
discovery. as F{'lder's species had never been recognized before, aDd his
localiti{'s ;ue frequently quitt' errOllE'OU!>. It is allied to charatlrias.
Stenoptilia zophodactyla Dup.
lh. Hudson sends a specimen of this nearl) cObmopohtan species,
taken near Wellington in November. stating it to be very rare. It has
not been previously recorded from New Zealand. It is common in Europe,
which is probably its place of origin; but I have also received it freely
from India. Australia, South Africa, and South America. The larva feeds
on E,ythraea. but must also be attached to other Gentianaceae, and is
probably artificially introduced. It may have reached New Zealand only
quite recently.
CARPOSI:>''1D -I.E.
Carposina morbida 11. bp.
e!. 26 mm. Hea.d ochreous-whit.ish. Palpi 2~, porrectetl. ochreous-
whitish. ba.sal half suffused with dark olive-grey. Antennal Cllmtions 4.
Thorax ochreous-whitish, shoulders with an ochreous spot. Abdomen
ochrt'ous-whitish. Forewings elongate, rather narrow, posteriorly some-
what dilated. costa gently arched, apex obtuse. termen straight, rather
olllique; silvery-whitish-ochreous, irregularly strewn with ochreous lIcales.
('osta and dorsum somewhat sprinkled with grey; a small brownish-
ochreous basal patch, edge parallel to termen; six small shots of
grey su:liusion on rosta bt'tween this and apex; tufts brownish-
ochreous. posteriorl~' white-viz., one bent'ath costa at -1, preceded
by 1.1 dash of bla.ck irroration, one beneath thi'! towards dorsum, ~
In..-gllr Ohe lD disc beyond these, preceded h.... a black dash o.a sub-
median fold. two towards costa in and beyond middle edged with
black beneath. one below middle edged with black above, snd a ridge
on t1'8ns'\'"erse vein, irregularly edged with black anteriorly, between
thesl' in middle of disc is an elongate patch of grey sufiusion; some
scattered black irroration crossing wing at i; cilia whitish, with two
greyish shades. Hindwings and cilia whitish. Under-sumer of fol"t.'-
wings and hilldwings largely elothed on anterior half with modified
paIli' yellow-ochreous scales, on forewings anteriorly su:liused with grey.
Lake Wakatipu (Hudson), in February; one speeimen. Ca.n only
be confused with exoc1uma .. but the c1 of that spe<'ies has much longer
porrected palpi (4).
TORTBJCIDAE.
Harmologa tritochlora n. sp.
~. 22 mm. Head and palpi pale ochreous, palpi 3. Thorax whitish-
ochreous, patagia sufiusedly mixed with grey. Abdomen ochreous-whitish.
Forewings elongate-oblong, costa. moderately arrhed towards base, thence
nearly straight, apex obtuse, termen slightly rounded, somewhat ob-
lique; whitish-grey suffused with pale brassy-yellowish, becoming whitish-
yellowish towards costa and termen, irrorated with darker grey on dorsal
half towards hase: cilia whitish-yellowish. Hindwings and cilia creamy-
white.
MJ<}YRWK.-IJescrlptiollll of lltw ZUlZulld Lepidoptem. 121
Lake Wakatipu, 4,000 ft. (Hudson), III Febrwuy; one specunen. Allied
to BiraM and aenea, but cannot be united with either; doubtless the ~,
as in those species, is very different, with dark hindwings.
OEC'OPHORIDAE.
Izatha metadelta Meyr.
Mr. Hudson suggests that pe:ronitis MeYl. IS the ~ of this &pecies, and
on consideration of the available material, having now several specimens
of each form, all the peronitis being ~ and all the metadelta !jl, I haYe no
doubt his view is correct, and they must be united as sexes.
GLYPHIPTERYGIDAE.
Simaethis zomeuta 11. sp.
!jl. 18 mIn. Head and thorax bronzy-brown, sprmkled With whitIsh.
Palpi with whorls of blackish white-tipped scales, base white. Antennae
dc\rk fuscous dotted with white. Abdomen dark fuseous. Forewings
elongate, posteriorly dilated, costa gently arched, apex obtuse, termen
bowed, rather oblique; bronzy-ochreous-fuscous; some violet-white irro-
ration towards base and termen; a moderately broad darker median
transverse fascia, angulated above mIddle, edged by obscure shades of.
violet-,vhite irroration, hecoming more distinct on costa, where the second
forms a clear white oblique mark: eilia bronzy-ochreous. with two dal'k-
fuscous shades, and white patches on outer half above and below middlp.
Hindwiugs bronzy-fusco.us, becoming blackish 011 posterior half; a rather
incurved white streak crossing dorsal half of win~ from ! of disc to toruus.
and some whitish irroration between this and termpn; cilia ,vhitish, basal
third dark fuscous.
)lount Arthur, 4,600 ft.; one specimel1 taken by myself in JanUd.ry,
not in fine condition, but twenty-five years have passed without further
captures. Near oombinatana, but distinguishable by clear white streak of
hindwings, and the joints of alltt>nnae are- rt"latively lnuch more- elongate
and slender.
Simaethis ministra 11. sp.
a. 9 mm. Head dark fuscous, face spnnkled WIth white. Palpi
v,ith whorls of dark-{UtlcoUtl white-tipped sca.les, base white. Antennae
dark fuscous, shortly pubeseent-ciliated. Thorax dark fuscous. Abdo-
mt'n dark fuscous, st'gmental margins partially white. Forewings sub-
oblong. costa mod<>rately arched anteriorly, a.pex: obtUSf.'. tenneu slightly
rounded, somewhat oblique; dark bronzy-fuscous; five very undefined
irregular transverse shades of whitc irroratioll, first three rather curved
or bent, fourth forming a clear white spot on costa be~·ond middle and
then a fine silvery quadrangular loop passing behind a transverse linear
discal mark of white irroration, fifth straight, interrupted above middle;
two or three silverv-metallic scales before termen above middle: cilia.
white with dark-fUscous shade (imperfect). Hiudwings light fusoous.
becoming darker towards termen; dorsal half with scattered whito scalE'S ;
a well-marked irregular white streak extending across dorsal half of wing
from disc at ! nearly to tornus. its lower half approximated to termen;
cilia white, with fuscous subbasal and post-median shades.
Mount Holdsworth (Hudson); one specimen. This and the two fol-
lowing species are closely allied and very similar. agreeing toge-ther in
122
haymg the antennae of a shortly pubescent-ciliated, whilst In S. tllarmarea,
which is also very l:Iimil.ar superficially. they 11.1'<' ciliated with long
fascicles (3). ns is usual m the genus. S. m1'mst'l'a differs from the
other t,,'o m huying the antennae wholly dark fuseous, the forewingt.
obviousl, broader anteriorly. with costa more arched than in either of
the others. the whIte markings of forewings less defined. tho white
&treak of hindwings bl'Oad(.'r and more irrE'gular. I'eaching termen at
t trom apex and contInued almost to tornus.
Simaethis microlitha Yep.
a
Q. 9-10 mm. Head and thorax llToratcd WIth wlute abo"\'e: scaletl
of palpi longer and more projecting than in (VI'uiloga " antennae dotted
with wrote: abdomen with segmental margins strongly white. Fore-
wings more narrowed towards base than in miniBt'l'a. fasciae of white irro-
ration more strongly marked. fourth slender, but more sharply marked
and brightly sil,ery-metallic above discal mark and at apex of dorsal
section. Hindwings with white streak slender, regular, well-marked, ex-
tending t across wing from ! of disc to middle of termen and thenre
running partially interrupted near termen almost to tomus.
Arthur's Pass, 3,000 ft., in January; two sp(.'cimens. In my descrip-
tion of this species I included also the follo"ing, which I now separate
from it. and therefore specify the more characteristic points which diB-
tinguish the true ftticrolitha from the preceding and following species;
the character of the marking of hindwings is the most obvious of these.
Simaethis analoga n. sp.
O'~. 8-9 mm. Head dark fuseous, face and sides of crown lrrorated
WIth white. Palpi with whorls of dark-ruoous white-tipped scales, base
white. Antennae dark fuscous dotted with white, in 6 shortly pubescent-
ciliated. Thorax dark fuscous, somewhat sprinkled with white. inner
edge of pata.gia white. Abdomen dark ruoous, segmental margins strongly
white. Forewings rather elongate-triangular. costa gently arched, apex
obtuse, termen alightly rounded, somewhat oblique; dark bronzy-fusoous ;
three CUX'\'"ed cloudy transverse lmes of whit!' irroration on anterior half.
two posterior sometimes irregularly confluent; a white line beyond middle
forming s quadrangular loop behind a transverse-linear white discal mark,
upper side of loop silvery-metallic, lower absent, a silvery-metallio dot
on upper extremity of dorsal segment; a straight cloudy line of white
irroration from ~ of costa to tomus, interrupted above middle j a trans-
verse silven--metallic mark before termen above middle: cilia white with
two thick dark-fnscous lines, and dark patches at apex, middle of termen,
and toxnus. Hindwings ruoous, beooming dark fusoous posteriorly j a
very short white detached transverse mark before middle of termen, and
sometimes a dot on toxnus; cilia white, with two thick dark-fusoous lines.
Mount Arthur, 4,000 ft., in Jannarv; ten specimens. As explained
above. I originally regarded this as a furm of w'cro'fIi:IAa, but now think
it distinct. Doubtless more species of this group will be discovered in
the mountains. and an attempt should be made to ascertain their food-
plants.
Ereunetis acrodina n. sp.
~. 14 mm. Head whi1.ish-och.reous, hairs of forehead slightly mixed
with dark fuscous. Antennae ~y-wb.itish, basal joint with a blackish
MEYRICK.-lJescriptions of New Zealalw Lepidoptera. 123
spot. Palpl whitish, second joint streaked With dark fuscous aboye and
beneath. termmal joint dark fusc'ous towards base. Thorax whitish-
ochreous. shoulders with a dark-{uscous spot. Abdomen ochreous-whitish.
Forewings elongate, narrow, costa moderately arched; apex round-pointed,
upturned, termen extremely obliquely rounded; greyish-ochreous, with a
few dark-fuscous scales; markinp;s fuscous mixed with blackish; four
oblique patches from costa, more or Ipss confluent with a broad irregular
submedian streak from near base to apex. first near base. second broadest,
before middle, third narrow, fourth reduced to a streak; an irregular dark-
fuscous apical spot surrounded with white: Clba whitish, with an inter-
rupted black subbasal line, and fuSCOUB post-median line, tips fuscous at
apex. Hindwings grey-whitish; cilia whitish, at apex With two dark-
grey lines.
Wellington (Hudson); one spccimel l • Iutel"mcdidte between prebil'lfill
(which has hitherto stood rather isolated) aud '''''lUI
itel1n.
Taleporia Rb.
The genus Ta1eporia haa not previously been identified fl"OID the
Southern Hemisphere, but the following species agrees fully with it,
except that veins 7 and 8 of the forewings are separate, whel'eas in tht'
typical European species they are stalked; in this group, however. this
character is of little importance, and I have no hesitation in regarding
the species as a true Talep()ria. a. very intt'restinp; discoyery.
Taleporia aphrostcha n. sp.
<1 22 mm. Head, palpi, snd antennae dark fuscous, antennal clllll.-
tiona 2~. Thorax dark fuscous, with several whitish dots posteriorly.
Abdomen dark grey, somewhat whitish-mixed. Forewings elongate, rather
narrow at base, posteriorly dilated, costa gently arched, apex obtuse,
termen obliquely rounded; all veins separate; white, mixed "ith grey
m disc and towards costa, coarsely reticulated throughout with dark
fuscous; the white colour forms a more conspicuous quadrate spot on
dorsum before middle, including a dark - fusrous dorsal strigula, and
preceded and followed by irregular dark-fuscous spots: cilia fuscous,
basa.l half spotted with white. Hindwings dark grey; cilia fuseaus.
!j1 apterous, active.
Hump Ridge. Invercargill, 3,500 ft. (Philpott); a pail' in De{'emhel'.
HEPIALIUAB:.
Porina copularis n, sp.
~ 38-40 mm., !j1 44-50 mm. Head and thorax pale ochreoUti, some-
times partially tinged with fusoous. Antennae in ~ shortly bipeetinated
with fiattened-wedgc-shaped teeth. (2), Abdomen pale ochreous, in !j1 in-
fusca.ted except anal tuft. Forewings formed nearly as in wnbraculata,
but costa more amuate; pale ochreous, in !j1 tinged with fuscous; a white
dot finely edged with dark fuscous in disc at !, and an elongate mark
beyond middle; in ~ sometimes a smaller dot beneath submedian fold
rather beyond :first; a faint pale irregular bllluate transverse shade at f,
sometimes marked with 8. few indistinct fuseous strigulae, and sometimes
a. series of indistinct fuscous dots beyond this: cilia whitish-ochreous,
barred with ochreous or greyish-o{'hreous. Hindwings pale flw>ous tinged
with ochreous; eilia 88 in forewings.
West Plains, Inwreargill (PhIlpott); five spe{'imens.
124 TranRaction~.
XICROPTERYGIDAE.
Sabatinca Walk.
This generic name supersedes Palaeo'micfa Meyr., but I haye formed
a new genus. JlJicropa'l'duliB. to contain i/,(Jrozena Meyr.
Sabatinca caustica n. sp.
1. 9-10 mm. Head and thorax bronzy-orange-ochroous, thorax some-
times marked with whitish. Antennae ochreous, towards apex blackish.
Abdomen dark purple-grey. Forewings ovate-lanceolate, costa moderately
arched. apex pointed, termen extremely obliquely rounded; violet-coppery-
ochreous. in one specimen largely suffused with whitish; in one specimen
a spot of da.rk purple-fuscous suffusion on dorsum. towards base, one in
disC' beyond middle. and some irregular marking towards termen. and in
the whitish-suffused specimen the dark purple-fuscous suffusion forms a
blotch alonp: anterior portion of costa connected with a large oblique blotch
in middle of disc. a streak along dorsum from base to f. a subterminal
fasc1a enclosing a white llpot on costa. and a mark along termen in middle,
but in the other two specimens there a.re no markings: cilia golden-
ochreous_ Hindwings deep purple; cilia pale golden-ochreous.
Seaward Moss, Invercargill. in October (Philpott); four specimens.
The amount of variation is remarkable. but all the specimens were taken
together. a.nd are undoubtedly the same species: the shape of forewings
is oharacteristic. being more pointed than in a.ny other species. The species
may be placed between ZOt'IOtloza and M'1IBm'gyra.
Sabatinca incongruella Walk.
This na.me supersedes ckalcophanea Meyr.
Sabatinca calliarcha n. sp.
~. 12 mm. Head lliI;ht bronzy - ochreous. hairs extremely long.
Antennae pale ochrE-ous ringed with dark fuscous. Thorax clothed
Mnmf'K.-lJesrrzpflOIlS of .rtll' Zea[tl/u/ Lupiuopte)·.L 125
WIth long bronzy-ochreous ham" benE'ath which 1& a whIte Lent stripe
on each sidc of back. Abdomen blackish, apex ochreous - whitisl).
Forewings elongate-ovatc, costa moderately arched. apex obtuse, termen
very obliquely rounded j yellow j dorsum suffused with ferruginous-
brown, with H few black scales on cdge j four golden-whitish streaks
from costa between base and ~ converging towards posterior half of
dorsum, first edged po&teriorly with ferruginous-brown mixed with indigo-
black, hardly rea.ching dorsum. other three margined on both sidl's with
ferruginous - brown streaks and on costa. with black, second and fourth
reaching dorsum, third rea.ching a.bout half across wing; posterior area
ferruginous-brownISh somewhat mixed with pale yellowish, with an me-
gular black dot in disc ali i. and four black dots on costa edged benea.th
with golden-whitish j a thick black streak lying along termen from near
apex to tomus, edged with ochreous-yellowish a.nd interrupted to form So
long upper a.nd short lower portion, upper portion including two ~olden
meta.llic terminal dots; cilia light ochreous-yellowish, with a violet-coppery
basal line edged externa.lly with grey. Hindwings deep purple. disc and
veins blackish j cilia blackish-grey.
Bluecliff, Inverrargill, in December (Philpott); one fine specImen.
This is a beautiful and remarkably distinct species, showing some super-
ncial a.pproximation to Mioropardalis aor(X1;ffl(J, but structurally a true
Sabati'l&Ca in all respects. 1 regard it. however, as the earliest form of
the genus. I entertain no doubt that other forms of this primitive family
remain to be discovered in New Zealand, and, as they are amongst the
most important and interesting elements of the fauna, it is very desirable
that collectors should make special efforta to find them. Probably the
larvae feed on da.mp mosses, and Conifer forcsts a.rc the most likely
locality, especia.lly in the early part of the season, perhaps before
collectors usually take the field. The perfect insects fly in the sunshine,
but in partiall~' shaded places, and arc sometimes extremely difficult to
see.
ADDE::-l'DUlI.
Since writing the ahow. additional matlmal h&i> bcrll submitted to me,
which includf>s thl' two fonowin~ species ;-
Eucosma querula n. Bp.
S~. 21-28 mm. Hcad. palpi. aud thorax dark fuscoU&. Abdomen
fuscous, not hairy. Forewings elougatc-triangular, costa. gently arched, in
r1 with very short and narrow basal fold, apex obtuse, termen rounded,
rather oblique j purplish-bronzy-fuscous suffusedly mixed and strigulated
with dark fuacous j costa obscurely palc-strigulated on posterior i; a
whitish or ochr(.>ous-whitish dot in disc at : j two or threc variable curved
transverse series of small dark-fuscous spots or dots betwc.>en this and
termen ; cilia fuscous. with darkt'r line near base. Hilld wings fuscous.
posteriorly sometimes faintly darkl'r-strigulated: in c3 without speoial
cha.raeters ; cilia pal(' fuscous.
Christchurch and Wellington. in April (PhIlpott, Hudson) j four specimens.
I have also two !j! from Queensland which I refer with little doubt to this
specipsj I suppose it to be indigenous in Australia (and very likely in some
of the Malayan islands), and to have been reoently introduced into New
Zealand. It belongs to a group of several Indian and Malayan speoies
which are almost exactly alik(' in superficial appearancl'. hut possess good
126
charactl'r:. for disCliminatlOll III the secondary sexual structures of the cr
-viz., the costal fold of forewings, the folding and tufting of the dorsal
margin of hindwings, and the plesence of hairy tufts on the:> abdome:>n.
Sabatinca quadrijuga n. sp.
~. 13 mm. Head pale-greYlsh. Antenlllle dark fuscous. Thorax
purplish. Abdomen grey, lateral claspers and bupraanal projection longer
and narrower than in caustt'ca. Forewings ovate-lanceolate, less acute than
in ca'U8tica, stalk of 7 and 8 extremely short; deep purple, irregularly mlxed
with coppery-golden, darker and bluish on costa; four subquadro.te
ochreous-whitish spots on costa between base and 1, larger anteriorly, and a.
dot towards apex: cilia grey-whitish, with several dark-grey bars. Hind-
wings violet-grey, darker towards apex; cilia grey-whitish, 011 costa barred
with grey suffusion.
Invercargill (Philpott); onE' spE"Cimen received through thl! kiudnt.-ss of
lit. Hudson.
Fam. LIBELLULIDAE.
Subfam. LmBlLLuLINAE.
1. Tramea sp., 1 !F (label No.2).
Somewhat immature, and of pale rolOl'&tion. The dark patch at the-
base of the hindwings is exceedingly 8lllall. and does not spread down-
wards into the anal area8 of the wing. The male of the insert should be
obtained, as from the form of the ~ appendages and p:enitalia. the species
could be determined with certainty. I am of opinion that this will prove
to be & local race of a. widely distributed oceanic species. The genus
is hlghly migratory. and one species is rapidly travelling down the east
coast of Australia, and getting a strong hold there.
Subfam. COB.DULIINAE.
2. Hemicordulia australiae Rambur (label No.3).
cr.
2 ~. 2 In good condition. & dark tt.lld handsome form, practically
identical in size and colouring with the speoimens found in the Sydney
distriC"t. This beautiful species. recogniza hIe by tht' brilliant metallic-green
TILT,n.R]) -/)rfl{loll-fhr, from tllr K rrmodec [,taMII. 127
frontal pa.tch on thc head. and the sharp spille on the underside of
the malc appendagcs, has never before been recorded outside Australia.
It rangcs {rom Victoria, through New Routh Wales, to northern Queens-
land, but does not occur west of the ma.in mountain-ranges. Its capturt'
in the Kennadcc Isla.nds IS therefore of conslderable interest. The genus
Hemicordulia is post-Miocene, so that the oocurrenoe of this species may
be taken as eVldence of la.te la.nd connection between Australia and the
Kcrmadecs. The species IS non-mlgratory, and does not oCCur in Tas-
mania, though exceedingly common on the northern short'S of B~s Strait.
The inference, therefore, IS that the Kermadecs may have been united
lD some way, possibly via New Caledonia and Queensland. to Australia
since the time (' Miocene) when Tasmania became separated. We should
also expeot, possibly, to find this species on Lord Howe a.nd No:rf.olk
I'liands. whose Odonate fauna llote still unknown. 1i
Fam. AESCHNIDAE.
Sub£a.m. Al!:SCHNINAE.
3. Hemianax papuensis Burmeister (label No. 1, ~). J
l~. Immature. but a nne specimen (appendages broken). Common
all over New Guinea and AustralIa, except Tasmania, where it is absent.
A strong flier, but not migratory. This reinforces the evidence of No.2,
HertWmaa: also being a Miocene or post-Miocene genus.
4. Aeschna brevistyla Rambur, 2 ~ (label No. 1, ~).
One specimen immature, one mature. Tllis inseot 18 found a.ll over
Australia, except in the most northern pa.rts. It also oocurs in Tasmania.
In New Zealand a somewhat smaller and darker form oocurs, which, though
-clearly conspecifir with the Australian, may be distinguished at once from
it by its abdomen being exoeedingly pinched at the third segment, and
its membranule verv dark. with only a. little white at the base. The
Kmnadf'(' specimens' are idf'ntical with the New Zpa.land form.
Fam. AGRIONIDAE.
Sub£a.m. AGRIOYINAE.
5. Ischnura aurora Br,ner (= I. aeUoata SelY<I). (lnhel No.4).
5 &" 6~. A very beautiful species, of wide distribution, ranging
froni the islands nortJJ. of Australia, through Austra.lia, to Tasmallia. In
Western Australia a dimorphic female, coloured. like the male, occurs.
The male has a bright-red abdomen shading to black. with a blue tip;
the ordinary £emaIl' is dull-blaokish. These Kermadeo specimens are
practically identical with any series of this insect taken round Sydney.
In conclusion, therefore. thIS small (.'Olleotion shows the Kermader
Odot&ata to have a strong Austl'8.lian element (three species out of five), "
New Zealand element (one species), and an oceanic eleml"nt (one species)
128 Tramartlons.
TIDb short paper contains a few miscellaneous notes that helve been made
durmg recent years on some New Zealand C'I'ust{l,(l('a. Though there are
many othet questions that requir(' to be settled, and several groups that
need thorough re'\;sion. it has been thought "'orth while publishing thC8~
few notes as they stand. though they are necessaruy somE'what discon-
llE'cted. donO. 0.('11.1 'nth s('sttered members of the Ctwdacl'a.
Order DEC'APoDA.
Hymenosoma lacustris Chilton.
Elrltluna \~) lacustril> Chilton, Trans. N.Z. Inst., vol. 1-:1., p. 172,
pI. 8. 1882. H?(,menosoma lacustris Chilton, Truns. N.Z. Inst.,
vol. 15, p. 69, 1883; Fulton and Gl·snt. Proc. Roy. Soc. Victoria.
vol. 15, p. 59. pI. 8, 1902; Chilton, P.Z.S. for 1906, p. 'i03, 1906.
This lWlaU fresh-water cl'&b was originally descri1ed ll'ODl Lake
Takapuna (or "Pupuke "), near Auckland, which is quite nea.r the sea-
coast. and for a long time this was the only locality from which it
was known, and it was a little uncertain whether it ,vas a genuine fresh-
water form or a relict species that had only comparatively recently dE'-
yeloped in Lake Takapuna. In 1902, however, Messrs. Fulton and Grant
recorded the spedes from Lake Comc, in Yictoria, and about the Sd.me
time I received s~yera.l specimens n'Om ~orfolk Island. Specimens
from a.ll these localities were examined by Messrs. Fulton and Grant,
and, although there are a. few shght differences, thestl were found to be
not constant. and they decided to consider all th... forms as b...longing to
the one species.
In 1903 two specimens of the crab were found by Messrs. Hodgkin
and Lucas in Lake Waikare, in Auckland, which is a considerable distance
from the coast; and in the early part of this year (1911) IL few specimens
undoubtedly belonging to the same species were sent to me by Mr.
Cheeseman from the Waipa River.
It seems evident from the above facta that the species is a widely
distributed inhabitant of fresh waters, and its occurrence in the fresh
waters of :Sew Zealand, Norfolk Island, and Victoria. presents a problem
of some interest in connection with the geographical distribution of the
C'1tlstacea. In connection with this point, it is, however, worth while
stating that the fresh-water shrimp in Norfolk Island and Victoria is
Xip1tooaris compreSBIJ De Haan, and is quite different from the species,
."t. cv.rtJi708tris Heller. which is found in nea.rl)· all the fresh-\vater streams
of New Zealand. and occurs also in tbe Chatham Islands.
UHILTON.-Note, on oomt .New Zealand CruEotaC6a. 12U
Ol'dN' AMPHIPODA.
and its inner wdorgill ('1l1111Ot l:>e clea.rly speno but it Il.ppeal's smooth then'
also. In his original des(mption l'.!tpbhill~ descriues the telson as haviug
.. the minute apex microscopically trid('utate," and figures it all distillrtl~·
tmlputate: in the .. Das Til.'tl'eich" uetlcription h(' simply says, •. apex
a. little obtuse," which perhaps more accuratel~- dellcribrs thc u.ppearlll1(·(·
(If rl.o telson in those specimenl! that I havp pxamine(l.
Hab.-Ha\U'aki Gulf (25 fathoJlll!). Paterson Inlet (10 fathoms). Tllk('n
ILIS(I III Nt'w ZPaland tleas by the .. Challeng('t" (2.000 fathoms).
This variety differs from the typical form of the species in t~le longer
uropods, which are fully half 8S long as the pleon; the peduncle 18 shorter
than the rami, and may be slightly dilated at the distal end; the outer
ramus is almost or quite as long as the inner, but slightly more slender.
and has long setae, usually at the end only; the inner ramus has long
setae both at the end and at a point some distance from the end.
Hob.-On Sphaeroma quoyana, Marlhorough Sounds and Hawke's Bay.
Also on specimens of the same species mm Sydney Harhour.
I have had specimens of this variety for severol years. The dif-
ference between it and the typical form of the species is sometimes so
distinct that I have at times almost been inclined to give it a different
specific name, especially as it appears to be always associated with a
difierent species of Sphaeroma. I find, howe-veT. that Inis pubescens found
on Spkaeroma giqas shows considerable variation in the length of the
uropods; I have one specimen from Lyttelton. which has them much
longer than usual, and approaching the condition found in the variet~·
now described, while others from Port Chalmers have the uropods much
shorter, with the outer ramus very small and only about half as long as
the inner one. I can, moreover, find no constant points of difierence
except in the uropoda, and therefore prefer to look upon the form found
on S. quoyntla as merely a variety of the species.
CluBB HVDROZO.\.
Order SIPHONOPHOR.\.
Physalia utriculus Eschscholtz.
Lesson, Voy. de" Coquille," vol. 2, pt. 2, chap. 15, p. 39 : Zoophytes
pl. 5, fig. 2. Haeckel." Challenger" Reports, 28, p. 351.
Cast ashore 011 Denham Bay, Hunda" Island. WidE-h' distributed in
thE' Pacific. •. •
186
Class SCYPHOZOA.
! Atolla br.
A single somewbat torn and wstorted specnnen. measuring 30 mm.
in diameter, with a heigbt of 15 mm. in the centre of tho umbrella, was
found on the shore of Sunday Island. It was so much injured that I am.
not quite sure even of the genus; but it agrees in so many fea.tures with
Atolla that I haw but little hesitation ill placing it here. I will not, how-
ever. nttpmpt to ~,\"E' a specific name to it.
l'lctRb H OLOTHUROlDEA.
Actinopyga (Muelleria) parvula SE'lenka..
JI. !lal:o-ca8lanea Theel: ~elenka, Zeit. Wiss. Zool., 17, 1867. .. ("'h.\l-
lenger ,. Reports. Holothuroidea, pt. 2, p. 198, 1886.
Fdtl'en speCImens were sent to me. Oliver notes that the .. colour
is dark brown to nearly black: common at Coral Ba.~·, under stones near
low-water mark; not seen elsewhere." In alcohol it is chocolate-brown
with a purplish hue. The majority are uniformly coloured, darker dorsally
and only slightly paler '\"'entrally; but in four individuals there is an
abrupt transverse line separating the dark anterior region from a posterior
paler region. In one specimen the change occurs at about I of its length
from the anterior end, in two others at t, and in one at " of the length.
Fl'Om the condition of the ventral ambulacra it appears that this hinder
end has been regenerated, for here the podia are in distinct narrow lines,
whereas in the normal darker part of the body these organs spread out
into the interambulacra, where there are about 15 in a transverse line,
instoad of only 2 to each ambulacrum. There is, too, a transition observable
as the ambulacra are traced forwards. indicating a gradual rt'sumption of
the adult condition.
Disirtoution.-Bedford, ill his report on the Funafuti Holothurlsns,
Rpeaks of this species as .. the most widely distrihuted circumtropical
species of the genutl."
Chirodota rigida Semper.
Semper, Reisen im Archi~l ut'l" Phihppinen, HolothllI1en, p. 18,
pI. 3, fig. 3; pI. 5, figs. 8, 18, 1868. Lyman Clark, "The
Apodous Holothurians." p. 117, 1907.
The wheels difier from those figured, in that there is a distinct con-
striction of the radii at their jWlction with the rim; but. as my specimens
agree in the general characters of the species, I have little doubt that this
is the correct detelmmation. Oliwr states that the ., goneral colour is
reddish-purple; it occura. in sand and mud under stones in rock-pools
and at low-water ·mark. It is not rommon."
LJ(l.-Meyer Island.
DiBtributiOfl.-Clark S&ys it is .. apparently well distributed through
the entire East Indian region." I
flElliHAM -fll"frttbl'frfes trom flu' Kfrmadfr Islands 187
Class SIPUNCULOIDEO\.
Slpunculus nudus Lmnaeu"l.
This Mediterranean species is widely distrIbuted; it heLt! lIeen recorded
hom Singapore, Japan, and elsewhere.
Lo(.-Sun<iay Island.
Collected by Mr. R. S. Bell.
Physcosoma scolops. Selenks and Man.
Phascolosoma annulata RuttLn, Trans. N.Z. Inst., 12, p 278 188U.
PhymosoMa scoWps Selenka and Man, ., Die Sipunculidell," p. 75,
1884. Physcosoma annulatum Benham, Trans. N.Z. IllBt., 36.
p. 173, 1904.
When I described the Sipunculids of New Zealand (Tran". :N.Z. lust.,
\·ols. 36, 37) I had not the opportunity of consulting Selenu's monograph,
which was only pnrcha'Jed by the Otago Institute at a later date. I find
now that our common Ripunculid, whioh Hutton described in 1879, is
identical with Selenka's P. Boolop!J, a very widely distributed speCIes, which
was described five years later. Hutton's brief diagnosis, depending only
on externals, is insufficient for identification, and so must. give way to
Ralenka's specific name.
I note that, although FIscher (Die Gephyrca., Abhandl. aus dem Gebiete
Naturwiss., 13. p. 10, 1895) regards P. 8~olops a'J a variety of the Medi-
terranean P. grallulatum, Shipley still retains it as 1\ distinct specie"l
(Willey, Zool. Results Rep. on the Sipunculoidea, p. 156. 1899; a.nd R"p.
on the Gephyrea, Pearl Oyster Fishery, Ceylon, p. 174, 1903).
It is evidently very common on the Kermadec Islands, for I have
more than fifty I collected on various p.uts of Sunday Island and on
Meyer Island in the ordinary positions-that is, under stonos in rock-
pools, in amongst coralline algae, &c.
The distribution is very wide.
Aspidosiphon truncatus Keferstein.
Selenka and Man, ,. Die Sipunculiden," p. 118, pl. 13, IB8~.
Of this identification I do not feel quite certain, for the convolutions
of the intestine are fewer, and the longitudinal musl.lle bands rathor
more numerous j but as our specimens agree in so many features with
those of Keferstcin's specios, and do not agree with any other description
to which I have access, I place it here. The differences are so slight that
I do not feel competent to differentiate a new spocies.
Loo.-Sunday Island, in coralline algae. Six specimellB.
Distributitm.-Mauritius, Panama, JapBn (Iketla. Journ. (.'011. I:kli., 20).
Class CHA.ETOGNATIIA.
Sagitta fowleri nom. nov.
Fowler, .. On Plankton Chaetognatha. of the Bay of Islands, New
Zealand," Ann. Mag. Nat. Rist. (8), 1, p. 240, 1908.
I received Seven specimens of a rather large Chaetognath whieh had been
cast ashore, and were somewhat injured, and had unfortunately been placed
in a tube rather too small for them, so that they nre not only damaged by
the sand, but also folded a.nd crumpled. At first I failed to notice the
anterior lateral 00, and took it for a species of K'1'ohnia .. but the formula.
given by Dr. G. H. Fo,vler for an unnamed species from the Bay of Island'!
j'1 fllIJlW'tW/lX,
agr~es so precj~ely with the Kermadec forIDs, and in SODlO rospl·(·tl> 11:1 bO ex-
ceptional, that I carefully went through all the sperimens a!!,uin. III only 0111.'
indh;dual could I detect th(' anterior fin, a.nd this cjuitl' plainh, ,llthoUl!h it
was folded against the body. Iu its extent it does not o.gl'ee with Fowlcr':-.
figure, though hI.' places a (~) ngaimlt his sta.tement ill thl' !I'xt. But mving
to the damagt' dOll(' to the post{'rior fin, and owing to the tenuity of this
ullterior fin, I should not presume to doubt Fo,,'ler's I:Itat(,)Ul'nt that thib
fin extf'nds forwards a& far as the level of the yentral ganglion. though.
1:10 far as my specimen shows it, the fill is of mUl·h 1('88 extent.
Fowler refrain('d from naming his two immature and somewhat
damaged specimens, aud did not even place it ill a genus, tho~h h(' srutes
that certain of its characters .. suggest ne:raptera," at the Scl.IDf' timf'
pointing out certain difIerencl's from that species. As the only genus with
two lateral fins is Sagitta, there is little doubt that he intended to comparl'
it with S. hnaptel'a, and I take the opportunity of naming it after }.im,
My specimens yary from 23-35 mm. in total length. with a diameter
of 2'0-3 mrn. Owing to fiacciditr of the hody, it fiat1f'ns casil~', and ha~.
liS I haye said, been crumpled.
The head is distinctly constricted from the body: till' <.:urn·d hooks, or
.. jaws," are 8 or 9 on each side; in one (,lise 8 on one side Rnd i' on the
other. They haye 110 distinct separatt' tip. hut th(' whole hook is gentl~'
curved and "dthout am' slllTations.
The frontal spines. ~r .. ,mterior te~th." arl' 011 ~ each ~idl·. thoulI.h ill
one case 4: 011 one sidp and 3 on the other.
The marginal spines, or .. hindel' series of teeth." form I.l. row of 3 ~hort
conical spines on the sloping anterior ma.rgin of tho head.
The tail fin is in all my specimens slightly notched; tIw posterior
latera.l :fin commences ra.ther ill front of the middle of tht· t~ul. ,IUd it!
widest just behind the !lUus. IX> far as the imperf(,(·t l'olldition allow"
one to judge, it has l of its ll'ngth behind and ~ in front uf thl' ItUUI:I.
The anterior fin is 0111~' 3 mm. ill length; it s('eml:l ,wll defined. ,llld
I fa.iled to see any f'yidl.'nc(' of its continuation fUT\nlrds; its antt'rior
margin is 10 mm. from tllt' tip of the heacl (the YOlltl'ul glll1~.lion hein~
about 6 mm.): its posterior limit if! 5 mm. in frcmt of thc' tllIus-th,lt i:.l.
rlose to th(' pol:lterior fill.
The fOl'lllula used b'\'" Fowl!'r iR -
35 20 ~
.:1.,
27 :W !I 3
,
25 20·3 R-!I :J--j.
Loc.-Sunda, Island.
Distrt''butiml.:_Bay of Islands.
{'I&llS EXTEROPxEt·tlTA.
Ptychodera flava Eschscholtz,
Willey. Q. J. )fie. Sci.. 40, p. 165. Punnl'tt, Entt'l'Oplll'U:.tll. :It'II1111U
Maldive and Laccadive Archip., vol. 2. pt, 2.
A single laceratoo bl'Oken indh·icluII.I, fonnel .. nlldf'l' stOlWI'I" ,It L'm-1I1
Ba.y, ~unday Island, Julr, 1908.
Dis'7'f'butinn.-Indian Oef'lIll.
HOUHEX.--E{ll'tltqllakf-lJril/ill. 1/1 tile S(lIIf 1t-II't'lIt l'al'ific. 139
The waves used for determmmg the origins are the prehmmary tremor!>
and the long wavcs; the methods for the most part trial methods, such
BS the differential method a.nd that based upon the interval between thE'
arrival
and of the
those PI waves -;:=~:ii==~==~=::;:i~=::::i~=:;
of maximum ,. 110 ,.. ..I
amplitude.
The results tor eleven 1
.' . . . . :...
.. .. I
earthquak~s in which the I'.J :::--::---f___:=-=+:;__-t----L..:.
data are sufficient to -
determine the epicentra
·1
.
..re given below. Those
call e d "approXlDl.8.te "
are epicentra. probably
:
I
('orreet within the limits
of error of the observa·
tions; those called
"probable" are epicentra
fOT which there are re-
sidual errol'S somewhat
in excess of the limits of
errors of observation.
lIt should he noted
that the method of least
squares camlOt properly
be used UlIlt'ss the phy·
lIieal conditions are ap·
FlO. 1.
proximately the same.
For instance, we cannot Earthqllake-ollglllS III the Sout~-\\ebt Pacific, 1\:110
use it in reference to (G. BoqbslI.j
• Approltlluate epicentrum.
l·quations based upon ob- C Probable epicen;rum.
servatiolls fronl StatIOllS , OrIgins prevloulioly foulld.
varying greatly IlL their
distance from the origm; it should be applied onl~' to deductions from
obselvations of wavus p,tSbllltJ, along the samE' paths, or, a.ssuming the
l:Iymmetrical distribution of the various strata of the esrth. passin~ along
paths of nearly the same length.)
Tho oligin in each case moly, of course, han bCl'll a more or k'8B exten-
sive mass below the epicentrwn indicated on the map (fig. 1). Tb.e mll.p
",Iso shows the positiolls of pl'e\·iousl..,- ascertained origins in Australasia.
llale POIoltlon ot EPle,,"truDl. Remaru
1910. Lat.
13 Jan.
3 Feb.
. ,H· H.
32C' ~.
lifo"E. .\pproxllDat.· ••
l."i3° E. PInbabl,,·
::s'ot It'lorUl.,d Ul Eurupo 01' othor
diRtant btation'4•
nscorded at Ill'll.l and (h~tant oria·
tion~.
80 lIaJ.'l:h
1 JUll~
UI-
18° E..
I). lbO° .E. .,
170" E. Al'pl'UXlDllitl." ••
DittC'.
Ib
...
27< K. 173· E. Probabll' .
29
211
" (Ill ••
(b) ••
11" K.
in!;: ~.
172° E. Approximatl' ..
1731° E.
.
Het\,wn CalDllbolll.J.uldt. Q.l1I1 Au
Hpod(.'b .ullln,,1.... .\11 'lto.til)l1~
7 &'pt. 32° M. liVe W. All ..1atiozl'.
{) Nov. 17" S. U17· E.
2tI
10 J~.
.. 5° S. 16Jo E.
2° N. 14tlo E. PtolJAhl,·
HooBEN.-l!:arthqualrl-orlgI'll8 m the South-u·e,t Pacific. 141
The most mt('restmg of all these eal·thquakes IS perhap& tha.t of the 9th
~ovember. 1910. thc origin of ,vhich appears to have been below the ocean,
It little to the south-west ot Espiritu Santo m the N('w Hebrid('s Group.
The 8B('ertained elements of the prehnllnarv tremor& of thiR Nrthquak..
wIth reference to ple\'('>l1 statIOns are giv(,l1 III thp tablp nplow.
Earthquuke of 9'11 SOWlibel. 1910.
(EpIcpntrulll, l7e~. Lat .. 167 0 E. LOllll. Tnnt' at Orip;m. 6h. 03·7mm.
G.Y.C.T.)
..I.lcuaI' -~ ---I - - -
I'IOG< ul Obo.PI\atlon, D,'ltll.llce Chordal TIme ofl v (Arc) "',
.md latitude . l.ollgltud. flOID I Dl&tauce 16 : 1+ ,]i'iom. ~:)
In,trulll,nt Ougru ,(lUiom.). min pel min. J14."r mil;
(X,iom). "
Apia (\Vleohert) ..
S Y d n e y (Wieohert
113< 41:1' B'I' 171'
33° .~Il' Fl.
4t>' \Y.
151 0 12' E.
I 2,296
2,452
2,284 , 06·7 , ill.1 I'
2,436/ 06'1! 766
761
761
and Milne)
Wellmlrton (~mnc) 41' 17' B. , 174" 47' E. I 2,800 2,777 07'351 767 761
Porth (Alilne)
Honolulu (Milnp) .•
.. , 31° 57' S. 115' 50' E.
,210 19' N. , 158" 03' W., 5,305
3,711
:;,209
5,521
I 10'0
10'8
I 804
778 756
778
Batavia (Wiechert) lie 01:1' S. lOll' 30' E. 6,317 6,063/ 11'7/ 790 758
Zlkawei (Wieohel1.) SI~ 1,')' N. , 121' 26' E. 7,235 6,850 ]2.-& 832 781
Vit'toria, B.C. (Milnp) 148' 24' N. 12S· 22' \Y. 10,056 9,030 14·6 923 I 836
lIa.ch'a,& (Milne)
Eclinburgh (MihlC) ..
.. 10 14' N./
I,');)n .;7' N.
77' 2!{ E. 110,283
S" 11' W. 15,667
9.201 I 11H,
12,000 2.2·1
902, 807
851 652
So.n Fernando, Cadiz gllo 28'~. I h· 12' W. 17,889 12.1i60 : 22'6 665
(MIlnl'l 946
1
NOTIl,-P b preliminary tremors; VI' velooity ot PI WS.Vrt..
It will be seen that in this case there is a closer agreement betw(.>on the
velocities (values of VI) for paths calC'ulatcd along the chord than fol' those
calculated along the arc; in other
words, that the chords represent a o
closcr approximation to the actual
pathil than the ares. (It will be
understood that the chord cannot be
the actual path of a wave passillg
through layers of varying density,
and subject, therefore, to refraction
at the bounding surfaces.)
It will be seen that the velocity v
(P1) of wans, calculated along the
chord, for places not more than 600
from the origin is about 760 kilo-
metres, per minute; that for places
betwpen about 600 and 900 from the
origin the chordal velocity is
greater; that for distances over 90° FIG. 2.
it is considerably less.
Possible Pa.iJls
This enables us to formulate a 1910, to W~Ili:cgOOD, of PI Waves, \I.h November,
OW; Perth, OP;
hypothesis illustrated by the dia- Zik&wei, OZ; Vlctnria (B.C.), OV; Sm
gram (fig. 2). Disregarding the sur- Fernando, Ill, or 222, or 81 8. 0, origin.
face rocks. which I have elsewhere
shown to be not more than twenty-five to thirty miles in thickness, we
ma~' assume a shen of much greater deJlsity about liOO miles in depth (AAA,)
1.42 T 1'(111 ~flctiollx.
tmd below that a shell of still greater density about 630 miles thick (BBB).
Below the last· named shell there seems to be a marked change of physical
condition-either the density is much less (which is hardly conceivable) or
the centrosphere (CCC) is viscous. I have drawn hypothetically the possible
paths of preliminary waves reaching the San Fernando Observatory from the
origin: (a) They may have been transmitted along a path approximating
to the chord Ill, but with grl.'atly reduced speed through the central
portion; or (b) they may have bel.'n transmitted along, or nearly along.
the path 2 2 2, as internal surface wavl.'s for the middle portion of the
path-that is. along the surface of the centrosphere; or (c) they may have
been transmitted along a path 3 I" 3-that is, along the chords 0 r, I" S.F ..
being reflected at r.
I put this forw'ard as a mere trial hypothesis, based upon the examina·
t'on of the records of one earthquake, and examined only paltially by other
re('orcis. It is, however. I think, worth careful examination in the li5lht
of all the available data of othl.'r earthquakes. I propose to makc such an
examination (which may last months, ur e,'en years), and hope to place
the results. whether positive or negative. 11l.'fore you on a future occasion.
I should like to express my appreciation of the kindness of the observers
in ('harge of the }Iilne seismographs at Sydney. Adelaide, Perth, and Christ·
church in sending me copies of their records and seismograms. I regret
that I have been unaule to obtain any of the records of the instrument at
the Melbourne Obst'rvatory. .
AR.T. XI\·. - FblctufltimlX ill the Level 0/ th,' Water /11 some Artelliflil Tf('{1i1
in the (,hrisfellllTrll Amr.
By F. W. HILUENDORF, )l.A., D.Sc.
rRp·"J IIPlol~' tIe Pidlo-.op/w'"llIl,tllllle 01 ('alltprblt'!J. 8rl, DptBlllber. 1911.1
All pa.rt of the actiyities of the Artesian Wdls Committee of the Canter-
bury Philosophical Institute. observations on fluctuations in the silltic
height of the water in l:Iome flowing wells ill the Christchw.·ch artesian arell
well!' undertaken early in Janunry, 1910. The records or the wells will he
dealt with separately.
(1.) LIS(,OLX COLLEGE WELL.
This ,,",,li is 341 ft. deep from the ground·level. which it! 38 ft. above
sea-If've!. It;s a 2 ill. pipe, Ilnd was sunk ill 181.13. The water rises to
about 8 ft. aboye ground-level.
There are in the district foul' oiher w('l1s of approximately the same
depth. The nearest of these is about three'lluarters of 0. mile' away. and
the next: nearest oyer a mile S,\\·RY. •
The ohsel'Yations were taken b~: means or u glass tube attnched to a tup
bored into the well-pipe. and tht> tube "'as backed by It wooden scalo
marked in centimetres. The h,draulic mlns worked by the well were shut
off for the purpose or ta.king tht' ohserYations, aud the watl'r in tht' 1uh~
,.llowed to come to rest. The OS("illl:ltlOlIS ceased in auout five minutetl. A
loose-fitting plug was placed in the top of both the ,vell-pipe and glass tuue
to prevent the wind hlowing down .md lI.gitating the lenl of tht' watE'r.
The readings were taken at 8 H.m. and 5 p.m., lllld only eleyell l'ead-
ings were missed dUl;nQ the year.
The. Jlo)lth7y F7uctuafiOOI.
DisregardinA the minor varitLtions, the well sallk grllduall:r from Januar~
to Juno, durinl1, which time it fell 24- cm .. or 10 iu. On the 10th JWIC
and tho fonowing dars, 6 ill. of rain fell Itt Lincoln, and th... well tht'n
started to rise, and continued to do so fOl' four months, during which
time it rose 66 em .. 01' 2 ft. 2i in., on l:In average of the weekly readingb.
The lowest illdiyidual readiug was 71·2 em. on the 4th JunE', and the
highost 141·5 em. on the 25th September and the 17th Octouer. This giYl's
a maximum difiercu('e of 70'S em.. 01' about 2 ft. 4 ill.
The following graph shows the static level of the well for eaeh month
during the year. lI.11 the readings for the month being o.vel'8ged to find
the level for that month. Below the graph of the static It'yels there ill
shown the l1lonthly rainfall at Lillcoln in inches.
",
100 I\.. j
QS / ill"
QO J '1 0
u \.. I t..
!fa
"'\.
...
I S·
'"
• 0
')$ 4"'
. 3"
a"
.
."
-
~'IO, l.-l:0liTIlLl."
-
A, DAGLS 01' HEWIIT OF WELL, AloD :llO!'TBI.l l'OT.6L1:1 (IF HAI.li·
on
!'\LL AT LI.liCOL~.
14:4: Trn.n1actwn,.
A study of tws graph shows that the ra.in~lls from Januarr to May
were not enough to balance the water drawn off from the reservoir sup-
plying the well: that the rains in June and July were sufficient to
replenish it; that the almost total absence of rain in .August waf> accom-
panied by a still further rise in the leV'el of the water, possibly indicating
that the heavy rainfall of tho previous months was still percolating tu
the reservoir; that the rains of September and October were accom-
panied by a slight rise, although they were almost exactly equal to
the rainfall of January and February, which were accompanied by a
fall in the IcV'el of the well; and that falls took place in NovelJl.ber and
December.
These last mcts, and also, in pali:, the rise in .August, are probably to
be explained by the great amount of eyaporation in Novembt-r, Dt-cember,
January, and February. and its smlllier amount in August, Septcmb"r,
a.nd October; that the evapora.tion might have an effect 011 tho
tiuctuation of the well did not suggest itself to me early enough for me
to install evaporation-gauges. It seems probable that the evaporation in
thp summer months ,vould exceed the rainfall, and thus assist the lower-
ing of the static level of tho well; while in August, September, and Octoher
tho evaporation would be very slight, and thus all thl' rainfall would be
available for replenishment of the reserYoir. Thc following tablu by
Greaves, taken from Walring1.on's "Phygical Properties of the Soil,"
p. 108, is instructive :-
I
-----
Jan. 2·87 O·it) .Tuh· 1·77 3·44
Feb. 1·60 0'00 .\.ug. 2·33 2·85
March 1·94 l·ui Sep. 2·35 1·61
.April 1·-13 2·10 Oct. 2·73 1'06
May 2·06 2·75 Nov. 2·02 (I'7l
June 2'21 3'14 Del'. 2'42 0'57
lOS
..,J
t .... , 'It "'6 III 1I.,.1L I. 10 ..... to 1~ #. U
=
IDO \ ,
\-.
"- ~
Q6
qo '\. J
I
I
" ,.,
\.
as . \ , 6"
"\ S· .
AQ f' 4--
1\ .$"
'70
I
""'- r--
l'
,"
,I I I . I 1
• I. o·
FIG. 2.-WEEKLY AVERAGES OF HEIGHT (IF WLu", AND WJD1IXLY TOTALb OF RAINFALL
AT LINtOLN.
It is clear from this graph that th(' wpll rISes whenever rain falls. and
that the rise in the well is approximately proportional to the rainfall.
This result was anticipated from the work of Hutton* and Speight,t but
it was considered impossible that the rainfall at Lincoln could be respon-
sible for the rise in the well there, since, as before mentiou('d, the well
draws its water from 341 ft. below ground-level.
Lincoln is situated on the Canterbury Pla.in, fourteen miles from the
sea. The plain is about fifty miles wide, and slopes upwards from the
sea to the mountains, at whose feet its le'\"el is about 1,300 ft. It is com-
posed of a coarse gravel interstratificd (especially in its coastal portions
near Christchurch) with clay, peat, &c., as described by Speight (Zoc. oit.).
On the supposition that the lower strata have been laid down at 0. steeper
angle than those now on the surface, the water-bea.ring stratum tapped by
the Lincoln College well should outcrop on the surlace of tho plain some
miles above Lincoln, and it would probably be the rain falling on this
outcrop that would supply the well. This idea is embodied in the following
diagrammatic sketch, where the heavy lines show the clay strata between
* Tran,. N.Z. IIl'lt.• vol. 28, p. tIM. t Trans. N.Z. In..t .• vol. 43, p. 420.
140 TI'IlIl'III'/IUII'
aud noting at which places the ralllfall most lll'clrly wrresponded with the
:8.uctua.tions in the level of the well. For tlus purpose rain-gauges were
msta.lled or eXlSting installations were used to obtain records of the daily
rainfall from the following places. RollE'stOIl, Lawford, Kil"Vlree, Dal'field.
Hororata, Glenroy, and Mount Torlesse. The pOSItions of these places are
shown on the following map. w'hich also shows thl' two rivers of the district,
The slope of the plain is from llorth-wl'st to south-t·ast.
FlO. ~.-lIAl' or POUTI'f' "r t'~,\T:cnllrm. I'L~n., ...lI'nW~G PO... lTW....b OF n.u~
b It"OEOO:.
I, Lincoln: !!, HolJt.,toll: 3, l..awfo.rd (half-wd" Jx.tWe1l11 'rcedon'~ ,\1111 Web!
ltE'ltonJ: 4, Kirwt'(': ,i, D.u'ticIrl: H, HOYtll'R.tol: 7. mcnroy; S, lronnt Tor.
Itoq\oe It\\"11 Jmt.... ,\110\1 S}Jriuefidcll.
Hrr.OF:NDOI!F.-ArteR/(1/I Jrell~ III thp (.'/11"/('/'1,,,/1 A'UI. 147
14!:.l TrtIIlMlrtlom•.
Unfortunately. nil the lecords (hd not begm at the begInnIng of the
year, those from DadieH !lot commencing till the 1st April. and those at
Rolles10n till the bt June. Valuable miormation was thereby lost. The
records are, however, eomplete and Meurate for the period" they cov('r.
In the a('companying p.raph (fig. 5) the ayerages of all the readings of
the well for each \veek for five months are shown, and underneath them
the total weekly rainfalls fOL each of seven stations, Rolleston readings not
having been commenced. Zero for rainfall is made a sloping lllle, roughly
corresponding to the graph of the well, for the purpose of bringing the
rainfalls - graph near to that of the well. to facilitate comparison. To
simplify the figure, rainfalls arc shown. for only those weeks from which
conclusions mav be dmwn. The falls of Glenrov and Kinvee were identical
for the weeks 'shown, and therefore these two 'stations are represented by
only a single symbol-vIz., dots and dasheB.
Starting with the station nearest the mountaius-viz., ~pringfield-if
the rainfall at Springfield (squares) for the week ending the 15th January
was responsible for the rise of the well shown for the week ending the
22nd January, then also the much heavit-r rainfall for the week ending
the 26th February must have been responsible for the almost imperceptible
rise for the week ending the 5th March. These two results are inconsistent,
and therefore it may be stated that the rainfall on which the well depends
does not fall at Springfield, nor does thl' water-bearing stratum tapped by
the well outorop there. Similar inconsistenoies may be noted for other
localities, as follows :-Hororata : In the week ending ihe 15th January a
rainfall of 1 in. is followed bv a rise in the well of 1·5 em., and on the 28rd
April a rainfall of 2; in. is followed by a decline of 1 em. Glenroy shows
inconsistencies for the weeks ending the 15th January and the 28rd April;
Darneld for the WE'eks ending the 2nd and the 231'd April; Kirwee for
the weeks ending the 15th January and the 23rd April; and Lawford for
the weeks ending the 15th January and the 23rd AprIl. But when we
come to e:mmine the rainfall a.t Lincobl and compale that with the subse-
quent rises, or arrests of the decline of the graph of the well, a remarkable
degree of consistency is diselosed. The graph of the sta.tic level of the
well is as neal'ly parallel to that of thc Lincoln ra.infall as could possibly
be expected under the circumstances, and, being give.n the rise of the well
due to the rainfull of the 15th January, the graph of the 011e could be con-
structed with reasona.ble accuracy from that of the other. From this it
is evident that the stratum tapped by the well outcrops in a district with
a rainfall during the months shown almost e:mctly equal to that of Lincoln.
None of the stations recording for mt.' shows such an equality, and so
it seems evident that the water-bearing stratum Ullder consideration out-
crops nearer to Lincoln than to the nearest of the stations. That station
is Lawford, nine miles away, and so one would probably be safe in saying
that the stratum of shingle 84:1 ft. under the surface at Lincohl reaches the
surface seven miles or less up the plains. This would place the outcrop
somewhere about Rolleston-a district noted for its loose shingly soil,
direM:ly underlaid by coarse gravels, with no interposing layer of clay.
I:!uoh country is absorptive of water in the highest degree, and an ideal
catching-area for o.n underground water-supply.
The surface of the land at Rolleston is 134 ft. above that at Lincoln.
The water-1x>aring straium then rises 475 ft. in seven miles, or about 68 ft.
to the mile. The sudact" of the plains near their upper limit has a fall of
about 60 ft. to the mile. \vhile hetv.·t'en Rolleston and Lincoln it is onl~'
HILC,ENDOllF .--.irtellia II II" ell~ I II fIll' (Jh 1'1 Sft·11Il rrll .11"1'0 149
20 ft. to the mIl!'. At the time that the fall on the surface of the plallls
betwt'CD Rolleston and Lillcoln was 68 ft. to the mile-that is. when Ollr
water-bearing stratum was depositt'd-th!' whole plain must have had a
much steeper gradient than a.t present. This would probably be due to
the much greatt."r bupply of waste to the above-gorge waters of the rivers,
so that in those times the present plains would hav!' been mueh more like
the present-day shingle fans than like plains. That the gradient of the
plains was once much stet'pe'r thlloll 110W ill provtld by the high terraces round
Woodstock, and by Racecourse Hill. d. Tl:'sil1ud.l sllingle mound some 60 ft.
high. The cuttmg mto their bl"ds of the prosent rivers is merely a continu-
ation of the process of lesserung the gradient of the plain. the- bed of the
Waimakariri being virtually level with the plains at their lower edge. and over
300 ft. below them at their upper limit. It is therefore in accord with what
I suppose would be the expectations of geologists that at one time the surface
of the plains should be much more steeply inclined than now, but that the
supply of wdste should be so great as to form a deposit sloping nearly 70 ft.
to the Illlle forty miles away from the gorge is perhaps noteworthy.*
It was stated above dB evident that the collecting-ground for the well
is nearer Lincoln than the nearest rainfall-station is. On the part of one
unacquainted with the country, a possible objection to this is that the
collecting-ground might equally well be more distant from Lincoln than
the farthest station is. The COWltry between Springfield and the West
Coast, however, consists ot mounto.ins of greywaekes and sla.tes quite im-
pervious to water in large qUllolltities, and, in any case, this water would
percolate out into the rivors ftowing at the base of the mountains. The
a.mount of water in the Waimakariri is, moreover, a gauge of the amount
of rain !alling on these mountams. and I have been so fort.unate as to be
supplied ,vith daily readmgs of the height of the rlVl'r during several months.
Most of the ftoods that my records show occurred nearly contemporaneously
with considerable rainfalls on the plains, and th(' subsequent rises of the
well could not. therefore', be btated as dependent on, or independent of,
the rises in the river. On the 23rd March, however (see the arrow-head in
fig. 5), there was a heavy ftood, sufficient to stop the mails at the Bealey,
but, as fig. 5 shows, there was no sign of any rise or arrest of the declinl"
of the well until rain foll in the second week after the flood.
Although it is impossible that the rain :£alhng on the mountAins should
dirtlctly find Its way into the water-bearing stratum tapped by the well, it
seeml'd quite possible that after reaohing the river the water might percolate
into su('h a stratum wherl' the river runs across its outcl'Op. This, indeed, is
probably the common opinion held; but the observations made do not sup-
port the supposition, as f.l.r as the well at Lincoln goes. Thl' observations on
• At the meeting at wlueh thIS paper W&!o r.md 11r. Spoight pointed out that the
oonulusion reached here is probably UI0'Jrreot. Hi... oLbervations on HI!." strata piercod
by wclls near Christchurch IIhows that the doep.l'yil1g strata are a1 practically the
.."me slope a... tho pror,ent surface of the plain. It I, a mattel· of common observation
that clay strata. though '·ommon near ChriE.t-
ohuroh, disa.ppea.r farther up the plain~, and it
It is probable tba.t Rollot-ton mo.rk., tho clitltance
from Lincoln at whiolt the oloy IItratum over tho
water·bearing stratum fadell away, rather than the
outcrop of a. rt·ries of ..trata ~guJar in thiokne1!8
from the base of the welJ to the outOl'Op. Thib
idea. is shown ill the aooomp.m;ring diapmmatic
llketch, where the arrow·head bhoWII the position of RoI1estoll. Mr. Speight'll in·
terprotation of the faots seems to me cor.reot, and iDvaJi,latt'~ tht" ,·oneluolion, abow
,bawn &8 to the fanner slope of thE' ~Ilna('e uf the p)ain~.
150 1'rc/llx(u·tinll '.
the height of the riYer wt're made w.i.th great care, readings being tak~ll eneh
(lay to the nearest inch. On compariosn with the graph of the static level
of -the well no agreement ('ould be observed in any case, and the perfect
indifference of the well to the flood on the 231U March is typical of this.
Another possible objection to the placing of the outcrop at Rolleston
ill that thil! has been done almost entirely on the slight rainfall at Lincoln
aud the heavier rainfall at all other stations for the week ending the
23rd April. This is quite true: but occasions on which the rainfall is
markedly different at different points on the plains are rare, and some
vears of obstlrvations may be needed to secure a confirmation, hy thi:l
method, of the conclusion- dra\f"n. In the meantime, the accumcy of th£'
rainfall recorded at the various up-plain stations is sufficiently substantiated
by their mutual agreement, a.nd the accuracy of the record at Lincoln by
wmpa.rison with that mad!! by three other observers in the neighbour-
hood.
Tlu' Doily Fluctuation.
EYen during long p"r~ods of steady decline 01' rise of the "'ell its static
I1'Ye1 showed ('omparativl'ly largt! daily variations. On some occasions it
would rise 3 in. in twenty-four hours (without rain), and would fall by
the same or a greater amount by the succeeding morning. Varia.tions of
2 in. on successivtl morningl! were common, and usually the morning read-
ings showed variations of owr 1 in. The irregularities of the static lewl
within short periods of time during which no rain fell led to an attempt to
correlate the variations in the well with those of the barometric pressurt'
I)f thto air_ At length it was found that by turning the barometer-readings
upside down and multiplying them by four a marked degree of harmony
between the gmph of the well lUld that of the barometer was displayed---a
harmony so consistent as to establish the mct that the level of the water
in the well and thtLt of the lUerCUl'Y in the barometer are influenced b'v the
samc causes. The accompanying gl'8.ph (fig. 6) shows this clearly. -
w ..... 111M
.II~~ :& ."Il.-
c.". 2 :!'i' Ht as 2' 2.8 ::I! io 1 rz;'C.11
g, iJ.Q~ll)
'\. ·_-11_6
~,
...".
911 ·ltt
,,
••
• 1\ ·11..
AS
•\ J i\\ /~\ ' ••It
If .. •
lr..
13
.'. I"
. -~ .
. .0-/ :
". ,.~
\.'
\ "Y- .
/\
: i\
o
~\
I
t tt
Ii
J 1_\\
\
\
3DoOO
'0&
_0'1.
ojS
Thl' ('velllng ru,t' 18 thus fcurly well marked. Durmg thc month!, of
Octobpl' and XOyemhel I personally secured that all the wells ill tht·
ueighbomhood WClt:' lUllDlllg continuously, with the exception ot onl'
(thrpl'-4uarters of a. mIle cIoway) which its owner was good enough tI
."hut off from i to !) c.L.lll. and 4 to 6 p.m. ever~· day. The readlllgb
dUling thesc two months ,vere taken exactly fit ROO a.m. and 5.30 p.Ul ..
so tha.t the well had an hour and a half to recover any rustUJuance
that might have been set up by the well whose :flow was mtemllttent.
That this int<.>rmittently flowing well could have any efiect on the
CollE'ge well. so far away, is questionable, and, in any case, it was not
(e\'cn during the month", I did not keep special control of it) usuall,'
running ill the morning or usually shut off in the afternoon. The intel-
ference of neigllbourinp; wells rna\' therefore bt' rejected as a cause of th,·
eypning rise.
Any ro:ru.tant variations ill temperature are similarly to be rejected.
I kept a record of the temperature of the flowing water just as it emerged
from the ground from the 10th to the 30th October. The temperature
varied from 12·81° C. to 12·90° C., and this variation was more probably
due to th.e effect of the air on the stem of the thermometer than that
of the water 011 Its bulb. In any case, the temperature never showed
any disposition to be regularly higher in the evening than in the
morning, and, if it had, a much greater rise of temperature would havt'
been needed to cause sufficient expansion of the water (inside an iron
pipe. on which the scale was carried) to accoUllt for the observed rise in
the static le\'"<.'I. The water in the gauge-glass is. however, practically the
same water all the time. and therefore takes on to a considerable degree
thl' temperature of the atmosphere. It varied from 10.00 C. to 23'9° C.
durmg the month of October. The higher readings were, however, on
all but three occasion& obtained in the morning, owing to the sun
shming on th~ gauge-glass and aboye-ground portion of the well-tube
in the morning and not ill th~ evening; indeed, the highest readin~
(23'!:IG C.) was obtallled in the morning, Ilnd on the samo evening the
temperature was 12·(1 ('. In any case, an average evening rise of
temperature of about 25c ('. would be needed to cause a 4: ft. column
of water (in a glass tube with a.n independent scale) to expand suffi-
ciently to account for the observed riso in the static level. A shrinka(l('
of the wooden scalp in the evening would also explain the rise: but
means to detect and measure this, if it occurred, were not at hand, and
the line of inYe8tigatioll held little promise. During the months of
October and Noyember, also. records were kept of the barometrIc
pressure in the mornings and eYenings, and it was found that the
readings werE'. on the average, lower in the evenings than the mornings.
The amOUllt of the decrease in the ba.rometric height in the evenings was
0{)7 in.. sufficient to acCOUllt for a rise in the well of 0·56 em., or more
than the actually observed rist'. The barometric observations were, hov,·-
ever. taken on an aneroid barograph, the mercurial barometer unfortu-
nately being out of repair. There is, I suppose, no question that the
temperature of a living-room is higher in the evenings than the morn-
ings, and I have rather good proof that increase of temperature de-
presses the record made by an aneroid barograph. The apparent fan of
the- barometer e3("h evening is, thE'refore, onl;V a temperature effect, and
1I1Ulot be used to ,·xplain the rise of the well. TIns tact 11> t'mphdSlzcd
iJy the following graph (fig. 7), obtained 111 Invt"rcarglll III HIlla it I"
pl'l"haps sufficil'ntly btrikillg to merIt puLhc ahon.
,
~.:.. _. -
A,... '2.1. 2B
- --
I
.lll. a .. Pl.,
... ",
I.
- a..
MN
..t.
ao-so
10·1$ ~ j
V r'" IV ~ ~ I
~~
V
iii'.
IV ~
manner a.nd to the Bame degree at! in the Lincoln wt'll, but there 18 a much
less decline in its static level during a similar period of aJm.ost similar rain.
(3.) There is no sign of floods in the Waimakariri in:B.uellcing the well.
l4.) There is no sign of agreement between the graph of the well and that
of the barometer, however the latter is mallipulatf'd. l5.) There is all
evening rise. Points (2) and (3) are illustrated by the tollowing graph
(fig. 8). \vhich is comparable to fig. 2. both graphs being on the same scale.
.~1. , ,IrAI"
c...... faa.... IS ~q 00J: ~ :n --"0 ~ 2.1t U.s :u. .I 5 14 f,:l ,IL 1'1.1 1 .......
lOS
:f' ~ ~
"-
1'00 ~
as
. \
i'- I\. j,
QO
" \ l/11"'" t\.. , .,
QJO 1"
• I"
11 • . II II I o·
FIG. tI.-W'nI:xr,y _-\'VIm.\OES Oll' HEIGHT 0.11' MUSEUM WELL \ND WBlCKl,y TOT.\UI OF
R \I!OO"FALL \T LINCOLN (] 894-!l.l).
The want of agreement between the gt'8.ph of the well and that of the.>
barometer may be explained either as the result of the Wo.imakariri assist-
ing the rainfall to supply the well, or as the result of the interference of
neighbouring wells. That such interference does take place has b,'en shown
in a prl'viuut! lIection.
The lack of pronounced declme during a comparatively rainless p~riod,
and the smallilcss of the annual variation (Wi in.), opens up serioll8ly
the question ILS to whether the Waimabriri does assist the supply of tho
.flowing wells in Christchurch. In favour of the raiIllall being the sol...
source of supply are the following facts: (1) The rise of the well after
rain; (2) the absence of effect of even the greatest floods on the river
(seo fig. 8, 1st December); (3) the diminution of the static level of thl>
wells as each additional well is put down. The Museum well has fallen
41 ft. in IHteen years, and there is a generally exp:ressed opinion that
all the wells ill the town are similarly affected. This would be thE"
natural efJect if there were a restricted supply of water, such as a rainfall
of 25 in. affol'ds. If the lowering of the static level of the wells is an
indication of the lowering of the water in the water-table at the outcrop
(and it is difficult to suppose otherwise). then the wells in the town are
robbing the crops in the country of the supply of water that they should
receive by capillary rise, a matter of some importance on light shingly ground.
HII,HKSDORF. Artesi(lll If",l" ill fhe CIII';Rtr/Htrt'/1 ArM 157
It has been often IIollSerted by mytlelf, along with others, thctt It IS meon-
ueivab1e that the rainfall should supply all the water outflowlllg at the
l.'hristchurch wolls. but I hllove madt· a calculation that. whatl'Vl'1 its fa.u1t~.
makes the case at least llot illconeeivable.
Population of Chrir.tchurch suburl!s WIthin the artesian areali
-i.e.. from SoC'khurn to New Brighton and froID Papllonul
to tht' Port Hill1l-R6.661 (say) 90. ()(}I)
Gallons of water used pel capita per duy, including hydraulic
lifts and cranes, strt'et- and garden-watering-
Auckland (1910) 58
Wellington (ma.ximum) 60
Dunedin (maximum) 61 ~
Say, 8vel"ag(' f01" Chritltchurch (where stl"eet-w!lterillA
comes from river) 60
Two rams at College lift wat('r an doverage of 22-1 ft., anu
waste water is sevell times thlt; pumped. .As this is
above average height. wo may say proportion of watt'r
used to that wasted .. One-tenth.
Then, total wat('r urawn from artetlillolltl in Christchurch
90,000 x 110 x 11 x 365 x 10
area pt'r year = "224i1 96.251.160 tontl.
Agail"-
Population having bet'n taken as from ~Ol'kbul'll to Nt'w
Brighton-
Length of catchment-area 10 miles.
First-stratum wells outcrop two miles up plain (Speight) :
deep wells (450 ft.) outcrop about eight miles IIoway;
:. width of catchment-area (about) 6 miles.
A.verage rainfall 25 inches.
1 in. of rain = in tons per acre 101
Then rain falling on catchment-area per year
= 10 x 6 x 640 x 101 x 25 96,960,000 tons.
If there is any approximation to accuracy in this calculation, then
(,lLoh additional well put down to any of the strata at present in use can
receive its wlLter only by robbing its neighbouts, a condition of affaits that,
in tho upppr strata has long ago been rea.ched. .As for the lower strata
they have probably not been largely drawn on so far. and there is eve1'Y
reason to suppose that there are still lower strata available but still
untouched.
In favour of the Waimakariri t\ssisting the water-supply a.re these
mcts: (1.) Water does undoubtedly percolate from the beds of some of
the rivers, as stated by Speight (loo. cit.), and I am a.ble to add that near
Bealey a considerable amount of the Waimakariri flows underground.
This water is almost certain to leak into every porous bed, especially where
the thin deposit of silt that fonns on the river's bed haa been removed
by scour. (2.) The great degree of constancy of the Christchurch supply,
and the smallness of the nnnual variation in the Museum well dUling
the three periods it has been under observation. I should. be inclined to
think that water from the!' river doE'S assist the Christchurch wells in
some degree, but the Lincoln well in no degree; but a longer period of
observations would b(' necessary to establish any opinion on thp matter.
158 l' IIl1lbC/CtWII ~
..,_ A iSl'
." liN
,. "I'
••
Itlt i\
t •
\ I.'
..2
M
""
1\
5'
,.'
: 1\ \ 'f\, lA It\ , r..-... A II'
."
II 3'
,
· ., 'V "i'\ ~\
3'
~ I\. V'
· ! \ ~
~
~
--
".
\_ A . ,i
"3"
, I'
... . i \ . • J \I \ , .. 1\
"IG. 9. - Hn ,PH .. ~ WELL \T BELF4.q<f (Fr'LL LINE) I:II C'ENl'I'lETRES, \"." til
\Y mIl \KARTRI (D lTTED Ll~E) IN FllET.
l'ONl'LUSIOYS,.
The well a.t Lincoln depends fo)" Its supply almost entirely on rainfall.
TnI' wells in Christchurch depend on ramfall, probably assisted by percola-
tion from the Waimakariri. Thl' wells at Belfast depend chiefly or entirely
on the Wa.ima.ka.riri. The rain supplying the ,,,ells of present depth falls
on the plains comparatively close at hand-say. "ithiu ten miles of the
tOWll. The discharge from the wells prohably lowers the water-table III
the country. The ba.rometric preBBure influences the wells.
At the close of a paper that is largely a compilation of the work of
others I hnl! a long list of helpel'S to whom to offer thanks. Mr. Speight
Bnd Mr. Symes have been good ellough to offer valuable suggestions during
the course oI thll work. The Oouncil of the Canterbury Philosophical Insti-
tute has voted money for apparatus. lIa.ny of the students at Lincoln
College, lIr. Speight. Mr. Symes. His Lordship Bishop Grimes. Mr. Crump.
and the V4'rl(f'r of the Presbytl.'ry at Lincoln, have ('ither taken weIl-
Hn.("R~"'D01ll' -.Il"tl'5l(111 Wllll~ 'II tile ('/lrlsfrl,Il1l'/' .1/1'11 15!l
observcltlOlls for mt' 01' defimtely placed wells at my U1I:IPObdl. lh. lflcl.,Y hell:!
supplied analyses of the gaseous cOlltcnts of well-waterb. Mr. W. Palnf',
telegraphist at the Bealev, has made for mt' ven careful mea.surement"
of the height of the Waimakarll'1. The followmg have supplied me with
ra.lIlfa.ll rt'coms f'lthf'l' for '!hort pl'flod" or fOl the whoh· vear: Messrs. G.
Gray and U. Rcnmc at LmLOln..J. Brunton fIJlcl R. Ellis at Rol1eston
Grrffith I'Inllth at Lawforu ..J. Wilsoll dot Ki:n'llec, J. Reid WIlsall ,\t Dal'
field. (J. Hall dot Hororata. W. Hall and G. r. Hunt at Glenroy, P. H.
Johnson at Mount Torlt'SBC. and. finally. the Govel'llment lIeteoroiogist for
several stations. Mr. Hog{/. wa" kind enough to mdke the calculatioll
concernmg the alteration in thl' volumes of the dIBsolved gases undel
changes of pressure, and Dr. Evans and Mr McLeod to provide materIal
for apparatus. To all these I beg to off!.'r my thankb. gg without then
co-operation this paper could not have been written ill its present form. I
have also to aclmowledge the IlSlllsiance giY!.'1l by the Obsl'rYcltlOllb made b~'
the late Captain HllttOll.
ART. XV.--A. NI!JW GenWl (lllIl some Neu' tJpec1es "I PI(mts.
By T. F. CHEESE~[AN, F.L.S., F.Z.S., Curator of the Auckland lUuseunl.
[Read belon the A.url:land in,tltlltr. '38t1i Soullliler. 1911.'
Many vears ago the latl' Mr. Buchanan ga\"e me two specimens of this
plant, coliected in some locality in the Kaikoura Mountains; and I have
qathered what appears to be the same at the Wairoa Gorge. In the
Yanual I included both of them with my .lE. laeta, although they ob-
viously differed in the much greater size of all their parts. I have now
received good recent specimens, collected by Mr. R. M. Laing. and from
their study have come to the conclusion that they represent a distinct
ipecies, although closely allied to M. Zaeta. I have much pleasure in
associating the plant with the name of Mr. Laing, who is so well known
from his long-continued researches into the Algae of New Zealand.
3. Corysanthes Carsei Cheesero. sp. no'".
Ab O. 'Wfl9Uioulata R. Br. differt floribus angustioribus. labello apiculo
minore, sepalo postico emarginato.
Planta perpusilla. acaulis. florifera 8-12 mm. alta. Folium solitarium.
membranaceum. ovato-cordatum. scutum. 6-10 mm. longum. Flos soli-
tarius, pro planta IDajusculus. horizontalis vel deflexus. supra folium suL-
seasilis. 8epalum posticum hasi angustum, tunc lato-cucullatum. spice
incurvatum et emarginatum. 8epala.· lateralia. parva, linearia. Label-
lum magnum. 10 mm. longum. orbiculatunl. marginihus valde im"olutis.
Columna Lrev_iij. cun"ata.
Hab.-North Island: Peaty swamps between Lake Tongonge and
the coast, Mongonui County; H. Carse and H. B. Jlatthell~fI.'
A small delicate species. i-i in. high when in flower. Leaf sessile,
H in. long, ovate-cordate, acute, membranous. Flo"'er sessile or very
shortly pedunculate, about i in. long, horizontal or deflexed, dull-purplish
Upper sepal very narrow at the base, then suddenly expanded, so that
the upper two-thirds is broadly oblong and hood-shaped. extreme tip
incurved and emarginate and slightly thickened and papillose. Lateral
sepals placed under the lip, small, narrow-linear, 4-5 rum. long. Lateral
petals still smaller, 3 rom. long. Lip large. tubular. the margins involute,
meeting behind the column and enclosing it, orbicular or broader than
long when spread out. extreme tip produced into a minute projecting
lamina, between which and the overhanging emarginate tip of the upper
sepal is the only entrance to the fl'Ont of the flower. Immediately inside
the entrance the surface of the lip is furnished with a broad patch of stiff
papillae all pointing towards the interior of the flower, and which is con-
tinued as a narrow band down the median line of the lip. At the bast!
of the lip the margins on each side are rolled up on themselves, thUti
forming two minute circular openings leading to the base of the flower.
Column short. stout, curved. Capsule not seen.
This is a very curious little plant, closely allied to the Australian
U. tmguioulata,- but, judging from Mr. Fitzgerald's beautiful drawing,
that species has a much broader flower, the upper sepal is wider and not
incurved or emargina.te jIot the tip, the projecting lamina at the apex of
the lip is much smaller, and the papillae within the lip are confined to the
median line, whereas they also form a broad patch to the right and left
of the median line in O. Oarsei. There is also a relationship to O. Mat-
tkewsii,- but, among other differences, it has a much narrower dorsal sepal,
and the lip wants the projecting lamina of O. Oarsei.
The numerous additions made to the orchid flora of the North Cape
district by Mr. R. H. Matthews, and the discovery of the present species
by :Messrs. Carse and H. B. Matthews, shows how much might be done
by careful investigation in most parts of the Dominion.
B.ABTRUM..-Roc!{8 of ]Jount Cargill. DIIIINlill. 16:}
... Trans. Au>I1:. Ass. Adv. Sci., vol. 10, 1904, p. 186; Dunedin.
t Quart. Joum. Gool. Soa., val. 62, 1906, p. 381.
t Park. " Geology of New Zealand," • 23.
§ Ma.NbaIl. " 0001010" of Dunedin." Quart. Joum. Geol. Soc., Tol. 62, 190tl, p. 390.
II" Geology of New Zea.land,"l.' 148. •
~"On GeOlogy of North Rea , Wa.ikouaiti,·· TrauR. N.Z. mAt., vol. 36, 11104. p. 418•
.... .Loc. cit.
tt Trans. N.Z. Iwrt., vol. 41, 1909. p. Ill.
6·
164 Trnn ~nrtloll ~
... « Qeology of Dunedin," Quart. Joum. Geol. Soc., vol. 62, 1906, p. 40'1.
BARTRu)I,-Roch of Jloullt Cargill, 1)1111('11111 16;)
PETROLOGY.
DEHORIPl'ION OF ROCK TYPES.
A. TRAC'HYDOLERITES.
(a.) Getu!:ral Lava T'I'ackydoleriJ.e.
In haud-lipecimen this is a heavy greyish-black fine-gl'ained rock show-
Ing fairly prominent feldspar and, in places. pyroxene crystals. It breaks
with a rough fracture. Feldspar and pyroxene show up prominently 011
weathering surfaces. This rock is describt'd by Professor Marshall in his
paper on the Dunedin tmchydolerites.*
The microscope shows a base of irregular feldspar laths, with enmeshed
aegirin«.>-aueite and other crystals. I.'n('losina model'at«.>ly abundant pheno-
Cl'VSts •
. A porphyritic character is shown by the feldspar. and to a less extent
by the nepheline and pyroxene.
The phenocrysts are sanidine. nugitE' (chiefly of various brownish tints).
l'f'sorbed hornblende. sodalite. nephl.'line. oli~ne, and occasionally aegirine-
augite and anorthoclase.
The commonest phenocrysts are those of augite.
The sanidine shows marked corrosion, and its margins are usually
dentate \Vlth aegirine-augite. Its twinning is by the Carlsbad law. The
extinction in many cases is parallel to well-marked cleavage, and in some
idiomorphic sections to the edge between the faces 100 and 010. The
crystals are usually small, but are found up to I) mm. by 4 mm. in size.
Clear glassy crystals are oharacterisitic, but both liquid and aegirine-augite
mrlusioDs are <'Ommon.
'" Tran~. AUllt. A"'Ill. _.\(h-. Hci., '1'01. 10, 1904, p. 18:.
166 Tramartion~.
The G,ollmillla".
A. network of predominant feldspar wraps around plentiful pale-green
..ll'!l;trIne - augite granules, a little fine nepheline, and a little iron - ore.
The feldspar, as a general rule. IS in poorly shaped untwinned laths.
Fluxiunal d.rra.ngement is rare. There is a little polysynthetically
twinned plagiocl£Lse feldsp!!,r a~o ~resent. The iron-ore is chieHy magne-
tlte in small squares, but llmemte 18 also present.
The nepheline is only distinguished by staining the sections. It IS in
nunute hexagonal forms.
:No cossyrite was observed in the many sections prepared of thIS rock,
but there is an abundance of the mineral in a similar trachydolerite from
lIount Flagstaff.
The granules of aegirine-augite are alwa.ys irregular, and at times
Simulate a mossy structurc. Apatite forms stout though never plentiful
prisms. O/(j,pr of C'rv.taUulation.
Some of the relations are uncertain, but the probable order is (1) olivine;
(2) apatite: (3) amphibole; (4) augite; (5) sodalitc; (6) sanidine, nephe-
line. with possibly anorthoclase and oligoclase. and then, in the ground-
rolSS, iron-ore, aegirine-augite, nepheline, and feldspar, in the order named.
Sections of a transition type of tT8.chydolerite come from a little east
ot the Main Peak. Olivine, in coarse aggregates of fresh rounded crystals,
with a. corrosion border of magnetite dust and aegirine-augite granules,
is very common. Pinkish augite has been corroded, and is edged. by
.1.ejli.rine-augite. Large crystals of resorbed amphibole are rare, but the
lllineral may be represented by numerous small groups of secondary mag-
netic material. Feldspar sometimes encloses this magnetite. Nepheline
is rare. There is a little very opaque coss~'l'ite.
The groundmass is very dense and fine-grained; it exhibits occasional
flow structure. Feldspar continues to be more important than the aegirine-
a.ugite. Ohsmical OhMactelS.
Two allalyses of the trachydolerite from two different localities are
appended, and with them. for purposes of comparison, two other analyses.
_-\. B. C. D.
50·43 -:1.9·02 51·86 50·06
18·00 19·50 19·87 17·00
3·78 4·37 6·30 2·96
5·65 6·60 8·11 5·42
2·91 2·14 2·33 8·61
&w
4'7~1
.m
1-70
.n
6·20
~U
3·40
5·76 7·35 4·88 3·53
1·37 1-18 1-48 4·85
0·38 Not det. 0·51 HuO 0·14
Not det. Not det. 0·36 0·66
Not det. Not det. Not det. 0·51
98·83 98·64 100·67 100·28
A. Trachydolerite, Main Peak, Mount Oargill. (.Analysis, J. Bartrwn.)
B. Trachydolerite, near Pine Hill. (Analysis, J. Bartrum..)
C. Trachydolerite, Dr. P. Marshall.·
D. Shoshonite, Yellowstone National Park.t
* "Geology of Dunedin," Quart. Joum.. Ueo1. Soo, vol. 62,1906, p. 407.
t &aenbl't!oh, .. EJ.eme.nte dl.'t' Gesteinslehre," p. 365. No. 13, 1001 ed.
168 J'l'flIIM,ction" .
41st) occurs ill a few large crystals The feldspars exhibit the same round-
ine: as in the general trachydolerite, but the edges art' sharply defined.
- Pyroxene and brown amphibole also form phenocrysts. The pyroxene
is generally idiomorphic pale-pinkish to pale-greenish-pink augite. It is
oOmmonly fringed by dust-like aegirine-augite. Aggregates of pinkish
augite are common.
In some sections nepheline forms important large well-shaped crystals.
~odalite in small flaky forms is moderately abundant. The augite inoludes
a, few apatite prisms.
The impenetrable nature of the groundmass is given it by the felted
dust-like grannies of aegirine-augite. Typically, no cossyritc is present:
but in a few sections, where the density of the groundmass is not so
marked as in typioal sections, a few opaqne-brown dendritic growths may
h~ of this mineral.
A few minute feldspar needles are scattered throughout. Staining
detects nepheline in the mesh of aegirine-augite dust in minute rare hexa-
gonal and squaro forms.
Magnetite is very soarce, unless it o(,('urs with aegirine-augite as a
rt'sorption-product of amphibole.
Ohemical Oha.racterll.
An analysis was made of this type, and comparison with the two other
analyses appended shows how closely it agrees chemioally with both the
trachydolerites and the traohytoid phonolites.
A. B. C.
54·24 56·19 55·10
18·08 20·25 19·25
~H8 2·76 2·77
3·53 2·32 1-66
0·88 1-12 0·83
5·01 4·30 lH4
5·01 4·19 4·68
7·29 6·33 7-41
1-79 0·65 2·19
0·63 MnOO·32
(1·57 0·48
0·09
0·54 0·4:1
Thus field relations gh-e no help ill drawing distinctions between different
petrolo¢cal types. but indicate that the origin of all is di:fferentiation of
the one magma. There is. however, a possible exception to this. for Illl
outcrop of nephelinitoid phonolite on Butter's Peaks may be a dyke.
Under the microscope the chief feature of the type is the nephelinitoid.
or cellular, structure of the groundmass. due to the nllmCrOUI4 minute
hexagons of nepheline seen under modE'ratc magnification.
The phenocrysts arc typicall~' allotriomorphic; the most common art·
pinkish-hrown augite. sodalite, sanidine, uE'pheline, and olivine.
The nepheline is large and well-shaped. but is crowded with minute
liquid inclusions. The crysta.ls of sodalite are yery large. and are usually
crowded with minute ~ra.seouil inf'lusions: they show good dode<:ahcdral
cleavage.
Sanidine is clear and glassy, hut exhibits shadoW' extinction. A feW'
characteristic anorthodase phenocrysts occur.
Olivine has (>ither 0. dense corona of aegirine-augite with maglH~tite. or
else a corrosion fringe of aegirine-augite and magnetite dust.
Pleochroic aegirine-augite is shown in tI few well-shaped crystalll that
have suffered slight resorption, and haye been edged by the common
pinkish augite. This latter variety sometimes encloses resorbed amphibole.
showing that it did not separate out till after, or was connected \nth, the
resorption of the amphibole.
The grOlludmass is holocrystalline but finE'-grained, aud generally similar
in minerals to that of the other types of traehydolerite.
Aegirine-augite in mossy granules is dominant; if often assumes a
lath shape, and then shows more or less parallel alignment.
In sections' of those rocks that, both petrographically aud in field
occurrence, approach the nephelinitoid phonolites cossyrite occurs plenti-
fully, but is absent in other sections. unless some minute opaque dentritic
growths can be referred to this minerol.
Feldspar is moderately important, and enwrapping laths show up
amongst the numerous minute hexagonal forms of nepheline. Yer~' little
magnetite is present: there are a few s(·atterl'd. fiak('s of sodalite.
No analysis of this rock 'Was made.
B. NEPS:ELIYITOID PHONOLITE.
As ""ould Le expected, in certain places this rock merges clos('ly IDto
the type of trachydolerite just described. In se.eral sections segregations
or inclusions of the basic trachydolerite previously described are .ery
typical. The~' a.erage about 7 mm. in diameter, and are most probabl~'
of the nature of segregations.
Leucite again appears as a subsidiary mineral. It is difficult to dis-
tinguish it from numerous other rounded isotropic forms that are judged
from their ready gelatinization 'With dilute acid, and from the high percent-
age of chlorine in the rock, to be sodalite. The leucite is in small rounded
or idiomorphic shapes, and commonly shows chllol"&cteristic radial arrang~"
ment of included aegirine-augite granules.
In hand-specimen this rock is very similar to the dense basic .ariety
of trachydolerito, but has a somewhat lighter colour and greasier appearal1ee.
It weathers very readily.
Under the microscope true phenocrysts are rarely set'n, unless in the
proximity of the 'basie inclusions, where pink augite a.nd oliyil1(, especinlly
are common.
BAllTRUM.-Rorks of Mount Cargill, D,medil/. 171
BASANITEs.
lIention has been made already of the so-called basanites. In the
hand -'3pecimen they are :fine-grained dense blackish rocks. showing few
conspicuous phenocrysts.
Besides the occurrences already noted, a basanite is found as extensive
rubble near the hypabyssal trachydolerite on Pine Hill.
Under the microscope these rocks show a typically dense base and ,\
paucity of phenocrysts; augite is the commonest of the few phenocrysts
that are seen. It is most often of a pale-pinkish colour, but a green augite
,vith pink botde1' and a purplish-violet variety are also present.
In the basanite found north of the Junction School a strongly pleo-
chroic mica is prominent. It occurs, along with a little serpentine. as an
alteration-product of the olivine; its p!eochroism varies from deep brown
to bright golden-brown. It is thought to be anomite.
Feldspar seldom is a phenocryst in the Mount Cargill basanites: a few
very large feldspars show albite twinning, and seem to be oligoclase. They
are, however, so crowded by augite granules and other inclusions tha.t an
exact determination cannot be made.
The groundmass is very dense, and is composed r_nainly of small grains
and squares of magnetite.
In the anomite-bearing rock the magnetite is less important, and an
interstitial feldspar is thc chief constituent, along with grains of colourlesR
augite. The augite is usually in fair amount in these rocks, but feldspar
typically occurs only in a comparatively few needle laths.
No chemical analysis of any of the basanites was made.
RELATIONSHIPS OF THE TRACHYDOLERITE AND PHONOLITE SERIES.
The analyses given in the subjoined table show how gradual a passage
there is chemically from the more basic trachydolerites to the phonolites.
Petrological characters also indicate that such a gradation js Dot a matter
of chance, but represents a differentiation of many types from the one
magma. In certain cases this is due to difierences in the rate of cooling.
All evidence from the Mount Cargill area would show that the Logan's
Point trachytoid phonolite is u. portion of the main alkaline magma, and,
in fact, a modification of the trochydolerites and thtl nephelinjtoid phono-
lites. Evidence from other parts of the distri<.'t dispels any idea of its
contemporaneity with these latter rocks.
A. B. C. D. E.
50·18 49·02 54:·24 54-·88 56·12
18-00 19·50 18-08 22·80 21·32
3·78 4·37 2·18 3·66 2·59
5·65 6·60 3·53 3·26 3·29
2·91 2·14 0·88 0·38 0·56
5·76 6'76 5·01 2·24: 2·30
4:·79 1-70 5·09 3·65 4·81
5·76 7·35 7·29 7·53 5·79
1·37 1-18 1-79 0·91 1·64
0·38 0·63 0·63 0·34
98·83 98·64: 98·72 99·94 98·66
A. Trachydolerite, Main Peak, Mount Cargill.
B. Trachydolerite, near Pine Hill, Mount Cargill.
C. Basic type of trachydolerite. Butter's Peaks, Mount Cargill.
D. Nephelinitoid phonolite. Butter's Peaks, Mount Cargill.
E. Logan's Point trachytoid phonolite, Mount Zion, Mount Cargill.
178 1'1'1111 "1I' /f)U ~ •
t
t-!UAXTATIVE CLASSIFICATION.
The quantative classification of this seli.es of MOWlt Cargill rocks has
been worked out by the method of Cross, Iddings, Pirsson, and W"shin!!-
ton.t The following is the r('8ult:-
1. General la.va trachydolcrite-
(,'lass II. Dosalane.
Ordel' 6. Norgare.
RanI! 3. Saleruase.
Subrang 4: Salemose.
Leaves 14-22 em. long, I-I! em. broad, fairly uumerouE., btrlCt. nano,,"
lmear-laneeolate, entire, slightly coriaceous, gradually tapenng to the
dcute tip, marked by clolOe parallel longitudinal glOoves or fint' wunkhngr:.
above and less prominently below; upper surface yello,,:illh-green. l.llabrou!l
or glabrescent, the tips bometiml's loosely tomentose on both I>urfacee..
under-surface, except the midrib, covered with loosely appressed pall'-vello\\
cottony tomentum; margins more or less recurred; sheathb about' 6 cm.
long, thin and membranous, glabrous on the inside, cottony-tomf'ntose on
the edges and outside. .
Scapes 1-4, rather slender, flexuous, blightly rIgid, nea.I1r tWIce as long
as the leaves, densely clothed with creamy-yellow loose cottOny tomentum :
bracts numerous, linear, thin, tomentose except on the IDlduh'l. graduall~'
diminishing towards the top.
Heads about 2~ cm. across; involucral bracts numel·OUE., IInE-ar. thm,
cottony.
Achenel! glabrous or slightly hispidulous.
Hab.-Sealey Range, Mount Cook dish'iot, ill tUbl:!ock meadow, about
5.000 ft.; T. F. Cheeseman, Mrs. F. Bowe. and D. P.
This species is dedioated to Mrs. F. Bowe, a keen observer dud ardent
lover of our native alpine and subalpine plantb. who first dIrected my
attention to it. Mr. T. F. Cheeseman, F.L.S., collected it a good many
years ago, and he considers it a fonn of O. Moltrol Hook. f. This -new I
am unable to entertain. It differs from O. Mom'oi in the narrO\\el. lebs
coriaoeous, more acute leaves that are green above a.nd very distllloth-
grooved or finely wrinkled; m the yellowish loost'ly appresr.ed tomentum
tha.t olothes the under-surface of the leaves and the sea pes ; and in thl'
more slender flexuous soapes.
Gentiana Mattbewsii sp. nov.
Planta subgracilis, ramol!8., glabeiTima, annua (?), Ij-2j dem. alta.
Caulia a basi ramosus; rami graciles, adscendentes vel lIuberecti, sub·
quadrangulares.
Folia radicalia pauca.. subrosulata. spa.thula.ta, tenUla, 2-4 cm. longa.
± 6 mm. lata, obtusa vel subacuta; caulina sessilia, late ovata vel ovato-
triangularia, subaouta, basi lIemi-amplexicaulia. 8-12 rom. longa, in PBlibus
distantibus disposita.
Flores liubnumerosi, solitarii, albi, 10-15 cm. longi. ramulos ultimos
terminantes.
Calycis lobi ovato-oblongi. subac-uti, corolla persistente fere dimidio
breviores.
Capsula. matura breviter rostrata. eorollam superans.
A rather lIlender branched glabrous annual (?) herb. 1!-2~ demo high.
Stems branched from the base and again more or less subdivided;
branches slender, ascending or suberect, more or less distinctly quad-
rangular from ridges running down from the balSt's of the cauline leaves.
Radical leaves few, subrosulate, thin, spathula.te. 2-4 cm. long, about
6 mm. broad, obtuse or subacute; ca.uline 8-12 mm. lo~. sesRile in distant
pairs, broaqIy ovate or ovate-triangular, subacute, ISt'mi-amplexicaul.
Flowers fairly numerous, solitary, at the tipr. of the ultimate branch-
It'ts, 10-15 mm. long, white.
Calyx divided for three-quarters its length, half as long as the corolla ;
the lobes ovate-oblong, subacute.
Stamens rather longer than the calyx-lobes. Ca.psule whl'n matUl'f'
one-qua.rter longer than the persistent nearly dosed corolla.
184 1'ran~actlfmf.
ART. XIX. -Thp Jliql'atiol?8 0/ the Pol.Vne8ians (wcllrdinq tIl fhp E"ine1k'I'
01 fheir Lflnquage.
By Professor J. lIAclrILLAN BROWS.
I Reua be/ol_ tllr IJ'fnlll'ltoll PIII[o,opltll'rti 8m:lPt!l. 6th ~ept.mller. 1911.1
IN th~ "Transactions of the RClyt~1 Scientific Society of Gottiu/l.cu" for
1909 there appears u. long paper on this subject by the late Professor
Finck. of Berlin. It attempts, as its title implies, to point out some of
the distinctions between the "arious languages of Polynesia. a.nd by this
means to indicate the lines of mi~ration that peopled the:' islands in which
they are spoken.
The gist of the arguments a.nd conclusions is given in the last two paged,
and is somewhat as follows: From the southern Solomons a really united
people shifted to the northem fringe of Polynesia on their eastward trek.
Before the expedition turned southwards to Sa.moa the ancestry of th('
present-day Ellice and Tokelau people branched off. The speerh of that
time possessed all that marks Polynesian as contrasted with the rela.ted
Melanesian. t'sperially the:' use of the old tr~al as plural, and the employ-
ment of sepamte possessive pronouns where once only a suffix ,vas used; it
was, in mrt, probably the fundamental Polynesian tongue. The use of air
for" a thousand" docs not contradict this. although it appears in this sense
only in Fakaofa. Futuna. Samoa. Tonga. U,·ea. and Niue; for the word
is, as thl' }laori au·hE' shows. common to Polynesia; but it was extruded
in the other dialects by mano. There was a long rest in Samoa, as is shown
by the u~1' of fnkelau for .. north" and tonga for south ,. in a ma.jority or
<t
the groups. words taking this sense from the direction of the Tokelau and
the Tonga Groups from Samoa. After a small colony had swarmed off
westwards to Futuna, the great eastward-going expedition went south-
wards to the Tonga Archipelago, as is shown by the use of 11, in all the
groups to the south and east for 8 in Samoa and its immediate neighbours,
and by roe use of toko as a personal prefix to words implying number and
quantity in all to the south and east for toka of Samoa, Fakaofa (the Tokelau
Group), Vaitupu (Ellice Group), and Futuna. After a. short rest in Tonga
the expedition went off eastwards. leM;ng a contingent which sent branches
to Niue and Uvea. III the Cook Group it made a long sojourn, and there
fonned the growld speech of eastern Polynesia; it changed 1 into r and
I into 11, before 0 and Il, brought the.> Bdnominal particles Ra and no into
use beside the older 0 and 0, and abbreviated the old possessive tou into to.
From this point YarioWl expeditions set onto One went to New Zealand
and the Chatham Islands and developed ]I for f before other TOWels than
a and 0,' it left before:' the counting by pairs arose that characterizes the:'
other eastern Polynesian dialects. A second went off south-east to Manga-.
reva; thence a branch hiyed off to Easter Island. farther in the same
direction, before the birth of the liuguistic neologisms that unite the di..'Llects
of the lIarquesas and Ha,,-aiian Uroups ,vith that of MangareYa, the forma-
tion of adverbs hy prt'fixing ma or InO to a noun, and the change of to'ktrlW,
into tokurau. It wa.s 10llg befoTe this northern expedition set out-long
enough to deVelop these peculiarities. The Marquesss Group developed
as linguistic cbaracteristirs the pronominal form toia and the further
duplication of numeration by pairs in the case of rau; (the~ equal to 4(0)
and mano (there equal to 4:,000) before sending off the Hawaiian branch.
Meantime from the Cook Group another colony Ilh-ed off to Tahiti, wh08t'
190 l' 1'1I1l~art/(J1l'
11> to s,n-.. Dwlekte ''') as contrasted Wlth the countles!> Yarlet,- ot not
merely ciIalects, but langua~es, ill the Melanesian region and the ·Malay.:m
region. if properly considerl.'d. might have saved thenl from the latter
mistake. Even the few centuries which thev seem to have in their minds
as COVerllll!, the history of the human race in Polynesu~ would ha"e developed
language& B& distinct as, say, French and Spanish, or English and German.
If we were to take into account the marvellous simllarity of the Polynesian
dialect& not only in phonology and grammar, but in vocabulary. spread
over a.n oceanic region as \vide as Europe and Asia combined, we would
not be :f.u Wl"Ong in concluding that there have been thousands of migra-
tion& from every island to every other island; in short, a new sketch-map
of the Polynesian migrations should so completely cross-hatch the central
PaCIfic that it would look bla.ck. In other ,vords, for centuries at least
intercourse must have been almost unbroken amongst all the groups. If
this means anything, it mea.ns tha.t for a prolonged p<>riod doll the Poly-
nesians lnUbt have inhabited .:L large island or a.rchipelago centra.lly SItuated,
and also quarantined from other regions under a social, if not politica.l.
s:"st~m that was pl'8(·tically do unity. The minute dialectic differences that
arose must have been kept in bounds by the constant social intercourse
that a single administrative system would allow-a system absolutely
different from that of Melanesia or of Malaysia. The differences are no
greater than those that separate the dialects of, say, Yorkshire and
Somerset. or Scotland and Middlesex.
The consideration of the culture conveys the sa.me impression; the
ethnological differences are as negligible as the lmguistic when placed beside
the points of agreem('nt. One can find as wide variatlOns of culture and
dialect in the purely German part of the German Empire. They seem
to have arisen ill the presence of each other, as well as of the predominant
community of culture. In other words, they must have slowly developed
during the immense period of time that certainly was taken to produce the
'Practical identity of culture and language. This identity would have been
shattered into strongly contrasted fragments had it been compelled to run
the gauntlet of the limitless variety of Malaysia and Melanesia, not to speak
of having to sail right in the teeth of the south-east trades, the only fairly con-
stant wind on that l"Oute, the contrary wind being brief, fitful, and cyclonic.
There is, of course, a striking similarity between the languages of Poly-
nesia, Melanesia, and Malaysia that makes many speak of them unitedly
as the Oceanic language. But there is a phonological gulf between the
Polynesian dialects on the one hand Ilnd the Malaysian a.nd, still more, the
Melanesian languages. Each of these two regions has its own range of
sounds, with considerable community; but Polynesian has the peculiar
and distinguishing BOunds of neither-it has the simplest range of sounds
that ever language had, all easily pronounceable by Aryan and, one may
add, by Japanese organs of speech. It has a similar contrast in voca-
bulary: with. anyone of the Malaysian or Melanesian languages except
Fijian it has never more than 20 per cent. of common words. It is the
grammar that has led to their classification as one language; for none of
them have practically any formal grammar-they all move in an atmosphere
of particles, and there is a very considerable resemblance in the particles
used. But this absence of formal grammar is the commonest characteristic
of crossbred languages - i.B., languages that have resulted from the per-
manent or continuous settlement of a masterful people amongst & people
liDauistically different; the formal graroD'atical peculiarities of both are
192 r, (I n ~(/( f /(1/1 ••
~"
I •
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....
: l
::
r>
-
-
...
=:
~
"
~
:::: :
S ~
5
WAITE.-Xott'b on Sell' Zealand PIsllfl'. 195
The outer bronchial is free only m its posterior h.a.l£, the anterior portion
being a.dnA.te to the ceratohyal. This attached portion only bears gills:
they are much sma,ller than those of the other drches, on which they are of
considerable length. There IS no trace of paired arrangement in the gills
of this outer arch. A paired disposition is apparent in the gills o{ the
two middle arches, for, though the rays are set in continuous series, they
are of heteracanth nature. The inner arch is wholly adnate to the memo
branes at the lower part of the tongue, and is fully furnished with gill-
rays. The gill-rakers are spiny tubercles; there are 12 on the :first arch,
one of whioh is on the upper limb, just above the angle; the rakers on
the median arches are in two rows, arranged alternately, there being 19
on the second arch.
Fins.-Some idea of the character of the dorsal tentacle will he derived
from Clarke's figures, but as it \vas evidently imperfect. and is even more
complicated th.a.n drawn and described. the following des('ription will not
be out of place:-
The tentacle lies in a deep groove between the supraorbital ridges. its.
l>ollbous base bemg rather nearer to the mouth than is the eye: the shaft
is very stout, and it tenninates above in a larlle semi'lpherical bulb, ita
total length from base to summit being 1·7 in the length of the head.
From a cup in the summit of the bulh arise!> a freely movable stout
tentacle, which divides at a short distance above ita insertion, each branch
throwing off 2 smaller twigs at about half its height. Inserted in the
bulb and behind the cup are 2 thick b 'anches, which, however, arise
from a common base: they become flattened distally, and each, after
throwing off a twig from ita inner side, divides into 3 arms; these are
again subdivided, but the divisions are not the same in the two branches
The illust:ra.tion accurately depicts the condition. Also, on the hinder
part of the bulb, but nearer its base and sides, are two other small twigs.
The word "frond" would perh.a.ps be more appropriate, for the whole
tentacle may be likened to a plant of F'UCtI.8, the so-named twigs being
quite like the fronds of a seaweed, while the main and secondary stalks
answer to the stem and branches of the plant. There are, in all,
20 terminal fronds, and the distal portion of each is nacreous white,
a.nd is no doubt luminous in life. When the tentacle is bent forwards
these luminous tips dangle just in front of the mouth, and are no doubt
very efiective lures. It will be apparent th.a.t the tentaole was in('omplete
in the type specimen, the stalk arising from the middle of the cup bE.'ing
absent, and doubtless leading its author to conclude that the substance
within the cup was luminous. though he does not a.ctually say so.
The dorsal fin has a slightly more forward insertion than the anal. and
has one more ra.y. The first is simple, the other four beinp; dividE'd nearly
to their bases. The third is the longest. being 3·2 in the head. The
last ray is connected to the peduncle, just free of the upper caudal ray.
The anal ia very similar, but the first two of its 4 rays alO simple. The
pectoral is short and rounded, and is placed nearly midway between the
cmd of the snout and the base of the caudal rays. The caudal is large
rmd rounded, arising from a very compressed and short peduncle, whose
depth is equal to the longest dorsal ray.
~f'mOu,.-The skin is soft and loose, warty on snout and chin, and.
~ting the top of the head, cheekA, lower law, and all parts in front
bhenof, studded with round cartilaginous acutest each of whi('h bears in
if8 centre a h.a.rd low thorn with roots rndiating into thE' body of thE' SCllte.
WAITE.-NoteB on .Vtu· Zt'aland {I'tBke •. 197
Some of the scutes are much larger than others, and their exact number
and disposition are shown in the illustration. The covering of the main
stalk of the tentacle is formed of a mosaic of very small scutes. wbich also
hear spines, but they are reduced to hard tubercles.
CoZou",.-After long immersion in preservative the general colour if!
a pale-flesh tint; the margin of the jaws, the post-dental frenum, the space
around the eyes, and the wart-like elevation on the chin are brown; the
mid-line of the back and part of the stalk of the tentacle are also brown:
the branches of the tentacle are black, but their tips are white.
Some Measu"emtmt,.-Extreme length, chin to end of caudal, 410 mm. :
length as basis for comparisons, 270 mm.: length of head to gill-opening.
135 mm.; diameter of eye, 10 mm.; length of tentacle-sta.lk. 78 mm. ;
extreme len~h of tentacle. inclusin, 205 mm.
Fac." 195 J
WAITE.-...YofeB on .Yew Zealand Fulles. 199
pectora.l fin to a point in advance of the first dorsal COllI'; it thcm't' drops
to the mid-hue of the body and passes along thl' middlf' of thp caudal
peduncle.
OoneB.-The remarkable cones which give the fish such a sbiking appear·
anCE' are disposed as follows: The swellings on each side of the dorsal ridge
above referred to support 2 pairs of Iilmall size; the hinder pair lie at the
base of the dorsal spines. and are directed out,vards; the pair in front of
these have a more upward aspect. All the othel" cones exiRt on the ventral
portion of the body: the largest form a series of 5 pairs disposed along
the lateral margin, the centre one on each side bein~ the largest, and
directed strai~ht from the body, those before and behind being diver-
gent. A smaller cone is placed immediately in front of each ventral fin,
and a similar, though larger one, on each side of the ,"ent between these
"entral and anal cones. There are 3 pairs of much smaller ones, which
thus complete the vertical armament. In these latter each cone is set
close to its fellow. The mosaics in the mid-ventral line form small tubl'rcles,
but quite distinct in size and character from the true cones, which. as will
be seen, number 12 pairs-namely. 2 dorsal, 5 lateral, 2 subventral, and
3 ,'entral. The cones, which arise from an enlarged series of mosaics,
a.re as hi~h as, or higher th.a.n, their diameter, and are sculptured with both
radiating and transverse striae, the former being straight and the latter
wavy. The area between each radial is :flat. These correspond in number
with the basal mosaics, of which there are 16 surrounding the largest cone
-namely, that in the middle of the lateral series.
OoZnurs.-The ground-colour is brownish-yellow. and the markings
form wide open reticulations, consisting of a black line merging into bluish-
grey, which extends so as to nearly obscure the ground-colour. The latter
remains fairly pronounced on the cheeks, the lower edge of the caudal
peduncle. and an area at the base of the anal nn, due to the absence of
maddngs on these parts. The membranes of the first dorsal fin and of
the anterior ventral rays are black: the other nns are colourless.
Lengtn, 80 mm.
One specimen only.
There is a temptation to give the Pacmc fish a distinct specific name,
not onlv on account of certain described differences in the two known
individUals, but also in consideration of the widely separated habitats,
the one being taken in the Atlantic and the other in the Paomo Ocean.
The fish must have rather limited powers of progression, for its locomotory
fins are feeble, and the general conformation of the body is opposed to
even moderate progress. The original specimen was supposed to have
been taken in the surface-net, and, as the New Zealand specimen was
secured alive on the beach, it becomes fairly evident that we have either
two very closely allied species, or, like TetragOltllrlf8, a single species of
pelagic habit, of which examples have been obtained from both Atlantic
and Pacific Oceans without any intermediate occurrences.
The differences noted between the two specimens Inay be due to certain
chara('ters in the smaller one having been overlooked. It is unlikely, for
example, that the 3 spines preceding the anal rays were absent, or that
the ventra.ls had only "Ie nombre ordinaire de 115." Other differences
may be noted. in the descriptions of the dental arma.ture. The French
a.uthors apparently found teeth in both jaws, wherea.s my specimen exhibits
them in the lower ja.w only. There is agreement as to the presence of
teeth on the vomer, but I find none on the palatines, their presence being
200 'l'ran~action&.
• ... ~'9ier &lid V!!len~1IIIJiEis, Rist. Nat. Poise., vol. 4. 1829. p. 515, pl. 99 (0. r,(),,'.
/mJm). ' . .
t GiiD.ther, Cat. Fish. Brit. Mus., vol. I, 1859, P. 214; voL 2, 1860. p. 396.
t Lowe, .. Fishes of MadeiI:a.." .:m.
§ Goode and.Bean, Ocea.nio Iclth., 1895, P. 228, and fig. ,
II Boulenger Camb. Nat. HiBt.. 'Fishes .. 1904-. p. 68:~.
TRANS. N.Z. INs7.. VOL. XLIV. PLATE XII.
oS...==
....."
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0
iil
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~
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0
~
~
p
p
==
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Fuca p. :eoo ]
WUTE.-.'·ote~ on Xew Zta!lwd Flh/u, 201
the peduncle lOll!!, and narrow, its depth one-fourth more than the diameter
of the eye.
Scales.-Head ~enelally naked, but with soales on the operoles; upper
palt ot head with a. t3pongy porous integument. The body-soales are not
markedly deciduoug. a.re of moderate siY..e, and finely dentioulated; they
extend on to all the, ertical fins. The lateral line does not follow the curve
of the back, excepting for its anterior half, the hinder part being almost
straight.
Lengtk, 945 mm. The type was 990 mm., doubtless measured to the
end at the lon~el:>t caudal ray.
Colou7s.-Steel-blue above, silvery beneath.
The genus Eurumetop08, of which E. ioknstonii is the type and only
known species, may be thus defined: Body oblong. compressed; snout
obtu&e; mouth large; teeth present only in the jaws. Premaxillariel'
slightly protractile, maxillaries with supplemental bone; they are not
entirely ooncealed by the preorbitals when the mouth is closed. Opercula.r
bonet3 thin, entire: branchiostegals 7; gill·membranes united far forward.
not attached to the il'thmus. pseudobranohiae developed; gill-rakers long:
scales of moderate size, fairly adherent, lateral line not concurrent with
the dorsal profile. A single dorsal fin with about IX, 20 rays; anal with
about III, 15 rays; pectoral pointed, with 20 rays; ventrals below the
pectorals. Vertebrae 22.
The qenus appears to be sufficiently established, and finds its nea.Iest
ally in PsenopsiB Gill, differing in the larger mouth, the oharacter of the
maxillaries, the more adherent scales of relatively smaller size and their
development on to the bases of the dorsal and anal fins. The lateral line
is not concurrent with the dorsal profile, and the number of rays in the
vertical fins is noticeably smaller.
The following notes are supplied for the convenience of those wi&hing
to make a further comparison: In 1862 Gill* erected the genus PSe}i(YjJ61~
for Tracl,ynotlls ollonlolus Schlegel. a &pecies taken in Japanese r.eas.t
The affinities of the fish were previously recognized by Bleeker (1853),+
who placed it in the genus PS6nes. Regan§ has more recently added
Batltyseriola oyallea Alcock,1I from Indian seas, to the genus Psellopsis,
remarking. ., There can be no question that these two species belon!!. to
the same genus, although their relationship has not hithel'to been suspected.
and the two Rpecies are yery closely allied."
E...~PUNATION OF PLATES.
Pu..TliI X.
•!((lOfOldc11,1'!/8 flP1J1'lii Clarke. Les'S than half no.tural sizp.
Pu.TE XI.
OI£08OlIIa Iitlaltlirllflt Curler and Valenciennes. Nearly twice natural size.
PLATE XII.
EUfll.!netop08 jolltlBtollii Morton. One-fifth natura.l size.
ART. XXI.-Ncw Speo&e8 of Lepidoptera, wIth 'Yotes 011 the LarLoae and
Pupae ot some Sew Zealall(Z Butterflies.
By GEORGE HOWES, F.E.S., F.L.S.
[Read before Ule Otaqo lilstitulp. 1at August. 1911.]
line in ochre. The veins crossing this area marked in golden. Ciha dark
ochre, barred with brown. The markings continue on through the hmd-
wings, the oIlly difference being that. there is more golden colouring. and the
cilia are light ochre barred with brmvn.
The small size of this insect, together with the triangular - shaped
golden ma..rking cutting across the other markings, mal:es this moth
very distinct. It is with some hesitation I place it in the DasYllriB.
It mav have to be removed later.
Taken on the Game Mountains. near Nevill, 20th Novemher. 1910.
Morrisonia pansicolor sp. nov.
Three females, two males; 29 mm. Head dond thor.u ochreous, tllightly
tinged with rufons. Antennae filiform, rufous. Crests well de.fined, dotted
with rufous. Abdomen
ochreous, in ~ dotted
with minute dark specks,
ochreous-rufous in c!. with
dtrong crests, especially the
anal. Forewings ochreous,
suffused with rufous; all
markingtl rufous. Subbasal
line double. much broken,
double line at 1 bending
MOBIUSOlUA PANSICOLOlt. strongly outwards at centle
of wing. A mark on costa
at i, followed by two marks over reLLiform, \vhich continue through reniform
as faint jagged lines across wing. ~\.n indistinrt !luutt'rminal line forlllt'rt
by a series of dots.
Orbicular obsolete.
Reniform :filled with
dark rufous. Veins
faintly marked with
rufous. Cilia ochreous.
Hindwings ochreous,
centre of wing clouded
with rufous brown.
Discoidal spot well
defined. A faint series lIoUIUt>UNIA MOLLIS
of subterminal dots.
Cilia whitish-ochreous. with a darker line at base. Underside pale ochrtloUI:I_
OlllVed post-medial line across both 'vings. Reniforlll and d18coidallunule
well defined.
Taken in November, at Dunedin, at .• treacle."
The moth is so close to molliB that the first specimens I took I
thought were that species. Subsequent captures, which gave me both
sexes in both species, placed the matter beyond doubt.
My last illustration of molU8 being so unsati.t:lfactory, I am giving
¬her drawing of it, along with patliscolor. In mollis the reniform is
clear, in pcmiBCOlor :filled with dark rufous.
Morrisonia sequens sp. no'".
C!. 31 mm.; ~. 84 mm. Head and thorax grey, strongly crested.
Antennae filiform. AMomen ochreous grev. crests slie;ht. Forewings
HOwES.-Sew SJitcie~ of Lepidoptera 205
lJOBlliBOXU. PASCOEI. ;( 2.
edged with white, projected nom the head, and a similar hom from the
tail. .A white line from front of head along thorax, then splitting into
208
two thIn \.hlte hne.. to endose cl. dar1..-~Jeenl&h dOl'ldl hue A thIn led
hne m c.on] unctIon Wlth a whIte hne from tIp ot Iro'ital holU to tlr of tall
hom T\\o wmter hnes from wmg-cover to tIp of lhdomen A V.hlte
Ime ed!:.eJ both 'JIde'! \uth led from centle of wmg (.d~e not 163.c.hmg to
end ot abdomc I A dark hne with a. "hIte hue belo\\ .tlou~ the top of
"'mgs Yem.. of tutUle 'i\1l'~' deal Iv outIm.ed As the Int:.ect neared
emelgence thE' duk ::.pot... ')'1 111 \' m.!~ '1ho\\ed plllniv thloJ.gh the pupa
.uun
(..d"['E(TlO,\
In the ' IrJ.lll:>J.(.t on'> ot roe New Zealard Inl:>btute," \01 4) lUll,
pp 127, 12~ I find I have calelt.SsI) Wlitten ' hnes ' "hele It ... ho lid
bo "IDm' UutOltUJoHteh, thIS not only mel1..e'! the descllptlon le.ld
.... 1'Onsdy d'! to fhe v.ms;-e'\.p tnqe, llUt hab cl.l"o ml,:>lead those lespon",lblt.
f01 the lerlOUUc.UOLl ot the lllu. . tl thon .. '10 th. t tlleqt ha... e been }lllU1ed
mucll o\er thell II Itt I II ..,,/
By B C .\.STON F I (' , F C S
[R~ad bPlal 1/" II tlltn !Ion l·h,1 n.( pIlla" "'0 pt I ./tl, (JtfobeJ 1911 j
Pl.a.te... \..Lll XI'
CtPE TUR.4.I\.IL.Uo IS the "lOlth "estenl porot uong. CcIoPl P,,'h""'1 tiht. two
tapes lllelo,>'ng that noble stretch of wcltLr known as Pallise.l R,1\ TIlt.
rem.lolkable t!eolop;lCdl and bot'll leal flcl.tUl'e c 01 Tnra1..n&e hitherto apPGQl
to ha,e escaped the attentIon the) lllt.l.t a.nd It IS \uth the hopt" of
remedymg this neglett that thll> p1.fl.r d. \\rltten
The Orongolongo RlVer, neu the mouth of ~hILh IS 8ltuated Air Rldd!
tOld's hOME'Stead, cuts through cl senes of rl1bed beaches, nov. more or les'
obseuled by drlftlllg Blond or o"elgto\\n b, herbage Thev are, mOllO,\E"
('omposed of hner IDcI.tenal-lllosth (,Oc.\l"bll band-than those SJome I Cdochl..,
d. Imle neal"Cl the lape Thell" finer llatUl"C Ib ac.counted for by the fac.t thclot
the nvelS "ould bnng down q1lcllltltIes of fine debns, wluch '\ould be
thlO\\n up by the SC<lo Three beae.hes are here to be dlstInguIshed, and,
as their altltl1de IS the s'lome the, are undoubtedly of the same a.ge as thoDO
fhroc helea.fh.'l descnbed as beme: Dearer the sea The beach pl'f"sumably
elevated at tht' bme of the. 185:> ea.rthquake IS here very v.ell de\eloped
The nUluence of the imeness of beil( h parttcles on the flora. Wlll be presentI)
notIced It IS not until one has (.loBbed the nver, cIond proreeded a wIt
01 so towards Palhser Bay, that th~ eyo of the obselver IS arrested by thl."
e:1.tr"ordmary ph)BlOgraphJeal aspett of the eounny lymg betv.eell thE'
trac1.. at the base of tho steep hlllbide and the sea The trac1.. follo,,& clo
COUlse close to the foot of the Mia, about 100 £t above sea-level, and
betuen thts and the sea 18 a stff.hh of roc1..y country v.I.lVlng from 250 to
400 yards in mdth. and extendIng some t\\O or three llules, narroWlU~ to
nothm~ on thp further SIde of the ecl}lf' ThIs loeb plmll e.on'll'ltfl 1l1Obtl}
P \Tr \'III
...
.!
is f S
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PLITE XIV
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.r
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.III
j
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of large boulders 3 it. to 8 ft. in diameter, but running roughly parallel wlth
tho sea throughout the length of the boulder-strewn plain are two excellently
defined shingle beaches. These stand out most conspicuously, and form
Ilatural roadways along which one may drive. For the greater part the
shingle presents an appearance differing little from that of beaches which
.)ften eXIst now at the ocean's margt'. In many places the shingle is, how-
ever, overgrown Wlth MUelllet1.bec'kw comple.ra, or with grasses and other
plants. The main Impression left 011 one's mind is that marvellously little
alteration has taken pla.ce m the peophng ot these areas by plD.1lt&, and ill
the extomal appeara.nce of the shlllgle generally, 1ll the hundreds of years
which have probably ela.psed since each "'as rd.ptdly elevated. The slU'\"i.yal
of the 'beaches as shingle hwolves the fact that it is composed of the harder
p.:>l'tiions of tho country rock, and which would hence, in the equable
climate, offer a considerable resistance to the weathering influences.
isolated by bouldcr pla.ins on all stdes, little dust could blow in and form
. . Qil bctween the lUtersticcs, and without soil little atmospheric moisturll
could be retained. Only specially adapted shingle - plants, such 11'>
.l!ltd"Zenbec1.ia, could, therefore, hope to survive in such a. station.
Happily, we are not cntirely in tho dark as to the rapidity \'lith which
thcse beaches may be elevated beyond the reach of the breakers. It id
well known that the coast at Mukumuku was elevated 9 ft. during thl'
earthquakes of 1855 (see Crawford, Trans. N.Z. Inst., Essay, vol. I, p. 18).
Knowing this, the author carefully searched the boulder-strewn shore "
little above high-water mark, and was rewarded by finding traces of a
shil1gle beach about that altitude above high-wa.ter mark. Further search
nearer MukumulLl showed a long strip of shingle beach quite as well
developed as the oldor beaches. The fact that the sea is now breaking on
boulders and monoliths somewhat discounts the thought that beach No. 1
may be a mere storm beach. Exploring the country adjacent to the hills,
two much older shingle bea.ches were found. These nearly everywhere
bve been oblIterated by the debris carried dovm by temporary creeks from
the steep hillsides, the site of the older beaches being now occupied by fans
of angular shingle, mixed with finer detritus, many acrcs in extent, which
m.ly or may not support a flora. This recent alluvium has buritld these
two older beaches many feet below the surface, but where fra.gments of
them remain one is a.gain struck with the extremely recent appearance
·,f the beach. a.s Plate XIV, fig. 1, truly depicts. The interesting fact that
the younger of these beaches is that more thickly populated. by a flora is
probably accounted for by the difference in size of the component stones,
which explanation must also suffice for the fact that much of the newer fau-
material supports dense formations of herbage or arboreal 81owth. Five
c.listinct shingle beaches have now been mentioned, which for the sake of
ease of reference may be designated by the numbers 1, 2, 3, 4, and C, No. ]
being the youngest (the 1855) beach and No.5 the oldest. Observations
taken with a surveying aneroid show that the level of each of these beaches
is practically conSliant along its entire length-that is to say, bea.ch No. 1
is approximately 9 ft. (see Plate XIII, :&g. 4), beach No.2 is 40 ft., beach
No.3 is 60 ft. (see Plate XIV, fig. 2), beach No. 4: is SOft. (see Plate XIII,
fig.2), and beach No.5 is 95ft. (see Plate XIV. fig. 1) above high-watoT
mark.
The material of which all this elevated country 18 composed hall so far
been roughly classified as bouldem and shingle, but there is a third most
extraordinary component, the solitary monoliths which stand out some-
210 Transactions.
Kl:lUl:A.DEC I Sl.A~DP,.
'fhe conspicuouM te:~ture of the a.vifauna of the KerDllldecs iii its pau-
oity of land-birds, and what few occur there are of a de<>ided New Zealand
oharacter. There is pl'actically no Australian element represented.
The migrants inolude the two cuckoos mentioned abo\'e as occurring
in Lord Howe and Korfolk. Islands, and six Charadriiformes. None occur
in any great numbers; thus tlleir presence affords but little indication of
the island l~'ing in an ancient land-line. If a large area. of land evel'
existed in this direction, it disappeared before the present islands came
lDto existence.* The occasional and accidental visitors 111'(' all common
New Zealand species, and all except .AnthtlB nOllae-:ealandiat occur also in
Anlltralia.
The Kermadeo Islands are of a typical oceanic Chlll·llcter-tho.t is, they
were formed by volcanic aotion in mid-ocean, Ilnd have been popull\ted
THE LOUD HOWE - XOBFOJ.E TflLAND PRO¥INOE AND THE KERlI.l.DEO ISL.l.ND~
PnOnYCE.
Ha,ing' 0;;}10\\"11 b~' ,.Ill ulluly!:!is of their avifaunas that Lord Howe, N01'-
folk, and the Kel'mauec Islo.ndl:l fall within the limits of the New Zealand
biological rcgion, it remains only to oonsider whether they form one 01'
more Ilrovincc~ 01' subl'cgiontl of that region.
The I).,ifllunatl of Lorll Howe and Norfolk Islands are undoubtedh'
closeh" allielI. Of the resident land-birds two species are common, t:l
whioh must he added the endemio species of four genera - Gerlluone.
O,/anornamplluB, Turdu8, ZOBtel'opB-having representative forDls in each
island. }'Iost of tIle Hell- biIds (breoding), migrants, and occasional
visitors I1.re common to the two islands, but, from the manner of their
oocurre-nce. this iR not the case with the accidental visitors and the sea-
birds (visitorR).
With regard to the Rix resident land-birds of the Kermadec Islands, all
except Proatllemadera nOI'(le-zealandiae are identical with or closely allied
to species in Xorfolk Island or Lord Howe Island. But of resident land-
birds in Lord Howc or Xorfolk. Islands found also in the Kermadecs the
proportion iR very low. NO that the fact that five-sixths of the Kerruadec
land-birdN IlI'C related to IIpecies found in Lord Howe and Norfolk Islands
merely shows that in nIl prohabilit~·, as with the plants, some of the species
in the Kel'JIIlldt'cK which occur in both ~ew Zealand and Norfolk Island
ha.ve arrive 11 il'OIll Xorfolk I!:!land direct. The sea-birds (breeding) and
migrants chiefly belling til the same species as occur in Lord Howe ann
~orfolk Islnndll. TIlt:' occasional and acciden.tal visitors, like the resident
land-bil'ds. al'l' VI.;'I'~" few. n faot which can be explained by the isolated
position of the isl:mrlR.
The three iRlanns therefore fall no.turall~- into two biological provinces.
One includes Lord Howe and Norfolk Islands, and is charactorized bJ" an
avifauna containing foul' main elements: (1) an original element whioh
includes the species which came by the ancient New Caledonia - New Zea-
land land bridge (majority of resident land-birds); (2) Australian and
New Zealand species arrived subsequently by trans - oceanic migration
(some of the resident land-birds and all the visitors except migl'ants);
(3) circumtropical species (sea-birds. breeding); (4) migrants. The Ker-
madec province oontains only- (1) New Zealand (and Norfolk Island)
species arrived by trans - oceanic migration; (2) cll'oumtropicnl specieS!
(sea-birds, breeding); (3) migrants. Although the important and charac-
teristic element oircumtropioal species is common to the three islands.
and, in the ca.se of plants, formed the basis of. my subtropical island!:!
province,* I think now, on a consideration of the avifauna, that a more
r ill!fUI'II-JJ igralltll.
•\.relllLl·ia interpl'es. LimosR novae-zealandiae •
('haradrius dulllinicllK. Tringa canutuR.
XUlllt!nius varie~at\l'l. Gnllinagll austl'ali ...
X. c~·anopus. {'halcococcv.:s: lucidllll.
Erolia nUl·ita. Eud~·nam.;s taitenKill.
E. ft'l'ruginea.
r i/lif(JlII-Oualllllltul.
PUl'ph~Tio melanollotuR. Circus gouldi.
rhal'ndl'ius bicinctus. EUl';vstlllllUlI pllcificu~.
Xutophoyx novae-Illlllandiae. l'uculuH inlll'natuR.
X \'cticorax caienonicuil. ('acolllllntiil l'ufulm•.
AntlN superciliosa. Cm'acil1a l'ohusta.
Phulncrocol'nx sulcil'ostris. G1'8I1inll picnta.
P. llIelanoleuc3.H.
r isitCJrR-.AcciueII taT .
Phapli clegnntl. •\stUl' novae-hol1anditle.
Himantopus leucoceplulllllol. Haliaetus leucogaatel'.
Hel'ouias tiruoriensiK. Halinstllr sphenuruR.
Al'd~ttll pURilla. C'helidon neo:xena.
220
NORFOLK ISLAND.
Sea-birds-lJre.e.dillY·
PuffinulI liphennrus. Anous stoliduB.
P. assimilis. .\.. leucocephalus.
P. grisem.. Gygis alba.
Oestrl.!l:ltu philippi. Sula cyallops.
Hterna fuligiuuqa. Phaethon l"ubricaudn.
ProcelRterna ciut.·rea
Visitor,-8 e.a-birdlS .
Uestrelata macroptera. Sterna albistriata.
Puffinlls rerneipes. Ossifraga giga.ntell.
Megalestris antarotica.
Visitor,-Jligra"tll.
Arenaria interpres. Limosn novae-zea.landiae.
Charadrius dominicus. Chalcococcyx lucidlls.
~umenius variegatus. Eudynnmys taitensis.
Glottis llebularius.
ViBiforB-OCCaBional.
f'harudrius bicinctus. Phalacrocorax sulcil'ostriH.
Notophoyx novae-}lOllandiae. Eurystomus po.oificllfol.
Anas sUl'erciliosn. Circus gould i.
ri,itors-Acciden till.
Himantopus leucocephalus. Larus novae-hollnndine.
Plataleo. regia. Astur approximans.
Herodins timoriensis.
KERMAOEC ISLANDS.
Re8idtmt Land-bird8-Brttding.
Proethemadera novae-zeaIandiAe. Anaa auperoiliosll..
Halcyon ve.gans. Porzana plumbea..
Cyanol'hamphua oynnurus. , Hemiphaga. novae-zeall.l.ndia.e.
Or.IVl<:n -Bzrd~ of Lord You'e, .\"orlull.:, altd ll.ermadec islalld, 221
::lea-UI rdB-Breeding .
.::Itl'lllJ. fuliginosa. PUffillUS ~lJhellUl'u"
r:'I'gi~ ,tlba. P. assimilis.
Anous 1tlucocapillus. Ot.:strelata cervicali~.
Pl'ocel'lterno. cinerea. O. neglecta.
Phaethon rubl·icaudn. o nigripenni:..
:-! ula. cya.nops.
ViBitor~-Sea-bil"dd.
J)uption capensitl. PUtilllUb tenuirostri...
Diomtluea tl::i:Ulam,. ~'rtlgatn aquil..1
D. lIIdanophrys. Prion rlesollltIH•.
Phocbetl"ia fuliginosa. Ollbtn·luto. macl"opteru
Ptllugodromn. marina. :-Itema ht'l"gii.
:3Uhl leucogaster.
fis/fors-.UifjraM".
Charadrim. rlominicus. E.'olil.l. .W 'itt.
C. veredul!. Limosll novae-zealandillt!,
Tringa co.nutus. Chalcococc'i'x lucidu:..
XUIIlenius variegutm.. Elldynull1~:':I taitt>nMil"l.
Vi8itor8-0rrfl~lullal,
Circus gouldi. Anthus Iw\·ac-zealanrlill.c,
Zosterops caerulescens. Phalucroc'll"a'C sulcir"lIt'rh•.
ViBitors -Acci(lentnl.
Hypota.enidia philippensis. Demiegrettll. ~acl'a.
Porphyrio melanonotus.
Cd.nterbUl~· Ilwl W~btl,mu," pp. :HU-22. This i~, II.b fILl' nb 1 ha,'~ heen
ablc to glean, tht' total l~fclCnCtl in publibhed ltlports to thi~ intel t'bting
locality. The prebcllt account il> necebsd.lily imperfect, but it iN ba~eJ on
work dom: on llUlIlt'l'OllS vh.ih., 011 ho of which I "111> fortlluute ill having
the Itrlvi~ ana a~to.h.t.UlCtl of Dr. Mall!hall, to whom, ab "ell al> to MI'.
::lutel' fOl' valu ••bltl hdp ill illel1tif~'ing the rOfosil&, lily 'IinCelll thankR lire
tIUtl.
(jE:"l'ER.!.I. D.Ill:IUltIPTION OF THE LOC.o\J,Il'Y.
(See map.)
The Jibtrict rtlfel'l'ed to in thi~ account lies, 1 oughly, to tht: 1>1Iuth-
tld.llt of thtl point whtlre the ra.ilway 1 unning north frolll Chlit.tcliul'ch
Ol'ObbeS the Waipard. River, and f01 the purposelol of lllOle a«.CUl'ate Ildi-
nition the lower gorge of the river 1IlILy be taken as that part of its
course \,hich lie':! between ith junction "'ith the Omihi Creek <11111 thtl hca.
The creek joinb the ri\"61 about two lllil!::s below the railway·crobbing, unll
is historically important, since it flows from the swamp where Haaolt
obtaintlti his Glemnark collectioll of mOIL-l emains. TIlt' actual length of
the gorge is about fOUl wiles. Its bides are lllodel'ately ~teep, eabilv
climbed in lllllst plact!s, but unscalable in PIlI'tS, and ribing to an Ilverage
htlight of about 3;)0 ft. above the river-beu. 'l'he~' are higher on the
eastern siae, "hence thl!~' extend itS It IIb'etch of irregular dO"llb towlI.rus
the slopes of :Uount ('abll, which forllls the bouth-welltern buttrf::'Ss of the
Limestone Range. On the western side of the river the banks are not 10.0
high, anll they form part of the low do"nb I:Itretching to the houth-west
towards Amberley. The breadth of the gorge varie':!, but it usually pI'e-
bents a wide floor covered with Wiingle, on whioh the rivel' wanders. At
times, however, its brearlth ill reduced to about 100 yards, 01' even less,
and then the river-channel is more tit!finitel;v fixed. In some places the
stream forms well-defined loops or meanders-a notable one occurs about
half-way through thtl gorgtl-and it is now destroying the spurs which
project laterall~' from the high banks into thel!(> loops. Although the
",tream has considerablt! iall, the large amount of detrital matter which
it transports frolll higher levelb, anti ~pecinny from the banks of lool:le
..hingle bordering itll COUl'sa through the Waipara Plainll, hILS so dimin-
ished its vowel' of erollion that it has apparently 1 cached a temporary
base-l~vel, and this in I:Ipite of the fact that within fail'l;v recent tin1ell-
oertainly t!ince the Pleistocene-the con!.t hail experienced a distipct up-
ward movement .
.\. recent upward lllovel.nent of the coast-line to the north of the
Waipara has lJ~n recorued by McKay at Amuri Bluff (Ueport of the
Geological Survey, 1874-76, p. 177), whert' benchet! with Recent shells
al'e found at a height of :>00 ft. Evidence of the RaDle movement at the
mouth of the Conwa~' and at 1\[otunau ill given by Hutton (Report of the
Geological Survey, 1~73-74, p. 54), where the land has certainly risen
1;)0 ft., and, judging from the featuretl of the remarkable plain of marinl'
denudation noted b~' Hutton and examined lllore recently b~' the present
author, the elevation hall in all probability betln much greater. Renl-
nants of this plain ovel' a mile in breadth are to be found on both sides
of the mouth of the Waipara River. A little way back from the present
beach is an old sea-cliff about 50 ft. high extending along the coast for
several miles, and from the summit of this the land slopes gently back
for about a mile, the upturned edges of the beds forming the solid eub·
stra.tum. of the country being planed oft neatly b~' the former action of
the sea a.nd then covered with u thin veneer of loose shingle, some of it
SPEIGHT .-Lower Trai para Gorge. 228
LOWERI •WAIPARA
!&.
DISTRICT.
,. ~
Scale In Miles
FIG. 1.
'1J'1':PIMlrlllJn~ •
of lual"iue oI"lgin, IIncl of material \\ hicl! closely l'eseIllble~ the loells of the
Bouth-eal>!wrn portiCJn of the South IsI'lIld. 'l'his plain sloVes bllck to the
base of the low clowns near .\.llIberley, "hert! it reaches a height of about
1 jll ft. Hmwver, on looking up tht! CllaI>!t to the north from the wont h
of tltt! Wuipara, decided l"emnant!l of a t-.hore platform can be seen I~t an
!!btiUlatt!ll ht;ight of 230 ft. above the sea, and fronting the plain there
art' suvl'rul small remnants at lm\er levtlh. marking stationary pcrious
!luI'ing the vrolongt!d elevation, It is therefore ccrtll.in thllt a long stretch
uf coast-line has experienced the effect of this movement. 'flw,t the ril>(J
ip of recent date it! \'"er~- clear from the species of shtlils founu at MotullBU,
and also il'OIU tLl.' forIllS of the iltrell.l1l-vallers that haye been esta-
blisherl on the elevaterl marine shelf. With few exceptions, the streams
which I'un acrostl it have ,el'Y short courses, anrl are little more than
c\tcnrlerl gUllit!b or wa..houts: Through thE" some"hat 100KE" Momnau
llmrls, whloh form the s'llirl ba!le IIf the bnll, the~' have eroded deep
dumnel'l, at time$ liver ] 00 ft. in Ilepth, extremely narrow, and witl,
liiclelS so prccipit()ulo. that they arl.' I(lJ~(llutel:r illlPo.lI~ltble for long di,.tanceb.
Th(' whole plain itl dissected by thelU. and they render cOllltlmniclltion a
lUatter of difficulh" "here roud!ol and tl ncks do not e'l:ist. 'lhe district
furnilihes a flll)Rt rClllUrkabll.' ('}..amplt! or the effect of a recent npward
land-movement on the gradienl and crostl-section of the stream-channels.
The phenomenon is intensified by the Ilnifol'!l1 ReD.warcl dip at mocleratt"
angles of the beds llDller tile plnin, Imd tht! parallelism uf the strike to
the coD.st-lint'. A I'imilar phenomenon is to be obserl"cu near the mouth
of the Waipara. but the featllreR are not quite so perfect.
This plain of lllarille denUllation once erlended much fmthcr sea-
ward, and the small island at lIIotunau is a remnant of it. its Hat top
lIhowing a marked alignment of its surface with that of the coast-line
opposite. How far this plain extended seawards it is impoSilible to say at
prE"sent, but at the month of the Waipara the river-terraces appear high
above the present level Ilf the water, and are terminateu Ruddenly when
they I'each the edlle of the old marine cliff which mnrks thE' edge of the
coas1>111 plain. .tt a former period the river ruUl>!t have extended WllCh
further scaward, anll Howed on the top of the plain, the terraces with
their shorn ends giving J,ositive proof of its higher level and seMmrd
el.1:ension at that level. ..\11 river erosion was proceeuing the sea was
eating back the margin (If the plain, thus giving the streams a steeper
gradient and increasing their erosive power, and the truncated ends of
thE" terl'aces mark the limit to which the plain was destroyed. When one
takes into account their perfect ('()ndition he must conclude that E'ither
terraces are stable land-fm'JU!I or that marine orosi on on this Btl'etch of
coast haR been vary rapid and comparatively recent.
There is also evidence of a more recent land-movement still. Along
the bast> of the old marine cliff, referred to p1'eviouRly as bordering the
{'oastal pla.in, there is a p,trip of flat land consisting- of shingle-beds, sand-
dunes, and swamp, half II mile wide and but slightly raised above the
liea. It has, without rlouht, heen formed of detrital matter brought down
by the rivel'tI in thE' vicinity. IIllch as the A.shle~" and WaipBrR.. as well as
by the E.Ultlll titreuDlS which flow directly into the sea, their load of
waste beinl! distribnten 1l~' waves, tides, and currentfll along the base of
the old cliB. These nccUlnulationf. are Reveral miles in length, and their
Rize suggestll that there has been either a remarkable increase in the
Rupply of detritml or tbat there has been a small recent upward move-
llIent of the coast. TherE" ill no apparent reaSOli wh~' the 'ltreams should
hl\ve been 8uddenly 1\11'nillhel1 with an inereaserlloud of d(.>trihlR, nlthougb
8PUI.lH'l'.-I.ulI't"I' ""'PUIlI (Jol!/e. 225
~t Ul~bt be no!ed that they a.rc full~' charg!:!cl at thtl pr!:!~llt tim!:!, and it
IS tlUlte conceIvable that the~' could build up It t!hallow ~a-bottolll till it
was above sea-Itl\"el without any chauge in the lldv!:!l of the lallrl. I call-
not, however, think that this exphmation is altog!:!th~r l!atisfaetorv and
concluJe that a IS!Ullll and }Jroblibl~' clllltinuoul! u!,lift halt taken 'place
aftel' a co~uparatlvdy long peril)i! of btuLilitr, during "'hie]} th~ old
coastal plam was eaten back to the line IIf the forlUer liica-cliff.
.r udging ft'om the profiles of the ..tre.1!llt! joining the WaiV81'3. fro111
the :Banks of the Deans Range and elwwhcl e, thib movel1l!:!nt has extended
!ts f.,flects some distance i;nland, Tl,e lu(,p~ of thtl river in the gorgE>
ttsell. placed as they are m a !!ollle~,hat narrllw trellch, llla\" o'l'.-e their
urigin primarily to the fact that at a fortHer period the ri;'er reacherl
b8Foe-level, anrl that the gorge W8S cut ~ltl\\n t .. Ib, Ill'esent depth ,luring
a sub"e'luent period of elevation "hell the river had increased power to
corrade, anti that now it hlu; again .1lmost adjub1:ed its grade to thc load
it carrie!!, and all it!! er(,bive encrgT is (levoted to destroying the loops
th.lt it previoul'Ily- tormed. - It is -Very- ilifficuIt, --!n",wever, to correlal..
thebe effects with eel tainty.
ORI .... I:-i OF THl=: W.UP,\RA GORGE,
Th~ circumstances reRulting in the fOl'luation of the Waipara Gorge
furnii'lh on~ of those interesting problems with relSpect to drainage direc-
tion!! for which the North Canterbury district is noted. mf.'n tbe river
lea'l"es the hill!! between ~nt Bl'own altd t1e Dt:ElnH, and iRSUl'N fronl tlJe
middle gorge, whose existence baE> been largely determined by the great
~lid-Waipal'a fault, it pUl'I!ues a course lIf about seven miles acr08S the
Waipara Plains, and, instead of taking the eaRY path to the sea paRt
Amberley, it cuts a somewhat deep channel through the downs which
stretch south-west hom the tel'ruination of the Limestone Range, Here
it runs practica.lly along the strik.e of the beds which form this somewhat
elevated ground. At timet! it breaks acrost! the strike for a short dis-
tance; still. the coincidence:! is very marked, even when the strike swings
round through a right angle. When the river lea'l"es the downs and
debouches on to the coastal plain it pursues a dit'ect course to the sea,
still following the strike approximately. There seems to be no reason
from the present configuration of tIle ground why this difficult path
should havt! been selected whE.'n an ea!!" one was ready to hand, so that it
is apparentl:," one of the instances o(the allomaillus' behaviour of rivel's
which the diRtl'ict furnishcs.
The Waiau and Hurunui, a few miles further north. and even the
Waipara it~lf in its upper portion, have cut gorges through mountains
composed of hard greywackes and slaty shales of Mesozoic age when they
might easily have avoided the obstructions. The only satisfactory ex-
planation is based on the fact that they are instances of •• lIuperimposed I I
drainage, In late Cl'etaceous and eal'l~" Tel-ti81'~' times an archipelago
of small islands fOl'med of rocks of Lowel' Mesozoic age occuJlied the area
now known as North Canterbul'"Y and the Amuri districts, In the Rtraits
and bays among these i!Jontls, 'greensunds. solid lim!:!stones. mal·1s. and
loose inooherent calcareous sands and gravels ",el'e laid flown, so that
the original surface was completely mar.ked, ""hen the land 'WaR raised
a.bove the sea in late Tertiar,- timeR the cOUl'se of the streams established
upon it was largely determined by the form of the land as it emerged.
While cutting down their channels the stream!! removed a large part of
the veneer of loose and readily eroded material, encountered the h~d
underlying rockR, cut into thenl, and maintained their original direc-
8- Tr£lIl!,
226
tions. As time went on 11l0re and more of the loose sediments was re-
moved, and the original f01'111 o.f the In:nd-sul"face c~ll~}Jletely altered.
The existing Tertiary depos1ts a~'e m all probablht~· but a smull part
of those originally laid dow-n. for lsolated fragments of thelre ro~ks arc
found in various places high above the present vallt!y-fioors, .and In such
positions that the~' nla~' ·well ha,,·c .formed part of ~n extens1ve she~t. I
think that this is the bel:lt explanatlOn of the e",[)lut10n ?f the land ;n .the
district, as well as of the anomalous co.urll6b of tl~e r1",er8. A slmllar
explanation was also indicated b)' Captal~ Hutton lD a short parag.rap~
contained in hiil paper on "The Formation of the Canterbury PlaIDS
(Trans. N'.Z. lnst., vo1. :11. 1905, p. ·167). _.
It is quite possible, howe"'t!r, that the presenctl of the Lower "iIlpara
Gorge in the peculiar sit~ation in ~hich it now occurs may be due to a
small coastal sheam cnttmg: back lts head thl'ough the escarpments of
harder I'ocks, capturing the headwatel'S of othe~ small streams, and finally
tapping the W~ipa1"u.. itself; but t~e eAplitnatlOn based on the fact that
--it Is " case tit lSapel'lwpoSClt-'"' tlnrinaKc fih. the case bellt.
STlUTIGR.4.PIlY.
The quebtion of the strlltigraphy cnn naturally be elucidated by a
com}Jl~rison with other known localities. Fortunat~l~·. the Mid-Wo.ipara.
and ilie 'Weka Pass (in close proximity) have become classic in the history
of New Zealand geology, having been reported on by nearly all those
who have done field·work in this country. ~ thetle typical localities the
following ill a representa.tive set!ltenc~, starting frolll the top, of the beds
that have been recorded : -
8. Motunau or Greta Beds.-8ands and conglomerates, mostly ca}careoulI,
with 811e11l:l of MollltlBca in varying states of preservation, but
usua.lly fragmentary. The beds are generally loose and incoherent,
but a.t time concretion ary.
i. ;\lount Brown Bells.-Rough cnlcal'eoutl sandstones with harder con-
cretiollary bands. markedly fossilif~rous in places.
6. Gl'ey l\larl.-(il"e~· and greenish sanclstones and blu~ sandy and cal-
ca.reous cI8~·s.
5. Weka. Pass Stone.-Glauconitic nnd tllightly arllnaceous limE!stone.
4,. Amuri Limcstone.-Fornminiferal and argillaceollH limestone.
.3. Greensands.-:Uark(!dly glnllclInitic in the upper portions. anrl with
conoretions full of sam'ian remains in the lower part. These beds
are often argillaceous. ferruginoul'l. and calcal'eous, and at times
exhibit marked efHol"eRcence of sulphur.
:1. Oyster - beds, oontaining: shelll:! of OBtren, 00llrhofh/lra pa'l'asif7"cn,
fragments of Belemmtes, Inoerrnm1/~, and other RhE'lls.
1. Sands a.nd claYII with brown COllI and impul"E' ltmonite.
The lower pOl'ti?n of this .se;ies is more cnml-'letely developed further
to the north-east, In the OlUlht Creek and at Amul"i Bluff. According
to Hector, Haast, Hutton, Park, nnd perhaps McKar. the sequence is
broken by unconformities. placed in di.fferent positions by the different
authors, but it is very probllble that it is quite conformable throughout.
However, it is only the upper part of the sequencl- wiili whioh this paper
is specially concerned-that is, with the Motunau and Mount Brown beds,
and the Grey Marls and Weka Pass beds, which underlie them. The
whole of the bankl'l and terraces bordering the Waipllra River as it passes
through the lower gorge consist of the sands nnn conglomerates forming
SPEIUHT.-Lower WttiJlnt"ll Gorge. 221
the highellt members of the bequence record~u above; the othill' OOUI! are,
howll~er, in cloioll proximity to the river on its east!:!l"ll !lid!:!. '!'he former
will be reflll'red to hereafter as the Parcorn series, !leeing that their fossil
fauna shows a close relation~hip to that of the typic.!.l Pareora locality.
STRCC'l't:llE .\XD .Ann.\:li'I.lElfENT OF 1'FIE DEDS IX THE GORUE.
(See map &ncl ~eetio:J.).
At the junction of the Omihi Creek with the Waival'a Ri~er the beds
COlibist of !!and!!, sandy clay!! with concl'etionary band!!, and collglomerateb
with shells. Thtl strike is N. 5° W., with a westerl~' dip of about 22,0 .
.A. good exposul'e is to he seeu whera a recellt flood hall removed the
surface covering of shingle, and laid hare the solid bank jul!t opposite
the mouth of the Omihi Creek. In the Omihi itSlllf, just above the
junction, the beds :strike north-eallt nnu clip to the llol'th-west at 450.
It is evident that the flh'ike here swings 1'ound lIomewhat, a fllatul'e which
will be readiJ:' under'stood whlln the general structural filatures of the
locality are comi(lilred lllt!:!r, thil! IImoll movement being onl;'1' 11 pnl't of
olle of widll erient.
On following the rivt)r down from the junction, the high banb on
the east are founu to be ohHCured by soil anu slip-material, but after
going about :i00 yards the ail' ub~tlrved at the jUllction changes to
the south-east, with the lIame &trike as before. The stl'ucture il-l thutl
,
anticlinal, and tIle Sallltl anticline can be uistinctl~· trllced fOJ' over a
f
i
&
FIG. 2.-SmoTIox nOlI OMlHI ORBillt :rEBOU8H BILL'S BILL TO Su (FIVE MILES),
to thl::! 'iouth-cast of Mount Grey. but ill man~' cases the pebbles have been
weathered out of the conglomerates which form a fairl~' large percentage
of tilt:! bedR of Pareora age in the vicinity of the gorge.
A careful comparison of this list with thc list (If present specicij of
.1lolluBca fountl fOHsil given by Suter (Trans. N .Z. Inl:l1., vol. 42, 1910,
p. 8) MOWS that more than 30 pOl' cent. of the Rpecies given above are
now living in N'I.!W Zealand seas. Although the list of Waipar8 fossils will
no doubt be gl'eatly amplified by more careful searl.'h, the relative propor-
tion of ilpecies to those existing no,," is not likely to be much altered .
•Judging rrom this percentage, the beds should be cla&sified as Upper
:'\fiocene 01' Lower Pliocene .
•\. furthel' comparison with the list of fossilH foulld at the typical
Jlareol'a locality, in South Canterbury, shows that of sixty-foul' named
species given i~ the Waipara list thirty-two IU'e to be found in the lihts
of species collected at PartlOra given in Haaet's " Geology of Canterbury
and Westland," in Park's paper ., On the }t[a.rine Tel·tiaries of Otago
and Cnnterbul'Y " (Trans. N.Z. Inst., vol. 37, 1905, p. 530), and among
the fossils exhibited in the Canterbury Museum. This is at first sight a
HOmewhat small pl'oportion, but the forms common to both include a very
large number of characteristic species, and it is possible that furthel'
collection may bl'ing about iurther accordance. In any case, the number
of characteristic genera common to both localities renders it a matter of
certainty that the beds in the Lower Waipllra are contemporaneous "ith
those in the typical locality at Pareora .
.A further comparison with the list of the fossils collecten by Po.rk on
the Mount Donald escarpment (loc. cit., p. ;)40), and with the lists of
Mount Brown fossils given by Hallst (" Geology IIf Canterbury and WeHt-
land," pp. 306-11), IUld alRO by Hutton in hill paper on the" Railway-
cuttings in the Weka Pass" (Trans. N.Z. lnst., vol. 20, 1888, pp. 261-62),
shows a certain amount of agreement. of the Lower Waipara fOBSils with
those collected in the typical :\[ount Brown localities. There are, h()w-
ever. BODle differences, notably the absence of Brachiopods from the Lower
Waipara, in l1larked contrast to their ext1'8ordinllry nUlllbers at Mount
Brown. ThiN mll~' be (Iue either to the fact that the propll!, horizons for
these fossils have not been discovered in the Waipara, 01' that the condi-
tions for their existence 01' fOl' theil' entombment were not favourable in
that locality when the beds were laid down. The accordance of the foslli1
content is, however, sufficiently close to assign both tletll of beus to the
same age, especially vhcn the associated iossill:lpecies fl'om othel' 10cl\liti~s
of the same age lI.l·e taken into consideration. The stratigraphiClal rela-
tions o.1so strongly support this conclusiOl.l.
Since by fa.r the greater number of the fossils enumerated in the list
can be collected on one horizon in the gorge-i.e., jUlit above the Grey
};lal'l&-it ill l'enRonable to consider that the lowest bellR intersected by the
river are of the IIllme age as the :\Iount Brown beds, while the upper mem-
bers are probably of the same age as the llotunau 01' Greta beds. and the
confol'mity of the sequence in the gorge Bupporf.s the opinion of Hutton
that the Mount Brown bedl:! are the base of the Pareora system, and the
absence of Ilny unoonformity in the gorge also supports his contention
:-!P1UGH·r.-holilU W(1ipara (';01[11'. 233
that tht!le i!!nu loltratigraphical break bt!hYt!cn tiLt' hase of the Muunt BlOwn
heds an(I the top of the Greta bed!!. *
The circulllstances are also iavuurahle to the position maintained by
1£al'blla11, Speight, and Cotton in the paper on the Tel'tiat·~· series pub-
lished in last Teal"s " Transactions of the New Zt!aland InRtitute "-that
there is no stratigraphical break in Korth {'anterbury between beub at
the ba~e of the Waipara system characterized by the preseuce of saurian
remain!', Belemnites, {'ouchothyra parasiticfl, Ilnd vliriout! bpeciel! of Tri-
(Ionia, anu beds which have a fauna which lUll.,t be assignerl to the :\Iiocene
01' even the Lower Pliocene period. TbiR statement does not, however,
negative the exitltence of a palaeontological break.
COXCLt·SION.
For the purpollt: of aiding people who ",hill tu exnmine this interesting
locality, I make the following suggestions as to the means of visititlg it.
The lower portion of the gorge is best" orked from .AmberIey, which is
distant about three miles, with 0. good road suitable for driving 01' bicrcle.
The upper part can be reached 1110st conveniently from Wll.ipara, whence
a walk or ride of about two miles will bring one on to the upper entrance
to the gorge. If time it! limited, nnrl only one day is available for the
visit, the Glasnevin Rnilway-t!tation affords the shortest aIleI readiest access
to the mirldle part of the gorge. There is a good road learling from this
station to within a short distance of the place where abunuant fossilt. ure
to be found. At either Ambedey 01' Waipnl'n there nre hotels at which
accommodation can he obtained.
EXPLANA.TION OF PL..\TES.
In considering the map and section nccompnnring this po.per it mlUlt
be uoted that recent alluvial and marine deposits haTt! not been markeel.
It was founu imposl.!ible to do this accurately without examining almost
every acre of the eountry; only the underlring solid beds are, therefore,
represented.
*1n Tram. N.Z. lllbt.. \"'01. 37, 1905. p. JaS, Park HaYS, .. The Motunau ~ lie on
a. denuded surfa.ce of the lIount Brown beds, and the section is 80 c1ea.r that no doubt
can be- entertained ab to the unconformable relations of the two formations" ; but in a.
recent paper published in the "Geological Magazine" (5th December, vol. 8. p. 548) he
admits the ph;rllica.l conformity of the Mount Brown and Motunau beds. His exact
words are, .. The unconformity which I thought I recognized a.t Waipara. between the
Mount Brown and ::uotunau Iled~ may not l'XL'It, or, if it doe'!, it may be purely local.
In my clallllifi.cation of the .Ta.mger formations adopted in my • Geology of ~ew Zealand'
I have recognized only one physical break lin the Tertiary successiOll]-namely, one
between the Oa.maru and W&lpara. &eries. Nothing I have seen since the publica.tion of
that work has led me.> to a.lter the opinion I the.>n expressed." The author is therefore
glad to know that his position as to the conformity of the lIotunau and lIount Brown
beds iii quite in agreement with the most recently expre'lSlld opinion of Professor Park
on a somewhat important point in our Tertiary Rtratigraphy.
234 'l'ran lUll'! lon~.
I HAVE to thank Mr. P. Goren for identifying this !!pecies, and alsu
Professol' Kirk for his mun;r kinunesses tu me .
.lliUl.ls distinctuB is a small black spieler belon!o(ing to a genus which
includes two oth~r species-.Il. pal'ado:EuR and Jf. /lCllldageri.
hence when the door risell the projection catches the back of it and
keeps it at an angle of 60°. On 11l0st doort! these device~ are present ttl
.l certain uegree, but some doors are without them. It is difficult to
"a," whether 01' not these nbove-ruentioned devices are rnarltl on purpose
b~ the spider .
. Tile JIan.ner ill wlu'cll the 8pide/' makes ller Lid.-The spidel' begins
I,,· wtlaving a tag-like piece of silk on the hinge side. Having collected
fi;le pieces of earth and stones neal' by, t!he gums them one by one to the
tag. .After she has got a little patch gummed together she turns around
in her nest, applies her spinners to the little mosaic, and spins a silken
layer under it, which binds it tempol·al'ily. She then goes on gumming
the pieces together till the door-opening is covered. She then ngain
turns arounu, and spins another oovering over it. In thiR IItate the door
is fiimsy ano. elastic, and when the spidel' pulls at it from within (fig. III
it drops into the mouth of the tube. Some spidel'S perforlll their builll-
ing in a night. others in some dnJl!. They generally work at night,
although loIollletimes in daylight. The g,um appears to he exuded frOlll
the mouth.
TIlt' Tllbultll' ('(mfll of tlle :rest.
The depth and width of the tube val'ieM g)'eatly, dependiug. of course.
upon the size of Rpider. Generally the nelltlol are from 1 in. to Ii in. in
lenbrth, and Ilk in. to f in. in width, taken ltlngthwise at the lid. The
tube noes not remain a uniform width in its entire length, but widen~
at the terminus to ! in. (figs. 9, 10, 11, 12, &c.). The l'eRsons for thib
terminal swelling are t"o-firstly, til allow room for eggK nnd young;
antI, secondl~.. to allow the spidel' to move and turn around in the nest.
Linin[l of Tube.
The interior of the tube is lined with a layer of silk, which is thick
if the ground is dr;v or crumbling, thin if it is damp and firm. l'ht'
rim of the month of the nt!~t iR g-enerallY' thicket· than the other linings.
enough for her to pass through, but that after a "hile she ntlglects
to enlage one of them, and onlv attends to the othel·. I have set::n small
tubes with tvo '\"ery fine doors'j large tubes with ho proper 4io01"lo are
rare. Why the spider should build a door at each end of her nest is
harn to say. I believe that &he must lose sight of thtl faot that she has
,drcad" lllade one door, and, as there is no ending 01' terminus to the
nest b~lilt under a htone, &0., as in a normal nest built in all earth
bank, t.he naturally makes a door at each end of the tube. Afterwards
hhe Ub~ unly one door and neglects the other. This suggestion credits
the Rpide,· with little intelligence.
LIFE-liISTORY.
About the lllonths of February and March the female lays from thirty
to sixty small white eggs. These are plaoed between the wall of the nt'6t
and a piece of silk stretched across a rounded put of the tube (fig. 10, O.
the arrow).
The egg-bag is placed variously in the nebt, but ill fig. 14 (top)
the commonest position is shown. The piece which stretclles across the
sides of the tub!! is fin. in dIameter. Separate egg-cases, with two sides,
not connected to the tube, are rare. The young !!lllerge and lie dormant
inside the case. After a week or so tht! mother l"tmlo\"es the covering,
and often weaves in lieu of it a transparent filament-like veil of silk
over them (fig. 10, b).
After the :\"oung become strong enough they push out of the covering•
..nd wander out of their old home. Often some few remain with their
mother, and frequently ale met with as late as the end of April. Thebe
must be provided with food by their parent, for they are often a fair size.
When the young emerge from the parent nest the majority are pounced
upon and eaten by an eager, hungry horde of bank-inhabiting, vagabond
IIpiders. Few escape j those that do proceed to burrow their tiny tunuelR
and to make their nests. Thev bite out the earth with their falces, which
a.re very strong. Of course, the state of the earth deterrnines the length
(II time taken to make the boring. The doors of the nests of young
hpiders are round, very small, being ItiKS than ~1;r in. in diameter, anll
very diffioult to detect. As the IIpider ~rows she needs to enlarge her doO!.
and the door (fig-. 10, c, the arrow). In wllny cu~c" I:!htl is found dead
btiliind thi~ screen, ,,"liile her dOOI', aftPr warping with the weather,
allows the tlntranCtl of all sorts of vtlrmill-woodlice, ctlntipedes, aptera,
small spiders, and a large number of other forroN of insect-Iif!:'. Th~e
cannot reach her, for the screen I:!huts them off.
l.Tnless there is an ahl!ence of llloisture, hungcr haM no terrors for these
spiders, for they are easily able to exist without food for three 01' fonr
uwnths. FI'equently in a famine a spider devoUl"!l her neighbour, a hard
fight alwa~·s ensuing firt.!t.
I have severa] realiOllS to suspect that Pompilius jugar, &c., is a keen
enemy of .11. distincttls. I have caught P. juga:/) dragging a trapdool'
Rpider aoro~s a bank. '\'bether the spider was caught b~· the fiy by the
latter opening the uoor, 01' by the spiller jumping out to catch the fiy and
instead oatching n Tartar, I kILOW not, but I have murc than oncc found
a pupa-caselli a small Ichneulllon flr lying a1ll0ng the remain!! of a spider.
She will !:IelUom uash out unless the fiy is right in fl'ont of the
door. If it is too far away she would be obliged to expose her body
to danger while she reopeneu the door. As it is, her abdomen keeps
the door open, so she !:loon slips back. When the spider has eaten the
fl, she drops out of the door those parts which she discal·ds .
• I kept thirty spiders in a small box placed on a shelf. On the sanny-
da,s when the flies were about I would spl'inkle a few grains of sugar
in' fl'ont of eaoh door, and put the box in the sun. The unsuspecting
flies would come to feed on the sugar, and would fall easy prey to the
spiders. In w'inter, when few flies were about, on the fine days I would
oatch hou~e-flies and tie ootton to their ,vings, and make them walk over
the door. The spiders would drag them In, ootton and all. Next da~
the cotton, with the dry oaroase of the fl~·, would be found often an inch
from the door.
Spiders may be killed whilst attacking an Iohneumon fly inadvertently.
This would explain why nests, even in a plentiful insect season, are
found tenantless, exoept for vermin.
Even when not looking for food, spiders will be caught watching out
of theil' nests. Before they emerge at night they always reconnoitre fOl·
an hour or so. If a spider is alarmed she rushes up from the terminus
of her tube and proceeds to resist an entranoe.
of silk between tIle doo\' Ilnd 1illl, in order tu keep out an illtl'uder
(fig. 10, a, near letter D). HOUletillletl the~' do thitl whiM hibel'Ilating.
If the tube pierces thc bank in a (10,,"lIwl),1'd uircction, in ordel' to pl'otect
its ab(llllUen whill! IItill hol(Un~ the liel it woul(l be obliged to hinge thl!
door on tlu~ bottolll Bi(le of the open. Consequl!ntl~' thl' lid wuultl Ill'
continually falling open "iUI it.., 0"'0 weight. Hencl' the tube pierceI'>
the bunk ill all npwal'fl dirertion (fig. 14).
H.\BIT.oI.T.
This spidel' ill seldom found lU8n~' llliletl frum the ",en-couEtt. The!
tlpidel's prefer Il sheltered bank, ne'l"er, as far Ill! I know, burrowing into
the ground, as some otller h·ll.pduol' species.
Spiders of the sallle species frulll different localitiel! ufton III10W curiom;
differences in habits and manner of building their nestR. This is taken
into account when I ila~' that the l'clmlts may be slightl~, dit!similar if
s~idcl'l:l from provinces other than .\ucklnnd 01' Wellington are t!turlied.
I have been uisappointed to find that JI. dillfillrtuH iK fl'ee hom
paraRitic Aral'ill((.
I I
,. 5
°f"_~"
7
CD OiH]) <3
abc d
ca
e C!
r. h
9
MIG.!S DISTINCTt'S.
KIRK.-Heptatl·elll.\ cirmt.t ForMa. 241
With regard to the forking of the ventral aorta, it it! p~rhapt! \\tr1thy
uf note that ill two preparations that carry much injection the right
llivision of the aorta appears as though it sprang from the left.
Dorsal Aorta.-AE. in Bdellostoma, the efferent hranchial arterie!. u()
llOt run uirect to the uursal aorta, but those of each side run to a vt!t!r.el
that may, for the sake of convenience, be called a lateral aorta. This
uccupiell' a dorso-Iateral position parallel tu the dursal aorta, to which
it sends regular communicating vessels, Theile communicating vessel~
3re three in number on each side. The most anterior of these ve8~ls II.r~
behind the second pail' of gill-sacs, In" A Treatise on Zoology" (Ray
LankeRter) Goodrich l'epresents in diagram the circulatol'Y sylltelll of
Bdellostoma, the diagram being based on the figures of Miiller, Jackloon,
and Klillckowstronl, to ,\hich figures I have no accellS. The uiagrlllu
shows that the lateral aortae of Bdellostoma have 1110re connecting
branches to the dorsal aorta than those of Heptatrema, and that ill
Bdellostoma tbel't~ is communication anteriorly to the first gill-liaCH, III
front of the gill-sacs the lateral aortae of Hepfatrema can be h IlCtt1
forward nearly to the head. Posterio1'ly they turn inwards about abrt!allT
of the last gill-sacs to join the dorsal aorta, which has, of course, receivctl
all its blood through them. The turn inwards is made at a n(ltic~nhlc
angle, and the last efferent branchial veE.sel of either side may not be
rt'Ceived until after the turn is made (see Plat~ XVII, fig. 1).
The efferent branchial al'teries leave the gill-saci on the anteriul'
aspect. Those of the anterior pail' always, and those of the secolld pail'
nearl~· always, branch just as they leave the gill-sac, the two b1'ancheR
entering the lateral aorta separately, The third, and L!lollletimes even the
fourth, efferent artery of one side, or both, may branch in like fashiun.
In the preparation figured in Plate XVII, fig, I, the second efferent
branohial artery of the left side has a distinctly double origin.
From the median dorsal aorta vessels are given off to suppl~- the
notochordal sheath, the pharynx, and the body-wall. Thesc last. tll~
segmental arteries, often alternate with each other, but posteriorl~' tu
the heart a more 01' less regular arrangement in pairs becomes evi.lent.
One segmental al'tel'~' often suffices f01' two lUyom~res.
I have not been able to observe the blood-supply of the pl'o-ll~plll OM.
With regard to the nephridial system, each segnlental Ilrtery that C)'OSllCI!
the system sends, typically, a branch to the col'l'esponcling glolllel·ullls.
In cases-ann they are frequent-in which thel'e is no segmental artery
corresponding to a gimnel'lllus, a renal ul'ter~· runll direct from thl)
aurta (Ilee Plate XVI, fig, 2). On~ Clllle was observed in which a segmental
Ilrtery sendl! branches to two glomeruli.
Posteriorly to the heart splanchnic arteries run from the nursal aortn
to the intestine; the splanchnic artel'ies are numerOUII, and appeal' to
be one in each segment.
Numerous slende1' arteries pass from the dorsal aorta to th~ gonaclM.
These arteries are much more noticeable in cases in which there are
many ova forming.
Jugular SYBtem.-There is a large right jugular vein lying above
the lingual mass and beside the phal·ynx. It arises well forwal'd, abt'ellsf
of the fifth slime-gland. The vessels that contribute to it come from
the body-wall, the lingutJ.l mass, and the pharynx. Anteriorly to the
gill-sacs it resembles 0. sinus rather than a vein. In the diagram for
Bdellostoma already referred to, the anterior part of the right cll.l'llinnl
KIRK.-Hel'tlltt·~lU..1 cinllt:l FOJ·/Jft'J·.
EXPLANA'rlON OF PLATES.
PL..I.TE XYI.
Fig. 1. DiagrammatiC' repre'4entation of th(' cir('ulatory system.
Fig. 2. Part of the dor'!8l vessels and the nephridial system, from the doreal aar,ect.
a. Rena.l veinlet running from tht· dorss.l a~pect of a glomerulus to a seg_
mental vein in the bodv-wall.
b. Segmenta.l artery t-endinp;'bnncheh to two glomeruli.
Pr.\TI!l x-nI.
Fig. 1. The efferent bran('hial vessel. ani! the anterior part of the doreal aortiC' ~~ ,tl'lI1,
from the dorsal aspect.
Fig. 2. Th(' afferent branchial system, fronl th(' '\"entral aRpOOt.
Fig. 3. Right afferent branchial vewlk from till' right side.
Fig. 4. Anterior }lIlrt of pORt-eardinal ~y91:em, ~ho"ing connection of ~ht ..,jnu' \\lth
portal heart.
Fig. 5. The juglliar "ystem, C1iR~I'Ctetl fl'om ",mtral aApeCt.
---
TnAN~. _, •.
'" Z I"~T., PUTa XVI.
TRUis. N.Z. bST., VOl,. XLIV. PLATE XVII.
Fig. 4.
Fig. 6.
HEPT.~TRElIA. CmRATA.
COTTO-:sr.-Nott8 on JreUingtM PkUBiographu. 241;
CO:S'rE::;''rR. l'AGe
Introduction .. 246
I'3tructure .. 245
Land feature.. .• 246
Cycles of er06ion .. 248
Forms of the Ka.ukau cycle .. 249
The Tongue Point cycle .• 250
The present cycle .. .• 251
'rhe Port Nicholson ares. .. .. 251
Coa~t features .. 254
The clifts 254
The coast platforms .. 255
The Wellington fault 257
The fault-scarp .. 257
Xature of the mO\"CIDent 259
Other faults ., ~60
Changet! in drainage of the Karori·Kha.ndalJah or •• Long ,. \"alley 262
'rype of topography •. .. .. .. .. 264
l'3ummary .• ~&
INTRODUCTION.
IN the pl"epal'ation of these notes a detailed examination has been made
only of the district lying to the west of Port Nicholson, which for con-
venience will be referred to as the Wellington Peninsula. By means of
hasn' traverRes and observations m.a.de from a distance, howe,\'er. it has
been possible to reach general conclusions which, the writer belieyes. holcl
trut' for the whole of the district represented in the locality-map (fig. 1).
STRUCTURE.
With the exception of a few small patches of Recent sands and gravels
occUl'l'ing a~ beaches and river-flats, the rocks are a single series of sandy
argillites and fine- and coarse-grained greywackes.t They are closely
folded in a. complex manner, but, owing chiefly to the unfossiliferous cha-
racter of the rocks, the structure has not yet been unravelled. On an~·
cross-sectioll rapid changes in the direction of dip are the rule, but the
attitude of the strata is so much more nearly vertical than horizontal that
a":l far as their effect on topography is concerned they may be regarded as
vertical. There has been no folding of any consequence in more than one
." hen this paper was written 1he writer had not seen the criticism of Fell's paper
by W. ll. Davis in the Bulletin of the Am. Geogr. Soc. (voL 43, No.3, 1911. p. 190 •
Had. he read that a\ticle ea.rlier he "'ould have been a.ble to profit by severa) nluable
hintB gh'en by Professor D8\'is.
t Thi. thick, unfossiliferous series is correlated on lithological grounds with the
llaita.i system of New Zealand geologistR. which, according to Marshall C" New Zealand,"
Ha.ndbuch reg. Geol.• 7 Band, 10 Abt., p. 35. 1911), is of Trias·Jura age. The period of
foilling is believed to be late Mesozoic.
246 'l''I'ansactionB.
LAND FEATURE~.
The adjustment of stream-coUl'l:les to struoture, an arrangement which.
with exoeptions that will be noted, has been retained by existing streams,
points to prolonged exposure to suba.erial denudation for the duration of
at least one nearly oompleted earlier cycle of erosion. The existing topo-
graphy is composite, and has been developed during an uplift of at least
800 ft., and perhaps of 1,000 ft. or more. The amount of uplift seems
to have been nearly uniform, although probably not quite uniform.
over the area studied. During the uplift pauses occurred, some of whi('h
COTTO::>l'.-Notes UI/ IJ"elllflgtoll Pk!lfllograpky. 247
to over 1,000 ft., while a few peak£> reach to 1,500 ft. and more. The
texture of dissection il:l medium to fine.
Gycles oj E,·osioll.
All the forIlll> recognized. appear to be dne to stream-action alone. Three
sets of forms l:U'e recogruzed, corresponding -to different positions of base-
level, and It is possible that forms are present corresponding to other pauses
in the movementi:l of uplift. It is evident that such pauses did occur, for
remnants of coast platforms are found, in places, one above another (see
p. 25'5). Shortness of a period of standstill during which erosive pro-
cessetl work is not in itself a reason why the period should not be dignified
11Vi.th the name of .. cycle." Huntington and Goldthwa.ite* have pointed
out the analogy between the term •. cycle ,. applied to an erosion period
and the term •. life" applied to the period of existence of an orga.nism.
C< Life in one significd.tion is the complete existence of a normal organism
duri~ which it passes from infancy, through youth, maturity, and old
age to dea.th. The life of man in this sense is seventy years. In another
sense life is merely the actual period of existence of any specific organism.
All animal whotle life in thl' first sense of the word is fifty years may die
the day that it is born, but nevertheless we say that it has finished its
lif!". A cycle ill the first sense is ideal and can never be realized, since
infinite time wonld be required to reduce any land-mass to the condition
analogous to death-that is, to a plain at absolute base-level. In the
second sense any region that is subjected to erosion during a definite
period, no matter how short, passes through a cycle and can be de-
scribed in terms of age and development." The term" chapter," proposed
by Davist for an unfinished cycle, hal> not come into general use. A
number of such brief cycles, corresponding to pauses during the earlier
pa.rt of the period of uplift in the Wellington district, may have left traces
on the topography not yet obliterated, and the number of nearly flat-
topped ridges of varying height which lie spread out to the west of
Kaukau Peak suggests that in the future, with detailed work and accu-
rate mapping, some at least o{ them may be recognized with certainty.
The writer, however, feels justified at present in grouping the observed
features as belonging j 0 only three cycles.
The earliest cycle or which a record is preserved by existiug topo-
graphic fe.:l.tures will be called the Kauk.lu cycle. Base-level stood
perhaps 800 ft. or IWO ft. higher than at present.
To the next, 01' Tongue Point, cycle belong most of the broa.d
features of the landscape as seen from a high point of view. Base-level
stood about 250 ft. higher than at pr6l:lent, a.nd dUling the cycle the
most extensive of the elevated coa.stal platforms, including that at Tongue
Point, were cut (see p. 255).
It was between the Lukau base-level and the Tongue Point base-
level that other pauses occurred. that are mentioned above. They were,
no doubt, brief, and the Tongue Point cycle itself was of relatively short
duration. It is, however, of importance on account of the very recent
date at which it was interrupted.
There is, lastly, the present cycle, with present sea-level as base-level.
* Bull Mus. Comp. Z6ol, Ba.rv., voL 42, No. 5, 1904, p. 239.
t •. Physioal Geography as a. Univensit.y Study," JOUl'D&l of Ueol., 1894, p. 63.
C01'·.rOx.-Xotes on Tr eilill{Jlolt l'Jz!lIlIO!}rapltu 24:9
FIG. 3.-Vmw all' TlDiI EAST BBuoa: all' OJUllro BTBBAlII, LOOXING NOBTHWAlID :&'BOlli
TEll SlllIIMlT OF KA.UltAU P.a..UL.
A. graded rea.oh, at ita lowest point !SO ft. above sea·level, and inoised a.bout 50 ft. below
the graded vaIley.floor of the earlier OYllle.
Little is left of the SUliace belonging to the Kaukau cycle, and -per-
haps the most extensive remnant is a tableland nearly a square mile in
extent, standing 950 ft. above sea-level, which exists on the divide west
of the Makara Stream. In fig. 2 it is marked P. It has an undulating
surface of mature valleys and rounded spurs, appearing from a distance
perfectly plane. It is bounded on all sides by the slopes of young ravines
eating into it.
Many of the higher ridges show very similar topography. though no
other is so nearly plane. From these observations it appears that during
the Kaukau cycle the stage of maturity was reached, and that this nearly
plane area, P, stood not far above bo.se-Ievel. The height of Kaukau
and other peaks above it shows that the relief remained fairly strong.
It is not probable that this cycle was the one which began when the
folded range :first rose above the sea, for planation might be expected to
On the foreground .lnd on right a.nd wft a.re benches of the flood·plain of the Tongue
Point cycle.
.' . '
..........
DIVIdes In the Tongue Pomt cycle ha.ve In some cases been reduced to
a fauly low altitude. Where they stand only 600 ft. to 800 ft. abo....e
plesent &ea-Ievel they have been rounded and their slopes graded, and
lOck outcrops are few. Higher-stamdmg ridges aze more rugged, With
rock outcrops and sharpened summits, except where they are flat-topped,
a.nd forms of the Kaukau cycle remain.
The broad upland features in fig. 3 belong to the Tongue POInt cycle.
The Ohariu and other streams, howevel. shown in the figw.e have been
!evived, and reaches have been graded and WIdened WIth the fOimatlon
of flood-plains. These belong to the present cycle.
* Trans. N.Z. Inst., vol 4J. 1910. map and aectlOJl, pp. 337, 339.
252 Transactions.
• See F. P. Gulliver. ··Shore·line Topography," Proc. Am. Ao. of .Arts a.nd Sol,
vol. 3!J,. No.8, 1899, p. 189.
t .. The Outline of Cape Cod," Proo. Am. Ao. of Arts and SoL, 1896; reprinted in
Geogr. Essa.ys, 19()9. p. 090.
t .. G&lgra.pbie physique," p. 685; Pa.ris, 1909.
COTTOY.-.Y otel 011 II" rlll11!lfoll Ph/lllio/lJ"aph/l. 253
In the lower diagram Eva.ns Bay (opening to Port NIcholson) is on the left. and Lyall'
Bay (opening to the ocea.n) on the right. Spurs running down both to E\""a.ns Bay
and to the ocean have been cut back by marine erosion, and rock platforms indi-
cating their former area. have been exposed by a recent movement of elevation..
These are much more extensive at the seaward end, but even on the shore of
Evans Bay a moderate amount of cutting bas been done by the waves raised on the
waters of Port Nicholson by the prevailing north wind. The sand-bar joining the
island. to the mainland must have been formed at an early stage, for the spurs
running down into it have been almost completely protected from marine erosion.
Tne uppsr diagram is a restoration of the initial form of lIira.mar " island."
The case of Miramar Peninsula. is therefore one where island-tying has been
assisted by a slight movement of the lanl1. * It seems probable that with-
out a slight movement of elevation a shallow channel would ahyays have
ooen kept open through the ba.r by the tide.
In a. quaint paper by Crawford, t entitled "Port Nicholson, an
Ancient Fresh-water Lake," the view was advanced that the present
entrance had been opened quite recently by the sea, and that over a
dam of boulders in the E\"&ns Bay-Lyall Bay channel the waters of a
fresh-water lake formerly escaped and cascaded down to join the "great
Cook Strait liver."
The small cha.nnel appears to haye been formed by the drowning of
two small soeama, one flowing north and the other south. separated by
a low divide which is e,idently not deeply buried, for the spurs running
down from opposite sides into the sand-bar are not widely separated.
From the above description and from fig. 6 it will be gathert'd that
the coast-line of the downthl'Own Port Nicholson block is a normal
drowned coast, pasaing thl'ough the normal cycle of littoral erosion
which has reached the early mature stage. It is thus in strong con-
trast with the coast~ of the neighbouring high-standing blocks descrIbed
in the next paragraph.
(.'OAf,T FEATURES.
TIll' CliD8.
The actual outline of the coast of the high-standing block IS the
result of marine erosion working back from an earlier coast-line almost
certainly bounded by fractures. This seems to be the only view tenable,
for the amount of marine erosion Dl"CesSS1'Y to clit back the present
coast, with its line of lofty cliffs, from a coast-line of any other form
would be enormous, and seemb out of the question when a comparillon
is made with the recently revived conditIon of the land-drainage. There
is no evidence of a slow &inking of the land such as would be l'6quired
to keep up the activity of wave-action on a receding coast. The
dt'pths of hundreds of fathoms recorded within a very few miles of the
sonthenl COllst indicn.tc that the block to the south has sunk, and the
closeneSll of the hundred-fathoms line to the Webtern coast indicates sub-
14idence in that dilectioll.
The hypothesis of a fracture-bound coast gains further support from
the relation of the coast-line to stream-courses on the land-surfare.
The SlIver Stream (see fig. 2) rises at a height of 1,000 ft. only three-
quarters of a mile from the southern coast, and flows northward. The
western coast also cuts in along a north-easterly line, making an angle
with both the strike of the rocks and the stream-courses. The Ohariu
Stream, on the nOIth-west, like the Silver Stream on the south, rises
almost OD the coast, and :flows inland.
Cook Strait, wmch bounds the Wellington Peninsula on the WeRt and
south, has beell generolily regarded as the result of faulting since the time
of Hochstetter. whol:le views were followed by Hutton and more recently
by Park. Hochstetter's earl.V view· was that one island had been thrust
laterally past the othel'-that il:l, that the movement was of the nature
of It. "flaw." As has hee11 pointed out by Suetls, however, Hochsiettel's
later viewt was that Cook Strll.it owed its orij.,rin simply to the tlubsidence-
(If a mountain block or blocks, and he was ItWal'6 that the ('ontinuatio11
ot the North Island ranges is to be Iound on the same line of strike in
the Kll.ikoura Mountains of the South Island. This relstion is brouqht
out by Marshall'st ll1apl:l of physical features of New Zealand.
The west and south coasts present simIlar ft'atures. The only pro-
jecting points are those composed of resistant rock, usua.lly bands that
are hardened with interlacing veins of quartz, filling joints. The inter-
vening, less resistant rock ba.nds recede as bays of gentle curvature,
bounded by imposing cliffs. The larger streams emerge at beach-level,
in gorges revived and steepened by the rapid recession of the coast,
* Lecture ou the Geology of the Province of Nl!b.ou, 1859, reprinted in "Geology
of New Zealand" (Auckland. 1864). p. 106; c,ee al'lO Park's" Goology of New ZeaJBnd;'
1910, \'; 262-
t • Reise dar' Novara.' " 1864. Geol. vol. 1. p. 2.
t Lac. eit., pp. 10. 11.
COTTON.-.Yotl'8 on Wellington PII·/RlograjlJlll. 255
the base-level at the time when the streams of the district developed the
;,rreater part of the existing upland topogra.phy. For that reason the writer
ha.s named that erosion cycle the Tongue Point cycle.
The height of the shelf at its inner edge at Tongue Point is 240 it.
Iti slope seaward is at first 10", but lapidly decreases, and at the
end of Tongue Point, where the shelf is half a mile broad, it runs
gently out at an angle of 2' or 3".
The upper surface of the tilielf u. oovered by a veneer, 6 ft. or 8 ft.
in thickness, of gravels similar to those of the present beach. They
vary irregularly from beds of ooarse roughly rounded gravel and
boulders, matt-rial similar to what is being supplied to-day in large
qUllntities by the smaller streams, to layers of fine flattened discs of
l'etl.ch-shingle varying from the size of a threepenny-piece to that of a
lleJlDy. A layer of the coarser gravel is seen on the right in Plate XVIII,
fig. 3.
The varying height of the outer scarp of this ma.ri.ue terrace as
seen from the sea is clearly due mainly to the '\"arying breadth of the
portions that have withstood the action of the sea, the seaward slope
of the shelf being regarded as nearly constant. At the extremity of
Tongue Point it comes down to 170 ft. Beyond the next creek: to the
\\ est, where there is a. well-preserved. but narrower remnant, the outer
edge bounded by the present scarp is, as might be expected, higher.
It is evidently this apparent "ariation in the height of the shelf that
256 Trail ~art Ion,.
led Park* to remark that he had satisfied himself .. that it was not an
uplifted maline platform of erosion." It may be remarked that a
section, even on a vertical plane through a coastal platform, parallel to
the average direction of the coast must not be expected to yield a per-
fectly horizontal crest. It ought to show a convex crest opposite to
bluffs where the old coast approaches it, and a concave outline opposite
bays, 'where the old coast recedcb. To this initial irtcgularity there may
be added slight variations in the amount of subseq~~nt upl~ft. Rem-
nants of this terrace extend nearly to Cape Tel"awhlti, and It may be
seen also at Te Kaminal'u Bay, on the western coast.
There exists also a higher shelf, which was examined at Tongue
Point. It may be seen in fig. 7. Its height is about 450 ft., and, like
the lower shelf, it is covered with a layer of water-worn pebbles. Its
width at the powt examined had been reduced by the cutting of the lower
shelf to about 50 yards.
At Barmg Head, on the coast south-eastward of Pencarrow Head,
at the entrance to Port NIc1lOlson, similar shelves occur, t and again at
Cape Turakirae. They may be seen from the deck of 11 steamer enter-
ing Port Nicholson. The &ketch, fig. 8, represents them as seen from
FIG. S.-THB ELEVATBD COAST PLATJ'OJIlIIS BBTWlIiEN PDNCAJUWW HlIIAD AND BAlIIl\G
HB.\D. AS SBEN P'BOH 'l'HB SIGNALoSTA'l'IO'N ON Ml::aAMa PENINSULA.
Pencarrow Head in oentre, Ba.ring Head OIl right.
'" Trans. N.Z. !nat., vol. 42, 1910, p. 580, a.nd fig. 3.
t See Park, Zoe. cit., p. 580, fig. 2.
:j: See MarshaJl, lat'. t'lt., p. :..1.
Von'ox. .rote, ()/I lrelllllUtlJII Pll!lb/()"1'(lpllg. 257
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TB\M. N Z I"'ST, VOL. XLIV P[ATL XX1
ut! regarded a.s hanng extended at least a nnle out mto the Wd.tels of
Port :Nicholson, enclosing between them the contmuations of the present
gorges; and the COMt must have been cut back to a straight !me by
wave-action.
The problem may be attackecl in two ways: (1.) Search for the
lOck platforms which should remain to mdicate the former exten-
!:lIOn of the spurs. A glance at fig. 5 shows tha.t these are absent,
and that the deepest water of Port :Nicholson comes close to this shore.
Rock platforms. If they eXlsted, ought to ha.e been actually r8Jt.ed
above water by the 5 ft. uplift of 1855, but for nearly the whole length
of tht- scarp rocks ale not exposed at low water more than 50 vards
from the foot of me chffs. (2.) Comparison wIth other parts of the
coa.st-hne where marine erosion has been more or lebS effective m cutting
back the coast. The coast of the seaward end of Miramar Perunsula
(fig 6) may be considered. Here, indeed. bluffs have been cut back to
the extent of a mIle, as the exposed rock platfol'Ill at theu base Fohows,
hut the coast hat. by no means been rendered perfectly straIght. :More-
u.er, compared WIth Its actIVIty on the outer coast, wave-action Within
the harbour is extremely feeble. A safe compalison can therefOle be
made only with another stretch of coast withm the harbour. When the
eastern shore is examined it lOS found that wave-action has succef'ded
only m shaving off the ends of points. Fig. 11 represents the eastern
shore as seen from the signal-statton on lIiIamar Peninsula. Its
nrel!,ular base-lme may be noted on the map!!. figs. 1 and 5 It
should be noted that this side of the harbour is bounded by a fltnke
ridge, and that no spurs of any magmtude run down from It So a shore-
line originally nearly straight has been rendered but little straighter
by wave-cutting. Moreover, the increasing height of cliffs towards the
harbour - entrance shows that the greater part of the work has been
done by waves rolling in from the open sea. The western shore of the
harbour, on the other hand, IS affected only by waves raised on the
harbour itself. The effect of waves raised within the harbour is seen
on the shore of Evans Bay (on the left in fig. 6).
The conclusion reached is that the scarp bordering the harbour on
the north-west, with its straight base-line, cutting at an angle across
the strike both of the rock strata and of the drowned ridges to the
south of it, with its faceted spurs a.nd Its steep-grade gorges, is the
result of recent faulting. Fig. 9 may be compa.red with the sketches
and photographs of the Wasatch Range given by Davis,· and also with
the diagrams illustrating his theoretical discussion of the dIssection of
the face of a faulted block.t
• w. M. Dans, Bull. :Mus. Comp. ZOoL. Han•• vol. 42. No. 3, 1904, P. 153, aLd
pl. 4: &lid voL 49, No. 2, 19O.l, fig. 2. a.nd pL I, A.
t Loc. cit., vol. 42, No. 3, 1004-, fir. 6-9.
260
From the absent"P IIf ...lIpped material alone. the base ot the Wa&atc.i.t
Range, in Ct.:l.h. DaVIt. argued that the slope of t~e spur-facets now found
there mves the Inchnatiun of the plane of faultmg. In the case of the
W~l~on scarp. however, it is uncert~in whether a scree of slipped materi.ll
exists or not beneath the water and silt of the harbour. Nor can the very
e,en slope of the l.acets throughout the length of the scarp be take~ as an
mdication that they represent the actual plane of faultIng. Thell" slope
appear@ rather to be .• the angle of rest for the products of decay" of the
material of which they are composed. The writer cannot agree ,vith Bellt
that the slope is steeper than the angle of rest. It is clear that many, if not
all, of the clearlv defined, sharp-edged facets owe their actual form to wave-
action at their bases, the extent to which the scarp has been thus cut back
b~ing indicated by a narrow wave-cut platform at its foot. This, however.
seldom reache& a width of 40 or 50 yards, and part of it may represent a
le....elled-off scree of slipped material. It is now almost entirply covered b,
the railway-embankment along the shore.
Reasrr.s have already been gi,en for believing that the actual movement
has been subsidence of the blocl,: to the south-east (p. 258). It was
assumed by Bellt that the faulting movement was one of block elevation
and tilting towards the north-west, amI the Porirua Stream was cited as an
example of a stream flowing down the tilted back slope of the block. There
is no doubt, however. that the Pomua followed Its present course before
faulting took place. It follows one of the old strike valleys. In the valley
there is evidence of recent revival. but not such a~ ,vould be required by a
tilt of the magnitude assumed: it appears to be due solely to the general
movement of uplift which has affected the Wellington Peninsula. although
perhaps not everywhere by exactly the same amount. The drowning of the
lower Porirua. may be ascribed to a letl<;-extensive later tilt of a much larger
block of country (see p. 257).
OTHER FAULTS.
An origin by faulting is implied for some of the longItudinal featureq
of the Wellington Peninsula by Bell.§ and the line of the Makara Yalley
is inclllded by lIcKay' among" active faults and earthquake rents:'
The presence of many faults, and particularly of the last mentioned, is
revealed in natural sections. The three faults which lIcKaY·1 describes
as .• converging on . . . the capital of ~ew Zealand" can be recognized,
although it is difficult to see why they are to be regarded as the continuation
of faults in the South Island. The stratigraphy of the district is too little
known to allow an estimate to be made of the amount of movement on the
fault-planes, and the period at which the main movement took place has not
been ascertained. It can he confidently stated. however, for the whole of
the area west of the Karori-Khandallah Yallev that physiographic evidence
of recent faulting is entirely lacking (see pp. 26H4-). The boundaries
of the subsided Port Nicholson block may next be investigated.
. On .t~e map of Port; ~icholson given by Bell** there are indicated.
In addibon to the Wellington fault. 6ve other fault-lines bounding the
• Bull :M. C<)mp. ZOo!.. Har,· •• vol. 42. XCI. 3. lool, po 15~.
t Loe. tit., P. 536.
: Loc. rit., 1). 539.
§ Loco cit., sectioD. p. 339•
• 1 •• R~ of Geolupical E~pl(lrJ.ti"n.... lS90-IH: !R.I.p. p. I: WeilingtoD. 1891.
Lor. til•• p. Ut
•• Lot:. cd., I}• .137.
:261
uu\Vuthrown area. It is :probable tha.t these linea are only suggebtions, fOl
')n the accompanying section giving probable faults· two given on the map
elre omitted and another is introduced. As mapped they are nearly parallel
to une another, and appear to coincide with the strike of the rocks. For
those bounding on the east and ''rest the longitudinal ridges of lfiramar
Pemnsula and the Kilbirnie ridge to the west of it there appears to be
no e,-idence. The elongation of each ridge IS satisfactorily explained as
cotresponding to rock StruCtUl'e. Xeither lidge has, on either side. a
..tldi~ht or gently cUITed base-line. but sprawling spurs are giT'en oft (see
fl.;,:. 6). Both shores of Lyall Bay (fl.g&. 1 and 6) directly facing the ocean
to the south are bounded by clifis. That th&e are not fault-scarps there IS
abunUa.nt proof in the extensive rock platform& at their Lases, which were
rollsed a.bove the sea by the &mall uplift of 1S;S5. These prove a formel
101lg tleaward extension of the spurs. Where the tombolo (fig. 6) connects
lIiramar Peninsula to the mainland this has afforded protection from
mdorine erosion, and the spurs run far out, that from lIiramar Peninsula
almost meeting that from the mainland. To the north of the tombolo
m EYans Bay, on both shores. smaller scarps are found, fronted by
less-extensive rock platforms than those of Lya.ll Bay, all evidently the
work of the waves on Port Nicholson, the energy of which is very much less
than uf those of the open sea. They are, howeyer. suiiiciently powerful.
FIG. 1l.-E\STBBl!i' 8KOB:m 011' POBT YICKOLSO~, LOOXI:liG XORTB·lIoAST ROM THE
SIG:NAL-ST....TIO:li ON llm.uu.R Plil:l,'I:NSULA..
urged by the prevailing strong northerly winds, to account for the destruc-
tion of the relatively small bulk of the spurs and slopes. the removal of
which has resulted in the present scarped shore.
Similar arguments can be used against the probability of 11 fault bounding
the harbour on the east. The shore-line is fairly straight for several mil6ll
in the entrance, but the obvious reason for this is that it is the side of a low
narrow ridge, without lateral spurs, between two straight valleys. The
shore is subject to powerful wave-action, as it is not sheltered from waves
entering the harbour-mouth, and marine erosion has been able. by the
removal of quite a moderate amount of material. to cut a continuous line
of cliffs.
Farther north, towards the head of Port Yicholaon, the land is higher,
and no longer a narrow ridge. Torrent-gullies. opening to the harbour
aq small bays. are separated by tapering spurs which run down nearly to
sea-level without change of slope. The points only of the spurs have been
truncated by wave-action, and a marked decrease in the height of wave-
cut facets call be traced northward on successive spurs. This appears to
correspond to the decreasing energy of waves. ~ along the shore, with
* 1M. t'it., p. 1.
COTTO:S.--.\"OtIl8 Oil lflllbnntoll l'IIl/~IO[l/(ll""1 :!1)3
to the west, and finds its way to the sea as a tributary of the Karorl, haVIng
thus a roundabout course eight miles in length. As indicated In fig. 2, the
capture of the Silver Stream by the Karori i& n double one, two branch
ravines of the Karori tributary having successively tapped the course of
the clilver Stream. The floor of the old Long Yalley here stands about 840ft.
above sea-level. The deepenin~ of the captured 3tream a.t the elbow of
capture is 400 ft. or 500 ft. ~orthward from thil. divide the Kaiwarra,
which here occupies the Long \"alley, descendo; somewhat rapidly in a trench
incised in an older mature valley-floor. .At the upper reservoir (U.R. in
fig. 2) it follows entrenched meanders of small radius, and a portion of the old
flood-plain on which the meanders originated remains as a bench :far ahove
the present stream and at a height of 660ft. abov~ the sta. At this point .l
mature dry valley on a level with the old flood-plain bench, evidently the
old &tream-course, swings off to the north, while the course of the Kaiwarra,
:Hov.-ing north-east, is a young gorge. The sketch. fig. 12, shows the old
valley and the young gorge of the Kai'warra.
Following the old dry valley mentioned abo\-e. we find ourselves in the
broad mature valley occupied by the settlement of Karori. It has been
invaded by the head of the Karori Stream from the south-",eRt. as well
TYPE OF TOPOGBAPlIT.
_\. consideration of the courses of streams and the elongation of ridges
uf the Welli~on district lea.ds to the conclusion that, apart from local
complications due to unequal \"'erbcal mo\"'ement, the topography of the
!tOuth-westem end of the Xorth Island mountain-chain is of the Appa-
la.chian type-namely, an old, folded range subjected for a sufficient time
to denudation to bring about longitudinal drainage by subsequent streams
adjusted to structure, not following original synclinal folds, and afterwards
elfn"'ated suffiei~ntly to allow dissection b)' reviyed streams to produce a
surface of strong relief. The analogy with the Appalachian Mountains
must not be pushed too far. For example, planation in the earliest cycle
~ to have been far from complete, and the a.bsence of transverse streams
following antecedent courses is especially noticeable. Their unfortunate
absence accounts for the difficn1t\"" of railwav-construction between Wel-
lington and the western coast. In' spite, however, of the obvious differences
the remarkable similarity of our range to the Appalachians is brought out
by a comparison with Lesley's map of Pennsylnnian topography, repeated
by de Lapparent,* or \\ith the detailed maps of smaller areas gi.en by
&.lishury and Attwoou.t It may be noted that the" great Cook Strait
river" of Crawfurd,t it if existed, must have been trans\"'erse for part of its
eQ'UlSe; but reasons ha,·e been given above for believing that Cook Strait
is Dot a cll'Owued river-valle",.
• "lB.-OU! de Gc!ograllhie physique;' 1907, p. 613.
t U.s. Gaol SurveY. Prof. Paper 60., 1908. t"IpeClIalh· pl. 5 a.nd 56.
t'l'nDs. N.Z. Inst•• vol. 7. 1873. p. 448. •
C01'TOX -.rOttll 011 Jrdlillyrol/ P11'/~IU(/mJlII". 265
SL"1>D1ARY.
The following conclUSlons have been reached;-
(1.) The south-western extremity of the Xorth Island of Xew Zealand
is probably a horet isolated by subsidence of land blocks on the west and
on the south, and possibly on the east also.
(2.) The drainage-system has been ae~eloped by normal processes during
a long period of elevation punctuated by pauses. the amount of elevation
being at least 800 ft., and probably more.
(3.) The nature of the longitudinal drainage suggest!> that adjustment
to structure was established in an earlier erosion period.
(4.) A prominent feature, Port Nicholson. ha.s been ploduced by the
subsidence of a block along lines which, with one notable exception, have
not been clearly recognized.
(5.) This exception is the line of the Wellington fault, along which fault
scarp topography is well developed.
(6.) Recent changes of drainage have had the effect of destroying,
lather than completing, previous adjustment to structure.
(7.) This is attributable to the activity of transverse ",treams on and
near to fault-scarp"'.
MOST of the oVbters conbuuled in Xe\\ ~ulancl come from the Blutt and
Stewal't Islan.i. Owing to their COlllpo.ratively large size, thei! pleasant
flavour, and moderllte price they form u much-prized addition to the
ilietary of all classes. So fllr as the writer knowli, no analyses of th~
oystertl have been published hithertu.
The samples examined were procured from a fishmongtll' in the uaunl
way, and were then probably not more than three days out of the sea.
The anah-sis was begun forthwith, care being tuken in opening the
O~tel"1I not to a.llow particles of the shell to mix with the contents. The
amount of sea-water and other fiuid obtained on opening and draining
the oysters amounted to about 3 c.c. each, 0. quantity, however, "'hich de-
pends on the time elapsing between opening and draining. As the animal
dies it undergoes rigor mortis, or some analogous change, with the result
that more fluid can be drained ofij if heated even slightly the amount
is still ooore increased. In the samples analysed the opened oysters were
immediately drained under light pressure in a cheese-cloth, then minced,
ilried, grounrl in a cofiee-mill, and preserved in powder ionn.
Jletilollil.
Glycogen was estimated in the fresh material by Ptlugel"'" ulethod-
'·,t., the weighed sample was heated with strong KOH on the boiling-
l'rall~a('tloll~,
The datn obtaine.d trom analysis of the ventral part of t1u: h.1J al e
,~6 follows:-
Sample I,-Fat, 16';; per cent,; substances soluble in wiling "liTe!
fgelatine and salts), 3'7 pel' cent,; substances insoluble in boiling utI.!
~coagulated proteins, «c.'. 2'73 pel' cent. The water percentage W8' uot
estimated, These figures lire c,\lculated on the aSIJwuptiou thaT it wa~
the same as in the other parts of the fish,
Sample ~.-24/8 g11ll, ventral portion of frostfish ga~ 4'!l1\I-4I-!I'Il\.
ether-soluble iat = 20 per cent,
CrYBtalline Sub8tam:e.-On boiling il'eBb minced frostfilJ. "ith 96 pel'
ct!nt. alcohol. and allowing the extract to cool. a fine white cl'~'lItallinll
deposit fonued, rnder the mioroscope two types of crystals. appeared tn
be present; the more numerous were balls of fine. pointed needle.. ~ligllth'
bent or twisted 110 that ther resembled puff·balls, the othen were lUuch
.maller rounded clumps of indeterminate crystalline matteI', At nl'Rt
~ight they might be mistaken ior leucin and tYI'osin, When filtered and
allowed to dry in the a.il· the deposit formed a white powder. !Jasil\"
bOluble in watel', It gave no biuret or Millon'b reaction. and did n~t
1,t!lluce Fehling'b solution. Anullonia caused a slight precipitatl!, '\'\1U:11
tlhectly tested the powder gdve liistinct evidence of carbon, lJitrll~l'n.
olUlI phosphorus,
While frostfish i, uudoubtl!dly of high nutritive value. untl Illl I:).wl·
lent article of diet, the conditions under which the nRh i~ nhtllined. it~
ItlJubtful degree of freshneNS, its hi~h percentage of fnt \I hid. frulU it~
nil: nature is apt to become rancid, the presence of Q spetiul alcollol.
KIIluble Bubstance at present of unknown nature, all tend tCi makt, line
ca1'eful in advising ita lise for invalids. Parasitic worms-ocllllIll. ''',md.
,iI'
..nd coiled likl:! II. "atrh-'lJl'illl,l--occ1l1' fai1'lY often: the~' lire plObably
Iuite harmless
III. Kl"llAnA, OR SWJ;J:T POTATO.
The kumala. 01 }Iu.ori s,\\eet potato, i.. culti'f'at~d to a considerable
t:!xtent in the XOlth Island of Xe" Zealand. It seems to lit' the same as
-ht:! sweet potato of AllleIil.la and the Pacific Islands genelltlly, but some
..liQ'ht difiel'ence6 in the composition "ere found, and these ueser\"e to be
!Jut on record. Xo complete detailed analysis was made.
Oarbohlldrnte.-StD.rch, in tht:! form of granules which preSt:nt the
ll!>unl appearance or batata-t;,tarch, constitutes the most Important of the
~"lids. On hydrolysis it ~'ielrlb a ut:!xtro-rotatol'Y reducing-bugal'.
DHtrin.-Fresh kumD.ras Wt!IC extracted first with abllolute alcohol to
I ~mm'e sugar and other RubstanCt:!s, then "ith cold water after driving
INTROD lJCTION.
MONTAN WAX is a ha.rd yellowish material which, on account of its hlgh
melting-point, is used for raising the melting-point of stearine candleH,
and, on account of its low price, has also found some use as an adulterant
of beeswax. The wax was first manufactured from the brown coal of
Saxony and Thuringia, and more recently has been prepared from Irish
Ii ·tea.
~n the manufacture of montan wax, pyropiBBite is either extracted
with light petroleum and the soluble bitumen, obtained from the extract,
then distilled in superheated steam, the distillation being repeated until
a nearly colourless product is obtained, or the brown coal is itself dis-
tilled with superheated steam.
The method of manufacture was first patented by E. von Boyen
(German patent 101873, 1st July, 1897). In the original patent E. von
Boyen'" described the wax as consisting of two well-chara.cteri.zed sub-
mances-a.n acid and an unsaturated hydrocarbon. C. Rellt assigned the
formula CSSHSS02 to the above-mentioned acid, now called "montanic
acid."
E. von Boyent adopted the formula CiqHUOS for the acid, but now
stated that the other constituent is an alcohol melting at 60°, which is
readily attacked by sulphuric and nitric acids. He regarded the original
bitumen as an ester of montanic acid which is decomposed during distilla-
tion.
K. Eisenreich§ purified montanic acid by fractional precipitation with
magnesium acetate. He adopted the same formula for the acid &8 von
Boyen and Hell. He noted that the last portions of the acid to be pre-
cipitated melted several degrees lower than the earlier :fractions, but no
attempts were made to obtain acids of lower molecular weight from these
fractions.
To the non-acid constituent of the wax, melting at 63·5°, he assigned
the formula CuHuO, and supported the formula. by an ebullioscopic
molecula.r-weight determination, but could not find any evidence that
the substance was an alcohol,1I
.Ryan and Dllion" gave the formula for monta.nic acid as OuR.sOli'
TJle non-sapolllM1Jle pOltIon they found to melt at 58-59°, and the ana.lysis
gave 2·5 per cent, of oxygen, which a~ees approximately with the formula
C'JII~o() o£ Eisenreich, They also stated that no primary or secondary
alcoholic ~roup is prebent In the substance.
The work of plevious observers may there£ole be summed up 8.6
folloW6: (a.) Three investigators have a.SSIgned to montanic acid the
formula CuHnO!, while one investigator has assigned the formula
OUH&002 to the same acid; (b.) von Boyen first stated that the non-
acid constituent was an unsaturated hydrocarbon, afterwards that it was
an alcohol. Eisenreich and also Ryan and Dillon state that the non-acid
constituent is not alcoholic in nature, but regard it as an oxygenated
compound.
In the prebent papel' it is &hown that: (1) the" monta.nic acid" occurring
in the wax ltl 111 reahty a mixture of three acids-cerotic (OuRuO:a), mon-
tanic (OUH5602)' and melissic (030HSOO!) acids; (2) the non-saponifiable
pOltion is au olefinic hydrocarbon, probably OuRss or OuRu, both of
wh1ch. bein~ olefine6, would have, of course, the same percentage com-
position.
Oerotic and melissic acids were detlcribed by Brodiet as constituents
of beeswax, but have not hitherto been found in any mineral substance.
The separation of tllese acidl! was tedious, inyolving a series of over forty
fractional precipitations by magnesium acetate, wh61'eby the cerotic and
melissic acids Wel'6 obtained in a state of purity.
Montanic aciel, the Il.cid of intermediate molecular weight, was isolated
by conversion of the crude acid into its ethyl salt, and subsequent distillation
under reduced pressure. Some :fifteen fractionations were needed before
the substance could be considered pure.
After purification, the cerotic and montanic acids both crystallized in
pearly scales. Hitherto cerotic and montanic acids have been described
all crystallizing in needles; and the crude acids certainly do so, but the pure
acids crystallize in scales, and in so doing resemble all the lower members
of the higher fatty acids which have been obtained in a state of purity.
Tht're can he little doubt tha.t cerotic, montanic, and melissic acids belong
to thE' homologous series of the highel' fatty acids, and that these acids are
all normal fatty acids. A comparison of the physical properties of a number
of their derivatives supports this (Tables I-III, p. 285). In the case of
montanic acid it has been possible to show that the substance is undoubtedly
nOI'llUlI heptacosane carboxylic acid.
The OCCUlTence of Dlontan wax as the principal product of steam distil-
lation of bituminous coal is of great interest. Kra.m.er and Spilkert have
showll that fats and waxes, if distilled under pressure, yield mixtures of
hvdrocarbollB not unlike many natural petroleums, and they have suggested
that some petroleulnB at least owe their origin to the decomposition of wax
derived from algae.
At first it appears difficult to imagine such supplies of wax in nature
&8. by decomposition, would give rise to the immense quantities of oil present
in the large oilfields. Brown coal is, however, even mOl'e widely distributed
EXPERIMENTAl...
PART J.--THE COlllPOSITION OF MONTAN WAX.
A. TI'II!: .\CID (·O~S'l'lTUI<JNT;;.
• 'I'he fact that the proportion of hyUroca.rbon in oommorcia.l monto.n wax Us much
lesa than that of the free acids is Dot ~ for thl' physical properties of the hydro-
carbon are such ILl to Ioa.d to loss dunng the commercial process of recrystallization
from benzene.
This monta.n wax was obtained from Sohli.emn.nn and Co., Hamburg and London.
\ JOlU'Il. Soo. Chem. Ind•• 1909, P. 991.
Sci. Proo. Roy. Dub. Soc., voL 12, 1009.
RIGG.-Montall WtI:L. 273
) 9
860 820
I Mlxed III
l
sa~-89'
! j
I sa°-89° 86'5°-87°
llixed (11) Mixed (/1
k
88'lio 88"-89'
MELISbIO AOID
I
711 n 0
87"-88° 87"-88° 86°-87"
I 1IIxed (1111 I
1---1 ,-
t 110 fJ ., &
88°-89° 87'50 87'5°-8~'lio 86'50 -87'5° 85'5°-86'5°
:MIxed (k) Miud (t·) Mixed (tnl
MBLIBBIO AOID
7~78'6°, and were absolutely identical with th(' cerotic acid of beeswa.x.
This identity was proved. as follows :-
(a.) Both acids had the same melting-point, and a mixture of the two
acids melted within 0·1)° of the original acid.
(b.) The ketone prepared from each, by means of the catalytic action
of metallic iron, melted at 92·5-93°.
(c.) The ethyl cstel'R of each acid had the same boiling-point, 280°,
at 14 mm.
(d.) Both acids crystallized in pearly scales.
enlds Ethyl Montaflilte (Al61Rng-polllt. 66"). All distilllloticlns done under a. pressure
of 14-16 mm.
A B C
98&'-_ 196'-310 811)-
I I I
t1
--,:.-----ro- --~J I
~ 1
I
t1 bl h' rl
-1l98" Jlbted (1t.1J -81tr U1udlr) -8lO" -I!OO' BeeI4ue
_L Mhec1(&I) 1IW'd(;o'I)
-J, - ,j4;
~
--T--,
-SOU Mlnd("" t~ c~ ell
L----I
-1I9'l
J.h~ecl(&I)
-811t' IoiI"l:ed(Cl -810" -1!80' Beeldae
I Mu.ed("'"
.it aT ~ de Alo
- _ IIII>.ed(a') -!190' -8O!F Idhed (bII)
,;..
llfJred(ab) bT !i& ;7 ~ it!
-800 -81l¥' MIWlrJl -810- -8!Z3' BesulIl8
)i[iJed(a l ) I NlSedlbU) I
1>10
-900>
Jl b11
I
-811lP KWdlcll)
I
1..&
11'(0
1I1rad(a~J
JI
:19~ 161
I
CfI CO
ReaIcl1l88
,IeltlUlHlOlnt. N'eWngopomt, otS'l-6'1Il" J!I'I1l!1'!L 1ll~
8OG~81'6 B'1'II1'IoJIION'l'41i14'1'J1: lIBJ:.tllll4T11
'111'10 0II11OT4T1l
276
-1-- °
A.6 I -21l2 /15 mm. 60·5-61·5' !7IH-i!l'7° I 11
I
385 CJ6 IIao Of
A.9 I -302°/15 rum. 61-62° 81-82° 27 413 C27 H H 0 2
Bll -312°/15 nun. 66·5-67·5 83°
Q
32 427 ('~AH5602
08
09
I
-323°/15 mm.
Residues ..
16
12
liquor. To remove the la.st traces of acid me substance WI:I.S melted. a.nd
stirred into powdered quicklime. which was then slaked by means of a.
solution of potash. The po~ous mass thus obtained was extracted with
hot motor spirit which on evaporation deposited a crystalline substance
which aftm l'epeated crystallization melted at 59-60 and waq unchanged U
by further crystallization.
On analysis, this substance gave the following values: 0·1492 grams
gllve 0·4713 gramo; CO 2 and 0·1882 grams H 2 0
Calc. for c..:a.Ol' Oalo. for Ct,H56' Found.
C = 85·71 C = 85·26 C = 86·14
H = 14:·29 H = 14·74 H = 14·01
The analysis shows that the substance is probably an olatiw(' hydro-
carbon. This was confirmed by its behaviour with bromine water, which
was rapidly deoolourized when warmed with it. A rough determination
of the bromine absorbed was as {ollows: 0·25 grams hydrocarbon absol'bed
o073 gram~ blomine.
Calo. for C,28HsaBr2' Found.
Br = 29·0 per cent. Br = 22·6 per cent.
NOTJII.-BromiDa.tion was probably not complete. the rea.ction being only allowed to
prooeed for about three hours.
The molecular weight of the hydrocarbon as determined by the ebul1io
scopic method pointed to a hydrocarbon of molecular weight 380.
0·64 grams hydrocarbon raised the boiling-point of 8·5 C.c. of anhvdrom·
freshly dtstilled benzene 0.65°. .
Molecular weight = 380. Calc. tor CnHI56 = 392.
'fhe melting-point (59-60°), the analysis, and the molecular-weight deter-
mination all point to a hydrocarbon of the formula CnHu or Cu H 16 , but
It is only by the preparation and analysis of the carefully purifi~ dibrom
addition product that we shall asceltain whethel' the sub.:ltance contains
26, 27, or 28 atoms of carbon.
PART 11.-THE ACIDS 011' MONTA~ WAX, AND SOME COMPOU:sn& DERIVED
FROM THEM.
Since the ultimate aim of this research is to show the connection whioh
exists between cerotic, montanic, and melissic acids, it follows that the
physical constants of these acids, their melting-points, their molecular
weights, and the properties and physical constants of their compounds
should be aoourately determined.
The a.ccurate correlation of such data afford<;l :11.0 small interest to the
chemist, as has been pointed out by Krafft,'" Franchim.ont,t and more
recently by P. W. Robertson ("The Melting-points of the Allilides, P.
Toluidides, and N&fhthalides of the Normal Fatty Acids ").,t Further-
more, this series 0 fatty acids and their derivatives presents a group
unsurpassed in the whole of organio chemistry for illustrating the principle
of homology, and therefore it is desirable tha.t the physical constants of
all the memberd of the seried from the highest to the lowest should be
accurately determined and correlated. MOleover, it is desirable that the
proof of the constitution of the higher members of the series should be
demonstrated with M absolute rigidity all has been th(' case with the lowel
acids from acetic to arachidic.
A. CEROTIC ACID .\ND ITS DERIVATIVES.
Since cerotiC acid may be obtained from montan wax: only after a very
tedious and laborious PIOCesS, and since the cerotic acid of beeswax has
been shown to be identical with that prepared from montan wax, bees-
wax was therefore used for the pleparation of cerotic acid in large quantity.
The beeswax emploYEd for the isolation of cerotic acid was New
Zealand unbleached wax, which was obtained from a business firm
dealing in large quantities of the natural product. and was guaranteed
by them to be pure unadulterated New Zealand beeswax.
Th~ beeswax wa~ examined by Hiibl'!> method, which consists in
determinations - (1) the free-acid value, (2) the saponification value,
and the determination of th(' ratio of these two values.
The results are given in milligrams of caustic potash for 1 gram of
beeswax. In each determination a blank experiment, using exactly the
same quantities of alkali and alcohol, was performed simultaneously with
that on the beeswax.
The follOWIng are the results of aIWysis :-
Th°t.e :figures indicate that the beeswax nsed had rather a: low saponifica-
tion value, thus infelring the existence of much cerotin in the beeswax.
This supposition was strengthened by the fact that on potash-lime fusion'"
of beeswax, and subsequent isolation and crystalli.zation of the acids pro-
duced, a produot was obtained melting near the temperatur<' required for
cerotic acid
Cerotic acid was prepared from this beeswax by Brodie's method-
namely. extraoting quantities of beeswax with successive volumes of ethyl
alcohol until the £ree-acid value for 20 C.c. of the last extraction had been
reduced to 2·90c.c. ~ KOH. Four extractions wtre necessary to do thi'!.
The free acid. was precipitated by adding alcohohc lead aCtltate to the
boiling solution. The insoluble lead lIalts were :filtered off, and then boiled
out repeatedly with alcohol to remove impurities.
The purified lead salts were then decomposed by glacial acetJ.c aCld,
and the free acid, after washmg with water, was extracted with boiling
methyl alcohol in which, according to Marie, * melissic acid is insoluble.
The solution thus obtained was :filtered while hot, and the cerotic acid.
which separated on cooling. wag then found to melt at 75.3°.
The acid was purified by recrystallization from ethyl alcohol, acetic
acid, and motor spirit. An acid was thus obtained meltingt at 77'0°
(Brodie gives 78° for cerotic acid).
Attempts to improve the process of obtaining cerotIc acid by complete
saponification of the beeswax with alcoholic potash, followed by the precipi-
tation of the acids with alcoholic calcium· chlorIde, thus retaining the non-
acid substances in r:olution, were fruitless. It wa~ found that the calcium
salts in pre~ence of a saturated solution of high-molecular-weight alcohoL.
were easily soluble, a.nd that the precipitated calcium salts &lwa.y~ contained
a considerable quantity of organic impurities. Attempts with the lead salts,
using the same method, were also unsuccessful.
Cerotic acid has hitherto been stated to crydtallize in microscopic
needles.:!: Although this is the case when prepared by Brodie's method.
yet a careful fractionation of the ellter of the acid obtained by the above
method gives on hydrolysis a pure acid which crystallizes in pearly plates
from acetic acid.
A titration of the cerotic acid purified by fractionation of the ester
gave a molecular weight of 392'7, corresponding to the formula§ C 2C1 H u O"
thus confirming the formula of Lewkowitl:lchll and Henriques.-'
The anilide u. a white solid, fairly soluble in both alcohol and acetie
acid, but more so in motor spirit, from which, however, it does not
cr:vstalIize well .
. The crystals from alcohol were large groupK of fine needles, joined
together in tree-like formation. The yield of the anilido wa~ 55 per cent.
of the theoretical.
On one occasion a sample of anilide crystallized from ethyl alcohol
commenced to melt at 98·5°, but did not melt completely until a tempera-
ture of llSo was reached. After resolid.ifi.cation the sample melted sharply
at the lower temperature; and the sample when Cl'YstalIized from motor
spirit also melted sharply at the lower temperature. There can be little
doubt, therefore, that this anilide is dimorphous. The only other instance
I can find of an anilide exhibiting dimorphism if'! that of acetanilide. *
AflQZyaiB 0/ Owotamlide.
('al. for CssH:1I0.0aHsNH. Found.
N = 2·97 2·66
C '" 81·52 81-56
II = 12·10 12·32
Oel·owlle.-Two previous experimenters have worked upon the ketone
of cerotic acidt: BrUckner, by distilling the Jead salt of cerotic acid,
obtained a ketone melting at 62°: Nafzger, by the distillatioJl of cerotic
acid, obtained a ketQn~ melting at 92°.
By applying the recently patented method of T. H. Easter.6.eld and
C. M. Taylor~-namely, the heating of fatty acids with metallic iron,
whereby stearic acid yields 80 per cent. of stearone-the ketone of cerotic
acid was easily obtained. The details of the preparation are as follows:
9 grams of cerotic acid were heated for four hours with 0·69 grams of iron
filings in an air bath slowly raised to a temperature of 340° to ~moo.
Carbon dioxide was evolved when the temperature had reached 280°.
The temperature was now slowly raised until 3400 was leached, and the
ail' bath was then regulated and maintained at this temperature for four
hours.
The ketone thUl! obtained was purified by t.he following procedure:
I on was lemoved by boiling the ketone with dilute hyd:rochloric acid.
Free fatty acid was then rpmoved. by boiling with dilute caustic soda. The
soap thus formed was soluble in warm water, and was thus easily separated
_from the insoluble ketone. The ketone was now crystallized from motor
spirit. and a pure product was obtained, which had a melting-point
93°C.
The melting-point was not changed by furthel' ctystallization from
motor spirit or acetic acid. A 55-per-cent. yield was obtained by this
method of preparation.
The ketone thus obtained is a white solid, fairly soluble in motor spirit
and ethyl acetate, but spa.ringly soluble in acetic acid, from which it
crystallizes in feathery flocculent masses. It is almost insoluble in hot
alcohol, a saturated solution only becoming turbid on cooliug.
AlialYdiB 0/ Oerof,oIlP.
Ua.lc.. tnl' 0:11 H1020. Found.
C = 83·83 83·50
H = 13·97 13·US
Cerotone O:vi1l!e.-This compound itl neVi. It waR plepared a" followl!:
1 gram of ketone was dissolved in 150 C.c. of amyl alcohol, and one and
a half times the theoretical quantity of hydroxylamine hydrochloride,
with an excess of caustic potash to decompose the hydrochloride. was
added, and the alcohol then boiled under a reflux condense.r fOl eight
hours. It was found that unless prolonged boiling took place a poor ~;eld
of oxime was obtained. The hot solution was then filtered from the in-
organic salts, and the filtraie allowed to crystallize. The crYl!talb ,vere
filtered off, and recrystallized from ethyl acetate.
The oxime thus obtained melted at 77°, and the melting-point \\"a~
not changed by further crystallization. The oxime is easily soluble in
hot ethyl acetate and amyl alcohol. but is only sparingly soluble in hot
alcohol. The crystals from thE' ethyl-acetate or motor-spirit crystalliza-
tions are groups of radiating needles.
AnalYBi8 of Oerotone O:x;.,ne.
Ua.lc. for OSlH103N.O. ~'ound.
N = 1·87 1-58
C = 82·14: R2·15
H'= 13·82 ]S'1)3
Hervpentecolltanfl* 26 OZ.
This secondary alcohol is new, and was obtained by reducilljl; the ketone,
dissolved in amyl alcohol, with metallic sodium. 0·3 grams of ('..erotone
were dissolved in 150 c.c. amyl alcohol and boiled under a reflux con-
denser, while, at intervals, sm.all pieces of sodium. of total weight 2 grams,
were added over a period of five hours. The solution thus obtained was
shaken out with water in a separating-funnel. The solid ,vas filtered off
and crystallized from ethyl acetate. The melting-point of fhe alcohol
thus obtained was 97°, and was lIDchanged by further crystallization.
Henp61lteocmtyl ~cetate.-This compound watl obtained from the abo"e-
mentioned secondary alcohol by boiling it with It la.rge excess of acetic
a.nhvdride under a reflux condenser for six hours.
The alcohol gradually dissolved in the acetic auhydride, indicatiJl!l that
acetylation was taking place. The solution was filtered while hot. and
the filtrate, on cooling, deposited the acetate as a white solid. This was
recrystallized :£rom acetic anhydride, and after drying over caustic potaldl
in a vacuous desiccator, melted at 60.5-61.5°. The melting. point was un-
changed by further crystallization.
A'1UIlgBi8 01 He'fll/)enJerontul Acetate.
Oalo. for OsaH10oO:!- Found.
o 0:= 82·17 81·89
H = 13·69 lS'57
i
• Journ. {'hem. Soc., 1893, P. 466.
DaterminatioDa by the author.
Private communica.tioD, T. R. Eaaterfield and C. M. Taylor.
•
Previous experimenters ba.ve reported montani.c a.oid as oryst:aJlizing in -'led.
HIGG.-.llolZtan W' aa:. 283
Kelt.
Pt. Aoid. Anilirle.! Ketone. I lIDO.
_Ox- I
I
AmidE'. Ethyl
E'It(·r. IIMethyl EBtt'!'.
---------,--- ----- -------
I I
*780 Cerotic.. *98'0° "930 •• *77"
I
109° (Marie) *58·5-5li".. 6()0 (Marie)
"S3° Montanio 1*101'50 1*97'50 . . !.82;;0 .1110 .. *670 .. *67-67'1)"
-88'5°1 MeHssio.. .103" ,-99'5-10(,0 1.840 1160 (Marie) 7So (llar'e) 74'5° (Marie)
The first step (a) took place without difficulty; in step (b) the It hydroxy
acid was readily obtained, but all attempts to prepare the pUJ'e unsaturated
acid were unsuccessful, and step (0) could not therefore be attempted.
Le Seur. * in the degradation or stearic acid to palmitic, also experienced
considerable difficulty in the isolation of pure unsaturated acid (.1"" oleic
acid). He only obtained a 10-pel'-cent. yield.
The following is a brief description of the compounds isolated and the
e."tperiments maul:' in connection with the degradation of montanic acid
to cerotic :-
Bt'omlilontanic Aeid.-Hell and Sadomsky's methodt was used £01' the
preparation of this compound. 6 grams of ethyl montanate were ground
with 0·19 grams of dried red phosphollls, which had been previously freed
from phosphorous acid by repeated washing with water. Anhydrous
bromine was now added drop by drop to the mixture contained in a :flask.
There was no violent action such as Le Seur records in the case of the
fOl'lIl8tion I)f bl'Olnsteal'ic acid. The contents of the :flask were then
wmmed on the water bath for two hours under a reflux condense!'. The
condenser was now removed. and the excess of bromine allowed to escape.
The molten mass thus obtained was poured into cold water. The crude •
brommontamc acid was melted twice in fresh water to decompose any
acid bromide.
The crude brommontamc acid was then crystallized from acetic acid
and motor spirit. By this means a pure compound was obtained, which
melted at iFio C. The melting-point did not change on further crystal-
lization. ..
Brommontanic acid crystallizes from acetic a.cid in colourless hexagonal
plates. It is ea.sily soluble in acetic acid and motor spirit; the yield
obtained after two crystallizations was 60 per cent. of the theoretical.
CaJo. for C28HBBBr02• Found.
Br. = 15·90 15·81
Attempts to remove hydrobromic acid from brommontanic acid by
means of pyridine, quinoline, and a concentrated solution of caustic potash
did not result in the production of the unsaturated acid, as had been
expected.
The hydroxy acid could easily be obtained, mi.l:ed with the unsaturated
acid, by t.he action of SO-per-cent. alcoholic potash on brommontanic a.cid,
but all attempts to remove water from it by means of ortho-phosphoric
acid which had previously been heated to 200° were unsuccessful.
In conclusion, the author wishes to thank Professor EasterfieId for
suggesting this subject for research, and also for much practical advioe.
without which the writer could not have undertaken this investigAtion.
llllU,ht have dropped into the sample while it was being dt-ied, there bein~
.tlway& c. certaIn amount of copper and brass dust in mo&t laboratories.
For the work a room was therefolt:, het aside in which copper utensill:!,
11fdSS bWlSen!., und all apparatus cuntaining copper were rigidly excluded,
and .1.11Y b1'a&s :fittin~!> WE're coatefl with varnish. Further I:lpecimens were
uutu.med and analy&ed, great ral'C being tllken to ~ual'd agaimt aclventi-
tlOU!. entry of COpPl'l iutf) th(' l1e.say. The results of analysis of these
"'peclmen", d.l'C &liven III Tablet! 3 and 4. They &how that there is alway!:!
an exc&.i of copper (compared \\ith the amollnts found by the authoritiefil
quoted below) in the li\'ere. of e.heep sufferin~ from bush sickness, although
when thet!o liver!> aro extremely tatty the copper it! diminished; but if
calculated on the dried :f.lt-free liver the percenta~e is always excessive.
Analyses of the livers of healthy &heep killed for cOIlbumption have
!>llown, however, that a comparatiyoly large amount of copper in the
h,'!"r is. qwtc consistent \'lith health. llurtheJ', on do~ sheep and cattle
\, ith copper-acetate for over a year their livers were found to take U}J
much larger amountb of copper than any liver hitherto examineu. with-
out producing any of the final symptomt! of bll!1h sickness. Feeding
expel'lments conuuc,teu by Mr. H. A. Reid, F.R-C.V.S., at Wallace'\ille
Laborlltor:v, in which coppel'-acctdte ill small doses was given with the
food to &heep, showed that the liver could absorb lar~e a.moWlts of copper
anu remain healthy. The greater part ot the copper was no doubt elimi-
nated in the fdeces. 130me of the &heep died of a braxy-like disease, but
OthE'l'B remll.ineu healthy, and the experiments were dlscontinued after
they had been going on for seventeen montht:o. Expel'imentl:l with calves
and rabbits extendinA over a similar peJiod yielded negatin l'Csults, which
1
bile, pal'1:ly by the Ulin~ but, in lesser degree; .that the liVCl' retains the
copper with great tenaCIty and the pa.ncrea.s WIth wnost equal strength,
.and that the kidneys do not retain it as much as the other two O1'~ans.
The ner\"ous and muscular SYbtems do not contain enough to interfere wiLh
their action. Thev recommend numerous small dOHes in order to obtain
chronic effects. .My e:lo..-pel'iments show that the pun!.'l'eas doat! not contain
much copper. Max KleDlptner (Chum. Centr., 18!}4, il, 620; ab ..t. in J.C.R.,
1895, p. 821) records the following symptOIlll\ in poisoninl\ by sodium C'uplic
tartrate: Atrophy both in acute and chronic caset!; dlarrhoea Bome time
before death, and persistent vomiting, in chronic cases; enfeebled pulse
and laboured Iespiration. Wben subcutaneously injected into the plem a
the salt produces weakness and stiffness of the hinder extremities; the
same is noticed when the salt or copper haemoglobin is exhibited. In
one case blood and albumen were found in the urine. Copper was not
found in the blood serum, but in the colpuscles.
The question whether chronic COPPEll poisoning can occur in animals
is evidently doubted by m:my authorities. ~\.. Koldewo.y (Chern. Centr.,
1896, ii, 1041; abst. in J.C.S., 1898, p. 3H) states that no noteworthy evil
l"esults follow SID1l11 doset.! of copper (or even large doseR) in people in good
health, 01' in arumals that vomit rea.dily; long continuance ill the use of
copper, howev'!r, produces slil\ht degenerati\'e chan~es in the liver and
kidneys, which can only be detected on microscopic examination. He
doubts the emtence of chronic copper poisoning amongst workers in that
metal, any illness occurring probably being clue to otllel m'!tellic impurities
of the copper. L. Lewin (Deusch }Ied. Wochenschr., 1900,26,689; abet.
in J.B.C.I., 19, 1900, 1183) could not discover any phenomena indicating
chronic poisoning in copper-workers. The editor of Taylor's ,. Medical
Jurisprudence" (1905, p. 476) is evidently doubtful as to whether chronic
copper poisoning can occur in human subjects. This doubt should be
even greater as to whether the poisoning can occur in herbivora.
Dieulafait (Compt. Rend., 89, 453: abst. in J.C.S., 1879, p. 1020;
1880, p. 489) states that copper OCCUlS in all plants that live on prinlary
rocks, or on soils derived from thORe rocks. One hundred grams of rock
will always give 80 reaction for copper. Dupre (Analyst, ii, 1; abst. in
J.C.S., 187i, p. 511) states that copper is found in all vegetables atS well
as in animals, but rarely amounts to more than 1 ~ram per 100,000 (0·001
per cent.). DieuIa.fait states that 1 gram of ash from all plants growing
on primordial rocks give the copper-reaction, but plants grown on purl"
limestone contain but tra.ces of copper, requiring at least 100 grams of ash
for its detection. J. B. Harrison, in a recent report (1906) to the Science
and Agricultural Department of British Guiana, shows its occunencfl in
hundredths of 1 per cent. in many igneous rocks of that colony, and Hille-
brand (" Analysis of Silicate and Carbonate Rocks," Washington) con-
siders that it can be found almost invaris bly if looked for in the rock-
analysis carried out by the United States Survey Labolatozy, but it is
seldom reported unless extra precalltiOns have been taken to prevent its
entry into the analysis.
The few determinations which have been made of copper in the soil
and grass-ash of the affected country go to show that it exists in amounts
of the order of thousandths of 1 per cent, la.ther than of hundredths.
I have not been able to consult any of the original papers, owing to
the diflioulty in obtaining scientific literature in New Zealand. One is
unable to oonjecture how many specimens were analysed to supply the
above data. and whether the figures might be taken 80S 80 standard.
.At;TON.--(Jheml~tr!l of BlIIllz Sirkllel8. 291
'].' \BLE I.
Watt·s Wltb I
lloodo\l Ml!l!1-2.
vol.ll,
1•• 1148•
I
Wdtt'h I ".
Dlcholl.ll). Rlpo hrl'd, I o}o.....,
!t'.
(h'l:;':J:?,u!flW 110m
DlrtlClllPlv. DlCtlunnn I ('Iluntl~.
.U...·...... Tp Puk,'.
I I I
Pot&Sbiu oxide (KaO) 29·52 33·06 I
22·13 I 1;)·01 1645
Calcic oxide (0:..0)
lIagnesic oxide (MgO) ••
Ferric oxide (Fe 2 0 3) ••
5·82
2·22
0·59
I 8 14
3·47
0 23
9·13
249
0·62
I ;;·23
3·36
0·93
800
4·07
0·72
Sulphuric onhydride (S03) 3·52 I 3 9(1 .,1,·23 3·44 2·4.)
Silicic oxide (SiO a) .. 26·65 32·18 34-11 42·86 36·92
Carbon-dioxide (CO,) 2 01) 2·88 II 1-15 I 2·93 6·57
Phosphoric anhydride (P,03) 8·60 6H 5·56 3 83 3·79
Pot&llsic chloride (KOI) .. 17·86 4·87 117'40 IHl4 1619
Sodie chloride (:N'aCl) •• 3·09 4·76 3·14 4·10 407
Manga.nllde-oxide (1\In a0 4 ) 0·97
Carbon and undetermined 5·90
Alumina __ 1___ . 0':18
TULE 2,
I Curotld
Hrnlthy UDhoaJthy
Artery II Carotid Artely
of steer, of Steel,
W&lwetu, 119. F504. 1
.Ash-
Total ash .,
- -: 4-20 - -~.43 --I
IllIIO].uble in water 12·00 12·71
Soltlble 88·00 8729
Silk'a 106 2·77 loI):I
Sodic chloride ;)2·07 1it1·35 32·68
Phoflphoric anhyrlride .. 4·75 305 1071
Calcic oxide .• .. L'Ol 1·68 0·9.,1,
Ya.gnesic oxide 0-49 082 0·46
Ferric oxide ,. 7·64 3·80 :H2
Sulphuric anhydrille 6·13 6·90 3·86
Potassic oxide 6·55 3·99 2·23
Sodie o-dde .. 11·49 11·20 6·28
Carbon-dioxide .• .. •. 8·81 7,39 4·24
Perc=~of ash contOtituents in the dried
Silica •• 0·044 0·205 O·Hi)
Sodie chloride .. 2·168 4·322 2·416
Phosphorio anhydride •• 0·200 0·220 0·123
Calciil oxide .• .. 0·042 0·124 0·065
l!fagneaio oxide 0-0-20 0·060 0·034
Ferric oxide .. 0·318 0·281 0·157
Sulphtaic anhydride 0-255 0·510 0·2813
Potaasio oxide 0·288 0-298 0·167
Sadie oxide 0·4!78 0·833 0·467
-- ---------------- ---'-------
AF.TON.-Cli.end!t1·!1 of BtlRh Sic/meR8. 293
294 TIOT/6actwnB.
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At-oTON.-C!zemIBtTl/ of BZlsll Sicknes!. 295
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298 Transaction8.
TA.BLJ!l 6.
Boils j1iag
NOTE.-Ana.lyses made on the &a.mp1es dried at 100· C. No. LIl21 is from open
fern country; Nos. L1l22 IIoIld Lll23 are from dected fo:rest country whioh bas bel.'l1
oleared.
REFERENCES.
Ross, D. 4th Annual Report, Department of Agriculture, 1896, p. 3.
Pa.rk, A., M.R.C.V.S. 5th Annual Report, Department of Agriculture,
1897, p. 68.
Gilruth, J. A., M.R.C.V.S. 6th Annual Report, Depa.rtment of Agriculture,
1898, p. 70.
Park, A., M.R.C.V.S. 6th Annual Report, Department of Agriculture,
1898, p. 88.
Gilmth, J. A., M.R.C.V.S. 8th Annual Report, Depa.rtment of Agriculture,
1900, p. 186.
- - 9th Annual Report, Department of Agriculture, 1901, p. 66.
Aston, B. O. Hth Annual Repolt, Department of Agriculture, l!W6, p. 106.
Reakes, C. J., D.Sc. 18th Annual RepOlt, Department of Agriculture,
1910, p. 33.
Department of Agticulture Journal, vol. I, No.6, p. 457, 15th November.
1910.
Aston, B. O. Department of Agriculture J oumal, vol. 3, No.5, p. 394..
15th November, 1911.
F.ATHEn~.-CompoBition of ;Vitric Arid. 299
By H. T. M. FATHERFl.
Communicated by ProtcFosor EJ.r.terfield.
LReacZ bPjO'lB the R'plZitlglon PhlTooophital ,'Joeiellj. 4th Octobea, 1911.]
THOUGH the cOlllpo&ition Il.nd molecular weight of nitric acid are l-nown
to be represented by the formula HN0 3 , I have been unable to find in
any text-book a method of &howing to a class that the r,ubstance really
has the formula assigned to it. I have therefore worked out the details
of a comparatively simple method whereby the demoD.lltlat.ion may without
difficulty be carried out.
1. Strong nitric acid il:! prepaled by distilling c:l. mixture of sodium-
nitrate with an exce&s of 98 per cent. l:!ulphUlic acid at as Iowa. temperature
as possible, and the resulting acid iR further dehydrated hy distilling at
sbout 20 mm. pretlSure (vacuum of the water-ejector pump) with three
timel:! its volume of strong sulphUlic acid, Bnd condensation of the acid-
vapourl:l by meanr, of a good freezing-mixture. Diffusion of the aqueous
vapour from the pump into the distillate is prevented by means of a tube
containing pumice moihtened with suiphUl1c acid. An acid pl-epared in
this way will be found to be practically colourless, 8ud to have a specific
gravity ILnd titration value cOl'l'esponding to over 99 per cent. of pure
nitric acid.
2. As soon as the acid is prepared a number of thin-walled glass bulbs,
each capable of holding about 0·2 grams of acid, and blown on capillary
stems Itbout 3 cm. in length, are filled with the acid by placing the bulbs
"ith the open end of the r,tem downwards in a beaker containing the
acid and placin~ the beaker in 0. dC'licca.tor, which is then evacuated with
the aid of the filter pump. Upon readmitting the air the bulbs will be
founn to be completely filled with acid, oxcept for the presence of a very
minute air-bubble. The euds of the capillarieK ure now seall>4i over a small
:flame, aud the weight o[ the contained add Ilsccrtained.
3. The apparatul:I ill which the analysis of the a.cid is actually e&med
out is as follows :-
* Playfair Ilnd Wa.nklyn... Journal of tht' Ch('mical Society." vol. 11), p. 142.
ART. XXXII.-·PJte Zntel'actioll 01 IrM with the HigJler Fatty .J.C'lds.
Bv TnOM.\El H. .E.o\ATElIFIl1lLD J,lId CL..UIA ]\lILLICElI.oor TAYLOR. 'l\I.A. (Nllw
. Zealand 00\ t'l'nmcnt Rebearch Scholar)
llleatl before the Ji'ellill(lful£ J>hiZo80pl£1oo,l FJocietl/, 9th Augu.t, 1911.]
INTRODlJl"rroN.
IN a former papm'" It was shown that under tho action ot llletalhc Iron
abietic acid is rapldly deprIved of ita cal boxyl group WIth Tlrodllction of
a hydrocarbon. It is well known that thll higher fatty aCId& under con-
chtlOns which should lead to dcprivation of a carboxyl group yiC'ld kt'tonc'l
more easily than hydroc':l.l·bouF., accordil1~ to the equation
2X . COliH = XlCO + CO2 + HlO.
1t theroiore seemed probable that heatln~ the hi~her fatty acids wlth Iron
fihngs would be a simple method for obtainin~ ketones in good yield.
Upou heatJng &tearic acid witl, caRt-iron turnings to a temperature
of 360-365° C. it wos found that ovcr 80 pill' cent. of the acid was con-
verted to stearone. As the usual method of prepar~ the> ketone of stearic
acid only gives about 50 por cent. of the theoretical yield, the advantau;e
of the new procest! is obvious. Another J.nd equally important point 1"
that the quantity of acid which can be treaterl in one operation 18 almoEot
unhruited. In the ordInary process of dx&t1llin~ calcillm or harium stearate
with slaked lime under dimmished pressure from a combustion-tube the
quantity of ketone prepared in each operation is necest!al'lly small.
The method was also found to give good yields of ketone with lauric.
pQ]mitic, cerotic, montanic, and meliSSIC acids, so that it may be rega,rded
lib a general method .for the preparcl.tion ot the ketones of the saturated
fatty acids with from 12 to 30 atoms of carbon in the molecule. With
acetic, butYTic, phenyl-acetic, suberic. and sebacic acids no satismctolY
results were ohtained.
The ketone!'> of the hi~her uru.aturated £a.tty acids have not hithertu
b~en prepared. bllt the .• iron" method a.llowR thCl:!e compounds to be
ohtained without difficul1.y in the oleic series. In the linoleic serit's no
e.:l..-perlmelltt! have been made, owiug to the difficulty of obtainin.1t the acids
ill Ii state of purity. Durmg the progress of these experiments it was
pointed out by Mailhet that the vapours of the fatty acids from acetic
to stea.ric acid yield ketonf.>s if pcl.ssed OVf.>r gently heated .. reduced ..
nletals, includinl{ iron, cOllper, nickel, cadmium, and lead. There is,
however, an extraordinary difference between the catalytic action of the
.. l'8dllCed " metals (which a.re in gener.ll pyrophol'ic) and the same metals
in the state of po,vder. (Comp8J.e, for example, the inertnet!s of ordinary
plJ.tinum with the intense cata.lytic action of platinum-black).. Sabaticr
has, indeed, recently drawn IIttention~ to the fact that co reduced" nickel
exhibits quite Illiferent CcLtalytiC effectb upon mixtures of hydrogen anCl
ac~tylene, according to the conditions under which the reduction has been
c'uTied out.
EXPERIMENTAL.
1. Preparation of Stearolle. - Pure stearic acid is heated with one-tenth
of its weight of powdered cast-iron turnings ~o a temperature of 2800 C.
The temperature is then elowly rai&ed to 3600 , und maintained between
360" and 370" until evolution of carbon-dioxidc almost ceases-usually
abl)ut two hOUlS. The product is freed from iron by mcans of hydrochloric
or sulphUlic acid, and from stearic acid by aqueous alkali. The ketone is
mice Clystallized from light petlOleum with the addition of animal chllr-
COllI, and is then pure. The yield is 80--85 per cent. of that required by
theory. The melting-point was found to be 880 , as stated by Krafft. An
analysis gave- Found. Calcu1a.ted.
C = 82·71 83·00
H = 14·01 13·87
2. Preparation 01 Dihepta-decyl Owibinol. -1 gram ot Sliearone was dis-
solved in 200 c.c. of amyl alcohol, and reduced by the slow addition of
8 grams of sodium to the boiling solution. The secondary alcohol crystal-
lized out on cooling, and after several recrystallizations melted cODlltantly
at 89·5°.
Found. Ca.lcu1a.ted.
C = 82·31 82·66
H = 14·03 14-19
The carbinol yielded an acetic ester melting at 61 0 (not sharply) and
giving on analy~
Found. Caloula.ted.
C = 80·80 80·73
H = 13·51 13·46
3. Preparation of Oleone. - 5 grams of pure oleic acid prepared from
olive-oil, and melting at 14" C., was rapidly heated with one-tenth of its
weight of cast-iron powder to 240°, and then more slowly to 340° and
maintained at this temperature for two hours. The produC't was treated
first with acid, then with alkali, afterwards crystallized from alcohol, and
:fi.na.lly from acetic acid, until the melting-point was constant at 59-60°.
The yield was 10 per cent. of the weight of the oleic acid taken.
The same yield of oleone of the same melting-point was obtained in
an experiment in which carefully rectified oleic acid prepared from com-
mercial olein was employed.
A smaller yield of oleone of the same melting-point was obtained by
distilling pUl·e barium oleate in a partial vacuum to a temperature which
nnaJ.ly reached 450°. The distillate was rectified under reduced pressUl"e,
and the portion boiling at 280-3300 at 5-10 mm. pressure deposited oleone
on cooling. After several crystallizations the substance melted at 59°.
The yield was only 2 per cent. of the theoretical amount. Analysis 1 was
ca.:rried out on oleone obtained by the iron method, a.nalysis 2 with
oleone from barium oleate:-
1. 2- Calcu1a.ted.
C = 83·5 83·40 83·62
H = 13·6 18·10 13·14
The molecular weight by the ebullioscopic method in alcoholic solution
gave- M = 4:92 and 508. Calculated = 502.
Bromine absorption in twelve hours = 62-1. Calculated for 4 atoms,
bromine = 63·8
EASTEll"FIELD AND T.\YLoR.-Diyher Fatty Acids. 303
F::210a"7Zaru system
~EocenP-17Uocene
~VolcanEc
~Po'" "EOCene
ri:i'i:'1 Glaczal morcune
~ Plezstocene
o Recpnt
_Nephellmie
1 Inc.h = 8 mlle8
variety, with pronounced zonal and hour-glass structure, and with the
usual pleochroism, and occasionally shows the tlteel-blue birefringence
due to the high dispersion of the optic axes. A narrow margin is
usually dark green. The mineral is comp.letely idion:orp!ric. The n~phe
line is abundant, For the most part Its cl'ystll.llizu.tion was firusllCd
before that of the feldspar. It is largely altered to natrolite. The feld-
spar ~ much twinned on the albite and pericline laws, and has in
many places the appearance of microline. The extinction-angle, however,
proves it to be andesine. It was the last of the larger Cl'Ybtl1is to
form. There is here and there a small quantity of groundmass. It
consists of idiomorphic and often bent crystals of feldspar, probably
andesine, allotriomorphic aegerine, nepheline converted into natrolite, and
much apatite. Of these minerals, the nepheline was the last to form.
In some of the finer-grained spooimens the idiomorphism of the augite is
less pronounced, and occasionally shows ophitic structure with the feld-
spar, which is then distinctly anterior in crystallization to the nepheline.
The basaltoid forms of the rock are well exemplified by a large dyke
at Clarendon (analysis B). Here apatite is much less noticea})le. The
ilmenite and olivine are not more tha.n 0'25 mm. in diameter. The
augite is in modera.te to small crystals, sinking to the dimensions of
micIOlites, but always idiomorphic. Feldspar is not abundant, a.nd is
always in the form of microlites. Nepheline is quite abundant, and is
in the form of allotriomorphic plates enclosing numerous crystals of olivine,
augite, and ilmenite. Lava-flows that cover a considerable area of the
country to the west of the dyke are also somewhat similar, but are of
much finer grain (analysis C). The irregular plates of nepheline are in
these rocks extremely small and hard to distinguish except by micro-
chemical methods. This type of rock has previously been described by
Andrew, who, however, failed to distinguish the nepheline, though he
recognized that much of the rock was soluble in dilute HOI.
The type from Omimi is particularly interesting from the point of
view of structure (analysis D). The apatite and ilmenite have the same
features as before. The olivine, however, is in extremely small needles,
sometimes 1 cm. long, but only 0·08 mm. wide. The direction of neigh-
bouring crystals is in remarkably parallel lines in longitudinal as well as
transverse section. They are similarly oriented ovel' a considerable area.
The phenocrysts of augite have pleochroism, zonal and hour-gla~1! structure,
as in the Waihola type. A similar appearance of lattice structure in the
feldspar is also ~ery noticeable. The nepheline is wanting in crystallo-
graphic boundaries, and is usually.intergrown in complete micrographic
fashion with augite. In some instances at least this augite is in optical
continuity with the large crystals. This micrographic intergrowth is
sometimes found in the groundmass in an extremely minute scale, and
constitutes its dominant feature. The augite is sometimes &lightly greell
in its smaller members. There are minute crystals of feldspar and apatite
crystals in the groundmass. The intergrowth is of the same nature as
that found in the Lobauer Berg type, but is much more complete, and
is shown on a finer scale than in the German type.
.As at Waihola. the rock is associated with basaltoid types, the exact
distribution of which ani their relation to the coarser type of rock has
not· yet been fully made out by Mr. Smith. Here, however, there appears
to be a complete series through types with large allotriomorphic nephe-
line plates to types of a dense nature in which the nepheline is extremel~
hard to identify. .
).r\R~HALT,.-)leJllU'lillife Roc7~x ill .YelL' Zealrllld. 307
By G. L. ADKIN.
Plate& XXIl-XXI\',
THE diSCOVelY 01 evitlelice of formcr ~lo.cicl.tion in the Taull'ua Range...
\f'a.1\ made bv' the miter in Febl'Ua1"lT. HI09. and two vearb Jnter (~1nlcl'.
1911) fUl'ther diaco\'prie:. wcrE.' made alld the previo~ ones confirmed.
The ~lllciated areal:> and the memOl'ialb . f {Olmer froRt-action I:!O far di~
covcred are situatl,d on the highel'lt rangeb of the Taral'un8-\'iz" on that
part of the Dundas Range lying nearest the geo,-\1'1lphica1 centre of the
mountain-sy"tem. and on the Mitre-Holcl&worth Range, During thl'
maximum phase of glaciation the heads of five rlver-vallE'~·fo. wcre :6.11l'd
with glacier-ice: (1) Park River,* the mam trIbutary of the Waiohine-ih
River, named after the well-known New Zealand Il,ladalibt; (2) the rot-in
source of tho Waiohillc-iti River: (3) Dorset Creek, a left-bank trihutaryof
the Waiohine-iti River. named alter a pioneer explorer of the Taral'Uus;
(~) Bennington Cl'eek, a hibutalY of the Waingawa River. lil:>inp, in the
south-west foot of the Mitre Peak, named after a companion of Edward
Dorset: and {5} the l\langaterera Rh·er. another tributary of the
Waingawa..
The phenomeno. rt"sulting hom the plesence of glaciers now non-eDstCl.t
consist of-(l) U-shaped yalleYI:!; (2) glacial cirques; (3) rock basillt-:
(J.) glacial1u1.nl!;ing valleyr.; (5) fluviatile hanging yalleYI:I.
In order to give a clear and COl'lect impression of the extent and c1J..t-
rllcter of lormer II,laciatioll in the Taral'uas, the phenomena tabulated 'tt 111
filat be dealt with sl'rilltim, dond. thl'll the topo~raphy of the Park Vo.lIc~·
-the locality where ther.e phenomena attained their maximum develop-
ment-will be fully del:!cribetl.
* '!he river draining thi, valley ha.s hitherto ~ither been na.med nor shown on
any availa.ble m&:{I. On every ava.iJa.ble ma.p the ma.in BOuroe of the Otaki RiVQl' i..
represented 80S draining the &ite of the upper portion ot Pa.rk Va.lJ.ey.
PLATE 'SIll.
THANa. N.Z. IN&T., VOL. XLIV
FGI.... O iIJ'll
.\.Dh.I\ -Pili 1111 I Gl'l('wfIOIl hi the 'l'(lI'al ua Ral1"f\ 309
310 Transartions.
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PLATE XXIV.
~\IJKIN.-POln t!1 (/farllltlOll ill flu. T{lI'UI'IIt1 .Rall{I'. 313
The muin cirque at the hend of the valley hItS a diameter of lleatly half
a mile. The preeipiceK forming the bounding wall~ ot the cirque attained
a maXimt1m height of 800 ft. aboye Hs floor. Below the cirque the valley
i~ U-blJ.apecl fol' ahout two milt!t1, the ~eer lateral walls having a height
of upwards 0.£ 4.00 fi. The U shape of tbe valley is less pronot1Uced towarlls
it~ lower end, and h\o miles below the cirque the lattcc ltarJ'O\l"tI, ~radu
ally becomes V-~haped, and filiully gorged. The continuity of the bounding
precipicelil of the main cil'ql1e-which are best preserved. on the bouth-,...~t
face of .Al:ete Peak-is brokeu hy the three U-rillaped glacial hangin!l, valley,..
The largest of these lies 011 the south &ide of A.l'e1 e Pf'ak, and lUleS in a
A'R.!T£
'P."I(,,
MT
L.ANC"STEII.
I
•
M!"
ZlO"'A.
... ....
Ai\I!.Tl!:
"'£"1(
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ARaTC
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po"",,, "..,0 OT,,"" JO."...
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Mr DalIlA
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o , •
va" TICAl..
1001> _
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0111. ", ... C.
The most striking feature of the main U-shaped valley is the high
development of acreea. These bUl-y the precipitous walls to a height of
from 250 ft. to 320 ft. above the vallev-Hoor. Above the screes the lateral
walls rise to a height of from 50 ft. to' 100 ft. In the main cirque the Pl'e-
. oipices rise 290ft. above the apexes of the screes. Throughout the greater
palt of its length (he floor of the main yoJ.ley is loaded ",ith scree-material;
the bases of the screes on the one wall meet the bases of those on the other,
and the modern drainage-channel o{ the valley {ollows the line of contact.
The screea are now not in the course of formation, being clothed with
tussock-grass and subalpine srrub.
In the U-shaped section of its valley the Park River is actively engaged
in alterinp; the gradient of the valley-floor. In the main cirque it is an
aggrading stream, and has theJ.'8 formed all alluvial Hat several acres in
extent. Below this flat the river flows in a narrow channel of gradually
inereasing depth. Near the lower limit of glaciation this ohannel is about
20 ft. deep. and the rock Hoor of the valley, upon which the screes rest, has
been incised by the ri"er to fL depth varying from 10 ft. to 15 £ti.
The infilling at the head of the valley, and the excavation below, clearly
demonstrate that the "alley was overdeepened· by the old glacier. After
the disappearance of the ice the rock basin was probably the site of a small
lake until it was filled in by the accumulation of alluvium.
Such criteria of former glaciation as moraines, roehe '1ItOuWl'IlII6es, and
striated surfaces have not been found in Park Valley or in any of the other
glaciated areas of the Tar8ol'U.80s. It is highly probable that some of the
phenomena enumerated do exist, but in Paxk Valley, and in the other
glaciated localities also, the present excessive accumulation of scree-material
and alluvium precludes all possibility of their detection. The apparent
absence of a terminal moraine may be accounted for by the small size of
the glacier. It may be, however, that some of the angular debris resting
on the valley-Hoor near the lower limit of glacia.tion is morainio materia}
laid down during the slow but regular sru.'inking of the glacier during its
final retreat. Another suggestion is that the great piles of boulders that
encumber the narrow gorges situated immediately below the lower limits
of glaciation in Park Valley are the re-sorted relics of a terminal moraine.
According to this suppositiou, the terminal moraine of the old glacier was
demolished and carried to lower levels "ince the disa.ppearance of the ice
by the periodic floods of the modern river. In this wa.y the angular blocks
forming part of the moraine were rounded and transformed. into the
boulders as they now exist. The boulders in the gorges referred to are
very much larger and more numerous than any that lie within the
glaciated upper portion of the valley.
The following altitudes in Park Valley W91.'8 determined by the use of
an aneroid set by the trig. on Mount Dundas: The saddle in the watershed.
of ~ he Dundas Range a.t the head of the largest glacial hanging valley,
4.440 ft. above sea-level; the lip of the largest glacial hanging valley.
3,900 ft. j the lips of the twin glacial hanging va.lleys, 3,750 ft.; the centre
of the alluvial Hat in the floor of the main cirque, 3,380 ft.; the summits
of the precipitous rock walls of the main U-shaped valley-left wall
3,800 ft., right wall 3,670 ft.; the lower limit of glaciation (i.e., of the
U-shaped part of the valley), 3,000 ft. above sea-level.
ADKIN.-Jt'OI'm.e1· Glaciation in tlu Z'ararua R(lllue~ SUi
GENERAL CONCLUSIONS.
The former glo,('iers of the Taral'uas owed their existence to the then
greater elevation of the country and to the more rigorous climatic con·
ditions. At the present time the snow·line in. the latitude of the Tararuas
is about 8,000 ft. above BeQ-Ievel. "The late Sir Julius von Haast, in his
'Geology of Canterbury and Westland,' estimates that during the glacial
period the snow-line was 1,000 ft. lower thnn it is in New 7.ealand at the
present time."· This estimate involves only a slight reduction of the
annual temperature-a. reduction presumably induced by cosmic or ex-
ternal causes or conditions-and appears to have been based on such, other
factors being neglected. The evidence furnished by the configuration of
the bed of Cook Strait (as shown by soundings) and by the physiographyi
of the lowlands at the western foot of the Tararuas indicates that the ele·
vation of that part of the country has been reduced since the glacial period
by at least 1,000 ft. Taking for granted that these estimates are correct,
and that they represent the sum of the influences that lowered the snow-
line, the snow-line in the Tararuas during the glacial period was. in re-
lation to the present sea-level, 2,000 ft. lower than at the present time;
in other words, the snow-line of the Tararua.s formerly stood at a height
of 6,000 ft. above the present sea-level. But it is evident from the known
altitude and position of the extinct glaciers of the Tararuas that the lower
limits of the permanent snowfields that fed them were at the most 4,000 ft.,
and perhaps only 3,500 ft., above the present sea-level. By taking as
correct even the greater altitude-i.e., 4,000 ft. above the present sea·
level)-there is a discrepancy between it and the foregoing of 2,000 ft.
This lack of agreement between the tentatively adopted a.nd the actual
altitude of the former snow-line may be removed by accepting one
of the following amendments: that during the glacial pt'riod the
snow-line was lowered (in each case with reference to the present sea-
level)- (1) by more rigorous climatic conditions 3,000 It., and by the
greater elevation of the land 1,000 ft. ; (2) by climatic conditions
1,000 ft., and by greater elevation 3,000 ft.; or (3) by climatic conditions
2,000 ft., and gtt'ater eleva.tion also 2,000 ft. The last of these is probably
nearest the truth, since the estimate tha.t the snow-line Wo.s lowered by
climatio influences only 1,000 ft., as pointed out by Mr. H. Hill (Trans.
N.Z. Inst. vol. 27. p. 453), "is a very small one, representing, as it does.
only a difference of about 3 degrees of temperature; and this certainly
would not be sufficient to bring about a glacial climate in. the South Island " ;
and, of course, still less so in the North Island, unless it can be shown that
the latter stood at an enormously greater t'levation in the early Pleistocene
period.
The former glaciers of the Tararua Ranges give some indication of the
extent and nature of the Pleistocene glaciation of the North Island: they
show that in these respects it was limited, localized, a.nd moderate. On
oomparison this view is found to be in harmony with the known extent
• lL Blll, " On the BAwke's Bay Pleistocene Beds a.ud the Glacial Period." Tra.Da.
N.Z. Inst., vol 27, 1895, p. 4;P2.
t G. L. Adkin, .. The POIIt.tertU.ry Geologioal Hi6tory of the Ohau River. &0.,"
TraD8. N.Z. Inst-, vol. 4.3, 1911, P. 496.
316 Transactions.
... See Bell, Webb, and. Clark. 'Bulletin NQ. 3 (New Series), N.Z. Oeol. Survey,
pp. 31.32,1907.
t JameR Park, Tra.ns. N.Z. Inst•• vol. 42. H)10, P. 38S.
~ James Park, .. The Grea.t Ice Age of New ZealGnd.," TranI!. N.Z. Inst., vol. 42.
1910, P. 599.
WII.D.-Geology of file Blllff. 317
• Rutton, .. Report on the Geology of Southland," Rep. N.Z. Gool. Surv., 1871-72,
p.89.
t Pa.rk, "On the Geology of Bluff PeuiDSUla.," Rep. N.Z. Geol. Surv•• 1887-88,
p.72.
~ Hutton, .. Notes on the Eruptive Rooks of Bluff Penblsula," TraDs. N.Z. lDst..
voL 23 (1891), :r.- 853.
§ Rutton, • Correotions of the Na.mes of some New ZeaJa.nd Rooka." Tra.na. N.Z.
Inst., vol 31 (1899), p. 484.
II Hamilton, .. Notes on the GeoJ.osy of the Muff Distriot." Tra.Ds. N.Z. Inat.. voL 1&
(1887), p. 45t.
'II Thomson, J. A., II NoteD on some Rooks from Pa.ra.pa.ra., Blull Hill, and Wa.ikawa,"
Tra.ns. N.Z. tDSt., vol. 42 (1910), p. 33.
318 Tran8(u:tion8.
., ! -au
BLUFF
1~
'lutoniC_NOrite. Tewawae'
onic_.BaaicI ~
Hypabasat-Porphyry.
Metamorphic-Hombl."Je aohiat,
Amphibolite.
y 1 If ? 3°CIIS.
mOl'C, some twenty miles to the east. ThuB a consiclenble area of wa.ter
was cut of! and enclosed by the sandspit. As the sanrlbank received
further additions it increased in height and mass most rapidly at its
eastern end, wher9 its onward progress was stopped by the headland
alJ'eady named. Finally, at its lowest end-that is, immediately
adja.cent to Blufr Hill-the waters broke over the barricade. and restored
communication with the ocean. The ohannel was made a.oross the
intruded rocks, possibly a.long 0. groove commenced by the a.ncient dvers,
and long sinoe filled in with sand and alluvium. This channel, (,Ince
begun, was rnpidly deepened by the inrush a.nd outflow of the tide.
which in the middle of the channel travels at the present time at the
rate of eight knots an hour. lIuoh is the size of the basin to be filled.
Subsequently this large lagoon was subdivided by la.teral sandspits,
and Waituna Lagoon was separated from Awarua Bay, and a 'leparate
outlet was formed.
320 Transartion,s.
For the purpobll of tl~hCl'iptiOll the l'ouk~ of tIm Bluff diRtrict 11.1'('
con~~nicnth' diviuell into t"o main divit,iom, dihtillgllibhed he1'(.' at.
A Ilwl B .. '1'11cbe rlivihinnh Ill'C-A, tIl<' iglll·lItl., 1'0ckR; Il, tho Illeta-
morphic l'ucktl.
Each ()f the!:!u (livihiollb m.n· h~ lUl'tllt'l' divided into dUIN!h, t[L(' l'ockl-
of euch ClUbb ill tLc divito.ioll h~'ing ~ntil·ul.v llihtinct. Tho i~l1~"IlH l'ocks,
Division .\., cOllsii.t oi-I. ignE!Ou!:! rock!:! or plutonic origin; 11, 19DOOUI'>
t'ucks of h~·pD.b~·S8ul origin. Th~ tOl'lll "h~'pabyssIlI" it. }U.lt·o ut.ell in
the b,lIlle llell!:!C us Harker* uses it, hyvub~ !:!aal rocks COl'l cspon!ling in II
genCl'nl war. though not precisd~', with the group "gnngestoille " 01'
" d~ke-rocks" or Roo,enbu"ch.
The ruetamOl'phic l'ockt-, Division H, coutuin tim CltH.hUb, 11.1. followtI:
I, acid metamorphic rllck!;j II, bu.&ic metamurphic rockH.
An attempt will be 1111'110 btllo,," to Rho,," th·tt tht:.re is an intilll:.tte
connection betwo~n th~ acid tnl:!ta1U(J)'vhic l'lIckR (Divitlion B, Clab!! I)
and th~ igntlOUR rockH of h~·IIl.Lb~·bbnl 01 i~in, thl:! llletalllOJ'phio rocks being
derived from the h.'l'abY'lsnloues. Thil\! i~ not the view tnlton by 11l'Cviolli:I
invostigntiollR-c.y, Hutton Imll PUl'k-IJut tIle pl'~ht!nt tLeory is the
oulCOll1~ of the e-.:aminntion of material thnt had not pl'eviom.ly beoll
brought 1I1111el' i""l'lct:H1I. Thl' point will hi:! fllll~' llihOl1'l'll.,d ill nil' COUl'se
of this paper.
The l'elation~hip amollg some clnsses and the ahllellct' or l'elatiolltlhip
among others thull bl'ieB.~· mentioned llul.kes it difficult to suggellt uny
pcrfectly e.nti"fuctol·Y heheme of clnssification.
• Hazbr. A., " Petro~ for Studenta." 4th ad. (1908). • lOS.
t ltep. N.Z. GeoL Sarv,. "The Geology of BId P"nillllU~ .. 1887. po 89.
WILD.-Geolog1l of tke Blvfl. 821
Under the MicroBcope.-Exlilllino.tion of thin sections shows the rook
to be composed essentially of feldspar. augite, hypersthene, and horn-
blende, with magnetite aR an accessory constituent. The feldspa.r, which
on an average forms half the rock, occurs usually in plates, ranging in
size up to 1'4 mm. long by 1 mm. broad. It also is found in irregular
pieces enclosed ophitically by the ferro-magnesian minerals. It is a
triolinic variety, showing both ooarse and fine albite lo.lnellation. 'The
extinction-angle ranges up to 27°, this angle being the one recorded most
frequently in sections 8.11 nearly perpendioular as possible to the albite
lamellae parallel to the brachipinaooid.
Aocording to the statistical method of Michel Levy for determining
the feldspars, this angle denotes labradorite. In &ome cases, however,
an angle of 160 in found on each side of the trace of the twinning-plane.
In such a basic rock. this figure indicates andesine, and Thomson thinktl
this is the preva.iling species, though he affirms that .. probably more
than one variety of feldspar is present." Undulose extinction IlDd the
bending of twin lamellae in a number of the crystals give evidence of
considerable crushing.
Of the ferro-magnesian minerals hornblende is the most prominent.
It frequently occurs as a fringe of varying breadth bordering the crystals
and masses of pyroxene. In thetse cases it is a pale-green colour, and
rather feebly pleochroio on the inner margin, but in the outer portion of
the fringe it becomes more compact and denser in colour and pleochroism.
changing from yellow-green to browny green.
Often, again, the hOl'nblende occurs in masses apparently independent
of the pyroxenes. Under these circumstances it is oompact, brownish-
green in oolour, and strongly pleochroic. Thomson* mentions that «the
oores of the hornblende crystals generally consist of 0. paler variety in
optioal continuity with the green exteriors, so tbat the fonner pre-
sence of pyroxene is suggeRted." This point will be further discul'I8en
(pp. 331-2).
Both mOl)oolinic and orthorhombio pyroxenes are to be seen. Augite
(lCOIl" in rounded grains and in irregularly shaped crystals. It i"
oolourless and non-pleochroic, and, where fresh, shows brilliant polariza-
tion ooloul's of the ReCond order. But often it is cloudy, and shows signs
of decompoRition, which, no doubt, ultimately lZives rise to the particlt'S
of chloritE> l'E'corded by Huttont and'l'homson.*
Diallu.ge uillo occurtl, though somewhat spnl'ingly. It encloses minute
tabular 80ales of 0. rednish-brown colour, al'ranged parallel to the baMal
pla.ne, giving it the schill~r structure, which distinguishes it from a.ugite.
The orthorhombic pyroxene is hypersthene. In some cases it exhibits
schiller structure like the aiallage, but it may be distinguished from the
latter minera.l by its pleoohroilll'D. and by the fact that it extinguishes
straight. These characters also distinguish it from augite. The pleo-
chroism showed a or 11 brownish red, b or 'jj readish yellow, 0 or J
green, pale watery colours in each case. To determine definitely that
the mineral was not enstatite, many sections were e:ramined in convergent
light to secure an optic axial interference figure, and thus find its optical
character, but these attempts were unsuccessful. However, the pIeo-
----------------------------------~--- -----
• Tho~ J. A., ,. Notes on some Boob:from Parapa.za, Bld Rill. a.ncl Waib.......
Trans. N.Z. lDst.. vol. 42 (1910). p. 33.
t ButtoD, F. W., .. Notes on the Eruptive Books of Bluff Penilllltila." '.I'nw. N.z.
Inst., vol. J3 (1891). p. 353.
ll-Tfana.
322 l' /'(f /I ~llI'f ;01' 11
* Thomson, J. A" Co Note'J on Knmc Roob {rom Pa.ra.pa.Ta, Bllln Hill, and Waikawa.."
Trans, N,Z, TDI!t., vol. 42 (11.110), p, 33,
t Hamilton, .. Notl'q on the neology ot the Bluff Di~trict'" 'l'rallfl, N,Z. 1nst. vol. 19
(1886, p. 452, '
t Vo~, J. H, L., .. Ph),'loika.lisohe.('heluiRcho Gcsot.zu dcr KriHtalliHatioD folga in
Eruptivgest.eine," INoh. min, II, pt'tr, Mitt. 24. p,4.17, 1900,
§ HUtton. F, W,. "Correoti(111ll of thr Na.mf)R of IIOnlC Nl.'w Z('IIlnnll Itucks." 'rrllllll.
N.Z. In&t" vol. :n (IR9!ll. p. 4114,
WII,]) -fJ.l/olaf!!1 of fill I1fufj 328
'rhe ildpl'PllC{' tll,tt all the horllhltmdt! iu thi~ lock 11'1 ~econdal'y 1&
~llppol·ted II.) dt'loocriptiom of tlimilal aretl~ in vnriuuR parts of tIlt"
WOI'lrl. Huch ;U'ell!'! have becn (leRcl'iben h.r Il'vin~i' and by Williamllt
in .-\lllt'l'ic:t, by Phil1ipKt ill COlIn\ Ill, hy Rensch§ in Xorway, by
IJPlllllllnnll in RU~OIl~', by Beckc~ in LonN' AU':ltria. by Wadl!,\\,orth*'"
ann by Hawestt ill .\lIlutica, un 11 IlIOle rl'('entl~' b~' Harker;:j: in the
'I"est of Rcotlallll.
All regardb the CIlUl>ell that produced the alteration of the pyroxene
little is yet known. rfhe experiments of Mitbcherlich and Berthier (1824),
Hllstav Rose (11-131). and Pl'oIuhllorh FOUgl' and Michel Levy, of Paris•
.md the recent reHearcheh of Vogt. Joly. Cm~ack, DoeIter, Brun, Day •
.\llen, and othel'lI have shown that IlUl-£ite apvears to be the stable form
lit high temperatures and hornblende at low temperature!!. From this it
mlly be assumed tl.at un~· condition tendinf,t to facilitate molecular
readjustment at ordinary temperaturell mUllt necessarily tend to facilitate
the change from augite to homlllt'llflc.
These consideJ'Utionli inclined WilliallJb*~ to aHcribe thl' Ul'ulitizutioD
of some rocks to the action of great }lI'CRI'IUl'e. such a.s might be exerted by
the llplteJ.vn] of lll'ltmtnins. ann TJPhlJ1ftl1nlllmd Hatoh~~ rtlached similar
conclusions. Rubs~uently. however, Williamst decided that, though
pressure may, al1d doubtleRIi does in lUany instances, IlSRist in the para-
morphism of pyroxene in rocks, it cannot in all cahes he re~arded as even
II necessar,\' adjunct.
In the c ise of the plutonic matlto. of rock forming mIdi Hi11 the follow-
ing: pointll nre put forward merely as suggestions.
The magma was intruded at sufficient depth to allow of the iormation
(If a holocrystalline lllRSS by slow cooling. At the temperature of the mass
augite wall formed. When ol'dinary temperature way reached the augite
would tend to change to hornblende if conditions should change so as to
induce unstable equilibrium in the crystals so far as the molecular forces
were COI1COI·nl;:'l1. Such a change of conditions would pORsibly be brought
about by either or all of the following : -
(1.) Diminution of prellsure by rltllludation of the overlying rocks.
This undoubtedly took place. bllt whether it wouln tend to induce
1ll01CClllar readjustment is a matter for speclllation.
(2.) Movements of depression and elevation dellCl'ibed above.
(3.) Lateral pressure due to the folding to which the whole country
WIIS Hubmitted in IntI.' Palaeozoic or early MeHozoic times.
* Il'Ving. R. 0., •• Origin of the Hornblende of the Crystalline ROl'b ot the North·
wehtern States," Am. Joum. Sci., vol. 26 (1883), P. 32-
t Williams. G. H.. .. Tho G&bbrol! and Associated Hornblende BOl.'k~ occurrin£t in
thl' Nei2b,bourhood of Ba.ltimore, Md.... U.S. Gool. Surv.• Bull. No. 28. 1886.
t Phillips, Quart. Journ. Geol. Soc., vol 32 (lS76), P. 155. and. vol. 34 (1878), p. 471.
§ Reusch. "Die foesilienfubrenden krystallinen Schiefer von Bergen in Norwegen.··
Herma.n tr&Dsla.tion by R. Ba.ld&uf. 1883. p. 3.3.
I' Lehma.nn, "Untersuchungen llber (lie EntMtehung der altkrystlLllinilK'hen Schiefer-
gt...tt·ine." p. 190; Bonn, 1884.
~I Becke, .. Minera.logischl' und petrographi!lche lIittheilungen.·· vol. 4, p.357, 1882 •
.... Wadsworth, .. Bulletin lIuseum ('omparativt" Zoology of Unrv:l.rd College. Cam·
bridge." vol. 7, p. 46.
tt Ha.wes. G. W.• Am. Journ. !::lei. (3), voL 12. p. 136.
U Ha.rker, A., .Mem. GeoL Surv•• Tert. Ign. Rocks Skye (1904). p. 319.
§§ WiDiamR. G. R., Am. Joum. Sci (31. 28. p. 266 (1884).
"I~ •• Mineralogisrhl' und petrographischl.' :\littht'ilunl!l'lI," .01. 7. I). 83 (1885).
11-
324 'l'1'(lIlRUcfi()llh.
III thih vapt'l' tht· lIllllll.l .. 1l0l'Itt''' hm. htll'Ll ullol'll·(1 in nt'rol'clulll'C
with the l'ioWI! of HUl'kol' and Roscnh1ll,ch.
As has b~ell Jolhovm above. the Hlull rock ('tlut-I!dK I'hKl'ntially or 1\
soda-lime rel!lKp ,II' , a monoclinic PJ"J'O:IOUl', 1111(1 Ull fll·thol'holllbic
pyroxene. Both kinclH or p:rrox('nc ,\I'C rhanging to hornbll·l11h,. nncl 111
tl;e case of C1',' stub. will' I'e the change IH C·lllllplcU-d. it ill iml'0"Klbll' to SlI,"
"lu;!thcr the hOl'lIhlt'IHlt' iR nl'l'iv!'d 1'1'0111 ol'thm'hOlllhi!' 01' {"om lIlonoclillic
pyroxene. Howt>wl', 110 far al:! IllUY be jUflgotl l'l'om what remains, the
orthorhombic val'i~tv is dominaut. !tllll tlu' l'Oc], iM thC'I·(·I'ol·(' II nOl'itl:
I'ather than It gabbro.
Ohemical Oom,positiun,-If the chemical composition be appealed to,
as some authorll demand. it also will he found to HIl)l}lOl't the clnsRiticatioll
here Rugg~sted.
Si0 2
Ti0 2
AI 2 0 J 20·85
Fe 2 0 3 ... ·85
FeD 10·55
MnO 'I'racl('.
C8.0 H5
MgO 3·99
K 20 0,63
NasO 2·73
fI 2 0 and loss on i!(nitioll 1,00
Total 99·HI'i
:;PI'('IDC !!l'!l.vity, 2·6~
2. BaRil' Secretion.
At Htn.rling Point tht! nOl'itf' t'nc'loR~K II lIUlhh of II flal'k-I'nllllll·tlU )'l\th('1'
thw-grainecll'ork. Hlwl'ific g'·lIvit.,· - :J'O:JIi,
The size flf th~ IloaHH l'annot be millie out definitely. aM thcre il:! :t
o(,lLting of Huil at this lOllality. 'l'be outUl'OP is slltull, hchg ('SPORed. ill
lL cutting tthllut half u cha.in in It.'l1gth lind 6 ft. or H ft. !l('ep. But.
judgillg b~' Ihl' I'IJ"ition or the oU"Wl'OpS of norit(' nroltuc1 it, th(l Rl1l'fa(,~'
t'xtt'nt of tlll' IIlll!>h CUUIlot bll mOl'e thun n chain iTl (lilllll!'h"l'. .\\1 hl1lo1
1l11'enu~' hl'l'll btutt·cI. the nOl·ite vades much in 1(!\tUI'C from point tu
puint. ulIll ill MIlW' easel> approuchell to a material lIiUlilul' to that 1l()\\
I1n dll I' clebcl'iptioll. 'l'hiR led Huttoll to dploicribc RPvt'l'1I1 vnrictillll of I'ocl,.
fl'lInt tIlt' cli"tl'ict.
UncI!!} the micl'Ol:lCoIJC til(.' rock 1"·~Ht'llt.. It loIilllil;~I' ILlllool'mblugl' of
mill~ralH to the nOl'ite, but illN'!' ill II VI."I'\· unu.. wul·th" inul't'lule in the
propol'ticm ..i feJ'l'O-magllcsiun minel'nlR. 'H()J'llbh'll!l!:l' r01'lllR mere than
half the rock. magnetite iR nbunllant, there iM Il little PY"'o'tcne, and a
basic plagioclase, sometimel! containing neelllPti uf npatitl', forms the
l'('St. Tlll'l'c i" nn npprtJllch to a rough ~npisRic IItl'1lctUl'e.
Strurtlll'l' und Oraf1' of ('1'l/8f(lZli::afioll .-The I'uck lluH a. coal'seh·
schistose 01' gneissic Iitl'Ucttll'C, IIncl this tennN to ubscurll the order ~f
crystallizutiun, so that it ?annot 00 lUade out with nl1~· nOg'roo oi n.CCU1'3CJ .
Hurnblende l'Ieemtl to domlnate, but the edges of it" (.'l'~·lltt\lH are e:rtremel~·
ragged. In faet, 110 minel'al Illl1l he said til be icliumlll'phic. and thl'
Wu,D.--G!l(J10[1lI of 1M Blllt!
• Pl'rk•• J.... Note!> on the O~ulogy (I! 'Bluff P('mn...ula,'· Rep. N.Z. (.1(,01. Rurv.·
]Khj-S~. p. 72.
t Hurklor, .A., U Thl' Natural History of hlllE'IIU'" R(l('k"l," p. 322. Hloo.
326 TrOIl,a/'/IOII ,
100·12
t5.l'eci1ic grlo\'lty, 2·5.
A. l)ol'phJl'.\', 'l'e\\ uewlle PlJint, Bluff.
B. ll()l'ph~"l'~-' i\IitlhouJ'i. l'ompohed }ll'illCipull~ uf orthoclase and
quartz, with I:IOlllU lllicl'olillt!', pillgiucluse, and biotite, allll winoI' Ilccessol'.'·
minel'llih. (" .\UUh·HI .... of noek ..... " F.H (fenl. Surv .. 1·'. W. Clal'kl',
1904:,) .
C. POl'ph) l'~', :\Jount ZWll. (JOlltltins Hl'thnclasc, plng-ioclase, quartz,
biotite, apatitE', llltlgnetitc, and zircoll. (.\.wII, L. n. Ellkim.)
D. AlkaliQ'I'II11it }ltll'(lh~T mit l<:ilJHpr('n~l. VOll Orthoklll:o. Illlll Oliao-
klns; grul1l1mtlilllt.' "E'I!t'utlich QUlIl'tll unll _\ulll'tllOklaH. (UUhl'nhmcb, E ..
" Elemclltl· d~l' Hl·Htein:-.lC'Ill'l'." HlOl, 1', 20;;.)
A stud~' of th~Stl anal~'ses Ilhows that compared \\ ith tY}licall'orph~'l'it!s
the Bluff type is relatively rich in the ol.ides of the hU'letl clilciUlII, mag'-
nesium, Hud iron, and conespondingly pOOl' in silica, wllile the pro-
pOl,tioIl lit alulllilllt 111111 I,lknlieb iH about lIol'mal. Further, it oompal't!tl
favourably with the anulYRes of the qlllll'tz-pol'pliYl'itC's except 1111 regal',ls
the pPI'centnge of alUluinn. In nthel' words, the rock is minera.logicully
It. porphyry, but ohellLicull;v it telldM tOWIL1·tIS the porpb,vl'itell.
\Ve conclude, thercfore, thllt from the evidence of both chemicnl ILIItI
mineralogical compol:lition the l'ock is It porphyry whi('11 haq h('E'n E'm'ichcd
with thE' 1>111:101:1 calcium, lllagllesiulU, nnd iroll.
FUl'tliC'I' ('unl'lidcl'utiollH III'" nceCRHlIr"\' hefore the lIlcthlltllli' this enrich-
lIl1'llt elln Ill' ,tllIlil'll. (Kee]l. :l:H, '" The Ol'i~iJ1 of tile Homblen,lt!
Achists.")
nn'ltlIOX B.-TU}: llET.A.lrORl'HIC 1I0L'KH.
These are found along the lIOuthern blhul'e uf the harbom·. The out-
IlI'Op is exposed between the level!! of Iligh nnd low wnter. Abo'Ve high-
water mark, Uti has beeu pointed out above, is a plain of marin...
de-nudo.tion covered 110W by" recent alluvium. On the other hanll, thE'
lowest ebb of the tide fails to rlitlclose th~ limits of the outcrop, The
main outcrop begins at HE'udel'ROIl Street, and tltrikel:! 150 H. of E. fOl'
a distance of 15 chains. The strike then l"al'ietl to E.S E., aud continllE'~
so for other 22 chains. when Il. southel'ly b~nl1 nf the eonRt ('uh off tlu:
ol1trl'op.
32t!
'1\\0 cOlllplctl· IIICl'ieK uf hl:lctiuuK "ere mlu1!:' 1'l'lIlIJ VIII·h·t i l'~ (If rock
ohtllilled b.,· Cl'UHKillg the Rtl'ike at right llugleK. Hl'I'iel- .\ \\ III> obtaincc 1
1I101l1-! till' lint' lllarked An IIll tltl' lUlll' (fig, 2). H(,lieh B \\11,.. ohtainl•• T
nlollg tht· line VD.
~()tet- of the micl'ohCopioal ('XIIUliuatitlll of ~C1'it,,, .\ aro Ilpl'enc.led.
~\. 1, - Low-water mark. 1:37 ft. from high-water mark, FeldslllLr
phenocrYllts fairl~' abundllnt; cI01111~' uncl clt'~OUlP()Hl'd; ROllIt' cnntllin
neeelleH of hornblende, HOl'ublencle in foImall cl'YNtnls; eHp('('ially nUlllC-
I'OUK ill tIlt' vicinity of feltlRj)lu'H; plIl'lIUel IIl'l·angemcllt. A littll' biotitl'.
Gra.ins of quartz, Ulngnotih... IIml felcls}llli' f(l]'111 arll\l11l1Ulll'll'o.
A 2.-97 it. 11 llcl1 the Humc as .A 1.
.A 3.-67 ft. I'Ill'nOcl'YKtll of fuli1sl'llI', Mllllt· IIhol\ ill!! ('III·l:.hlld twill-
uinll. decompusiJlg nil ill ~-\ 1 and A 2. TIl~I'c'ahe oj' 11OI'1Ihll'Ilcle t'elative
to felclHplu' compll1't'c1 with .A 1 and A 2. .\ litHt· l·pi,(lut('. (lI'otmc1rnMR
as before,
~-\ '" .--:Ji ft. Hornbl(·nrlt· IItill more Pl'lIlJlillt·lIt. I"l'ld"}llll' pllC'IIOCI',\'htR
unU:.·h Klllaller uucl !lUll'l' 11 <'I'olllposcd. SchilltoHl' RI rtwhn'(' HIlII·kl'cl.
)1 agnetitu IIbundunt ill grolilulmulIN .
.A I). - i ft. llol'uh1t·lHl!· dOlllinaut; lmlg'ituclillul axt'l! (If cl'\'Ktalh
II l1 l'n11('1. XI) phenocl'Yhb, of fl'111:'1'1I1'. .l\inglwtltC' ahulIcllLut. ...\ 'little'
epiitot(', Gl'ouuclmnl!R ,(mill:' oj' f ..llihPUI· uIH1 qnal'tz. fe1clHIIIII' PI'('_
dominant,
~\ 6.-High-wlttel· Illurk. Rock chiefly hOl'llblcnde. SclaihtOh(' ... tI'1IC'.
tm"e peJ'fectly 11110\\'11. OllC wction Khowed rOnlalll11 of II felchlpar phEmo-
cryst. Gruins of fc·111111\a1', qlllll'tz, IllIn magnetite, uun othl:'l' nC(lI'KI-I()J'ieH
betweell C1'YEltalti of h01'llbll"ndl·.
~-\.R will btl seen from tbt.' map. tht· n Hl'l'ies is uctuully II cCllltinllutioll
of the A sedUR. It ill 110t neCCI:IHm'~' to cl('~l'ibe the rocky'in detail. Thl'~'
are pel'iectl>· RChilltosl:' ill IItructul'l:'. Hall the rock iR hOl'nblC'llnr. Th"re
~re llII phenocr~'sts of fe1dRpar, but the ~l'llins of ql1ll1'tZ :U1I1 fC'lclKI'1ir
In the gl'oundmnBR Ill'e olear, nO) thoug]1 nUl' to l'etll"'·fltlillizlltion. Biotite-
is more abundant. '
SlIUlllling IIp the I'OKUItS of the exnlllinnt illll of thiR f1l'I'itlfl of roclul,
"e nncl thllt-( 1) the p}ll:'llOCI'~'Rtfl of fc·J.1Ml11I1' 111'(' mOl·... IInei lllOl'e bl·okc.>Q
«lo"n the 11tlo,l'er th~~ IIl'r ttl thtl plutonic mabil; t2) the decOlllpobitiull
of the ft'ldH)lo,l' phl'lIOCl'yloltjo, l'Pl'l"I'''p,mds to an increase in the nmount of
Lombltmdc ill the I'ock,
'l'hetle j'nctH will bt:' 1I1111lc 11M: of when we discuss the origin flf the
hlllllblenfle Hl'Itish. Cp, :3:H').
('llemical Composifi()II,-.A. ~.1l1l1)le of blleOlllltlll ::\0, 4:, bel'ills .A., was
"Iublllitted to chemical o,nlll~hi", "jtb tbe l'ejo,ult given in the following
table, This nnnlysis pl'ol)nhl;; represents the n,el'age cOlll},otlitioll of the
,~chistb, though microllcopil'al t:!xamination of sediom, leaus one to expect
Ulore acid results in the \ldSe of the outer members of the Reries, alul
UlClI'P b'lBic r .."nlt.. in th,' ('11.'1(> or tho~f' 1ll'lIl'er tlw nO! itc
SiOs
Ti0 2
Al~OJ 18,66
.Fe20~ 2'4.3
FeO JO'83
MIlO
CaO 7·35
MgO 1-44
K 20 0·52
~a20 1·90
Loss ou ignit,ion , , 1'20
Total
Total
Spec in,' gt'a vity, :!.1l4:.
The Contact. of (md 1111 Ilurl/bltl/ti, .... d/lllt/J.-~\t tIle
tn,f, .J1lIJi/ubu{ilt-
line of junction these two rockH III·t· Itllll'l' l'llllil,\' )','"ognized ill hnnd-
specimens than they are under the microllc()pc, for tho caUhCIi that pro-
duced the metamorphisnl in both tendeu til hl'illg about all oxcllango of
material between the two. Thull the amphiholite iR riChl'l' ill feldspal'
where it is in contact with thc Hchists, and the achiRtll ILI'(1 l'olatively
enriohed with hornblende. .
~r~r, A., .. Petrol~ for StuclentR:' 4th cd. (190'1), TI. 70.
'II
+W~~ms. G. H., Am. JUllm. Sei., vol. 2K (18IH), p. 2111.
t (-leikie... Text·book of (I('ololzy." VIII. 2. -l-th ('II. 1190:1). p. 790.
\VII.H. -fleolof!!! of tlu' lJllljt. aal
Severtheleslo. "iJlce purhnps the TOajOl'it,r of nuthorities give pro-
minence ill theil' cll'finiti'ln of th i~ lIIilleral to a fibrous or acicular
Htructure, thiR tlil:lt,inction "ill be ob'll'J"\-cn in the present paper, and
til£' tlefillitilll1 lLlloptetl here is as followl:I: Ul'nlite, IInie-green slightly
pleoclJl'oic fibrol1H vltriety or hOI'ul,I('mll>, nerin'll fl01ll pyroxene, The
hornblende or th(' 1I000ite. tlu'l'efoll', thoug}! of !I{'conclnl',r Qrigin, ilj not
in th;" paper l'eferrcn to as uril.lite, fOl' it is a compact variety, rather
dark in COlOI11'. IInll Ktl'on~lr pleochroic. •
llralitc WUI'! TIl'st rle~criben by ChIlltuv nOF,l' frollJ a goreen porphyritic
l'ock at MORtovaya, Ilear Eknterinburg, ann at Kaminskaya., near Minsk.
in tho Ural Mountains. It ImR 'lillC'(, heen obsorved f1'Onl Hlany localitj('~.
The microscopical stuny of I'ock!! hus Rhown the IH'OC't'S!! of ;, uralit iZII-
tion ,. to hl' VC1'Y common, and ROme authors rego.rn muny hOl'Ubl('ntli('
rooh and bchists to represent altered p~']'oxene rock!! on a lu1'W' scull',
TIm crystals obtained Iroll! tho dyke at Green Point alford ubununnt
and (''(cellent matE:'rial fol' the illvcstigation of the ehangeN that take place
wholl lLugite iR convel't(:'r1 into lIl'alite. Theso cl'YRtlLls are Ulmall.\- ~hort
.11111 .,tout, lind ~llOW un cqual development of the unit ).lritol!lll'o (J 10).
tIll' OI'tllo),illucoius (lOU). antI the clinupilltlcoidK (010), "hill! till' mmal
tel'HliIlll1 faces, the }IIuli pyl'amids (ill), are also pl:'rrcctl~· formpd.
T\\iDlll'd forms, "itlt the orthopinacoid (100) as twilllling nllel ('OIupu.. i-
tion plane, arc ako quite COI.J1loon.
'fhe l'esults obtained frOID 1111 (!xu,lUiulIotion of sections of some of these
cJ'ystals cut in various dh'ections will nmv be given.
The Core of Augite: The iutl:lrnal core or augite is culoul'les!!, except
where recrystallization has commenced.
Cleavage: The usual cleavage-lines are not very distinct in sections
in the zone of the prislUs, though they are seen well enough in Cr088-
tlections. What is very distinct in sections parallel to tho ortho- and
clino-pinacoids is II. seriel:l of parallel lines which intel'sect thc cleavage-
lines at angles appl'oximatin~ 700 in sections parallel to the clinopina-
coid, and at right angles in Elections parnllel to the ol'thopinncoid.
These lines thus represent a series of parting-planes parallel to the
base (001), a not uncommon feature in Il.ugite.
Rcfl'l1octive Index: A I'ough Rurface in polarized lip:ht indicates the
usual high vulue.
PleochroiRUl: Kot noticeablv.
Cro!!sed IlicolR :
Tllterfel'Clll'e colourR: Bright tints of scconn order,
E,tiuctiolJ: III hootimu,; Iill(lOO) = 3i o .
b (010) = (10.
Alterafion- prot/urts witllill til, Milleral. - rl'hl''''~ are feldspar anll
hOlllblelJde in nbont equal amonnt nnd a little olh-ina. The decollI-
position he~ills llt puints on the cleavage-lines nnd prllcl!('ns most rapidly
in th~ direction of them, The hOl'llblvnde is 11111'1;: I!retlD lIud strnngl~'
pleochroic. It ext inguishes when the c1l'll.ynge-lineh (If the augite 11.1'0
parnllel to the vibratioll-nirections of the nicolFl, 1'ht' It!ldspar e'{tin-
guishes lit small angles. Olivine occurs in small gl'ains; it is vet·y
rare, Th(,l'l' is no trace of calcite, epidote, 01' chlorite, lIliuel'als that
are l'OUIll1111l1~' repm'ted n.s associated with such changes n.s lne her('
described. The fact that hOl'ublenrle and feldsplll' are alway!! associated
as decomposition-products in the interior of the cl·~-l:Itn.ls l!ug'lteKhI tbat
th ... TlII)t(,rlnl tlE'l'iv('rl fronl the <1ecompol!in~ aug-itt! is divitlt'll btorwecu
1'1'1l ,1.m'/1I) /I ,
them the culciUlll .11111 1ll,lgll1'>.IUIII going to\\Jnl" tlll! j',H'uHltlOll (It tIlt!
oue, ;he alkali!:'!> amI uluillinu io ihl: UtllE'I' mint-mJ.
Tho intel'ulll dacolllpo>.itioll oj' tlll' lIugile "C(OIl'!'> III 110 \\IIJ m:,sociall'c!
"ith tIlt' formntion of Ul'alitl:' 011 itt, uuu'g-im., 'I'll(' 1111111('1' of ul'ttlitc i"
(luita Ilif.tinct, UOIl shows nH ~I'11I111iillll t()\\:tI',I" tIll ]lI',)(111I'1to. HI' d('OOlIl-
position in tho int!!l'iol' of th,· CI',\ IItal.. ,
']'111' Prillge 0/ rmf,ft',-H.llIa "U~1l ill hl!> .. H~l'ltl'lll HI MlIlI'l ulog~,'
.. 'l'he cl'\Rtalf>, "hell dihtinct, l'daiIl tllt' !OI'!l1 or thl' OI'iginul minel'ul,
but havo' th(' cleLLvagll oi amphibole. 'fhe clumgt' llRU.lllr COlll1nC'nCOR on
the surface. tL'nll",flll'ming the outC'r lurer into fill :tllltll'gILtiC)]L o[ "lenuC'r
unlphibok prisms. parallel ill position tu OUell (JUU."I' anll 10 thu parent
pyroxene. When the change is complete tilt' clltiH' ('l'Yiltul ill 1llade lip I)f
.1 bundle of alUphiholo neullIc" 01' fibres."
Present Obsl'l'l'atioIl8,-1'!le 11'illgu of ulalite Val'lel> iu \\ itlth with the
t>ize of the cl'Yl:ltnl. il1dioo.ting that thl:' U,lllOUllt of changl' vurieK as tht'
t>urflice el.l'ohcll. usuun~', howc"l"cl'. it il:l notiu('d that the ohunge hal-
t;~kell J:lllictl mort! l'lI.piuly in the dirtlCtioll u[ tlLl' vel'tic;tl axis thl:ln ill
the ot!J(,I' dil'ectioI1R. [01' the fring'lI bordeJ'ing the terlllinntiol1K of tlll~
l'l'~'''hllh ill willel' than that bUl'd(')'illg pl'iRmatil' f; U:C''' ,
Stlttemcntto. lIh.,nt the paltlllci Url'1I11gcIllunt of till' 111'jlllllh of urulitt"
010 not find support in un exalUinution of the I!e(ltiullS of tho Hl'ccn Point
l'l'ystals, 101' the fibrell are Ileen to be arranged in l'aUiatillg' gl'llups which
Khow no signs of systema.tic arrangement. Between ol'ollRed nicols IIOlDt'
of these groups, or parts of the group"', are extinguiI:Ihed, while other
groups are not. Each fibre extinguishes at an angle that vlLl'ies from
150 to 180 to the direction of the longest axis, 110 that a flul'k wave-
traverses the group as tho nicols ul'e rotated.
In sections parallel to the clinopinacoid the gl'OUpll conwlOnly make
an angle of 45 0 with the edge of the crystal, meaRul'ecl eithol' in a +
Ill' - diJ·ection. In sections pal'll.llel to the DlIl.cl'opinucoid lind to the
base these groups ure commonly parallel, nnd the ftbl't'll illlow straight
extinction. This sl:)oms to indicate that the fibl'es al'tl u,l'l'angE'il hi
rnn-shaped aggregates parallel to the clinopinaooid. nnll making angles
of ahuut 4!,)0 with the macropinacoirl.
Rosenbusoh* stnteR that the fibre!! al'e pllralld, and that the vertical
tlXis is the same in tho pal'ellt minerul UK in the new one, Also, that in
the case of a twinned crystal the fibres of lll'nlite Htand in twinned
position 1111 oppoKite hiilcb of the twillniug-pllLlle, With l'eglLrll to this
Illttel' Htutl:.'ll1ent. Illl OCCUl'!'ence in one st.octioll Khows quit(' II (liffcl'(mt
~tn.te of aff.lirs. The twinning-plane is distillet enough in the augite,
hut disappeal'b completely on the verge or tho maHte fringe.
Oltemica), Ol~allgel1.-So far aR pl'eRent knowledge goes, the compolli-
tion of uralite is believed to conform nearly to that of actinolite. 'l'he
most prominent change in paSRing from tho original pyroxene is that
corresponding to the diffel'ellce existillg between the two species in
general-that is, an increase in the amount of magnesia and 0. decrease
in that of calcium. Analyses of tIle Bluff minerals are con11'al'ed below
with reswts given in Dana'R "Aystem of Uincl'lIlogy." In the abse.nce
of other medIlI'! of separntin~ the materials thE' following process was
'l'he anal., NCb ot the pOI·Vh., I ~', the llol'ito, Ilnd 1m intel'ull'diate member
OT the "chiRt sf-riel. are l'epputecl hert.! foJ' cOIupaJ'ison.
Porphyry. ~chist. Norite.
l::!i0 2 6;,60 61·00 48·10
A1 2 0 3 12'2!-1 13·66 20·85
Fe 2 O:. 3·15 2·4.3 4·85
FeO {·88 10·83 to·55
VaO 2·1:10 7·35 7-15
MgO 1·08 1·14 3·99
K.O 2·16 0·52 0·63
Na 2 0 5·tH 1·1:10 2·73
Loss on l!HlltiOiI O·lfi 1·20 1·00
99·88 100·33
Hpecltit gravlt~
.. Jiarlrer. A.... Petrology for Stullent... '. 4th I·d. (11101:1). p. 326,
t TeaJl, ,T. J. R ••• s On the MetamorphOAiq (If Dolerite into Hornblende Sohist, ..
IQtla.rl:. J01lrn. Geol. Soc., vo). 41 (188lS). p, 142.
ViIl.D.-(;.eologg of th~ BTllff. 33i
IlJlhtook the rock for I-Iyenite, boulderb ot which \\1:11 u tl.oughl to ue lUUlld
111 the HokollUi Hills, (If Permian age. It iii now known that there iF!
110 I·ock rel!elllbling the nol'ite in thu Hokonlli conglomerates.
III his "Geolog~' of New Zealnnu." recently published, Park makeb
lIO dcfiniw litn.tcment of the age of the norite intrullion, beyond discutll!liug
it 'under hil! Maul1pouri Hye.tcm, \\ hirh mclnu(·1'0 l'oE'l'it!H of C!:unbriull.
Ordovician, ana Silurian agu.
There ill, ill fllct, 110 evidence that U.CCl1l"llte1y fixes the age of thil.!
intrusion. We knm\ definitely that it is ~·utUlger than the intmdecl
rockR-that is, late or post Carboniferous. V l·ry probably the intrusioll
is connecteu with the wideKprelull:lit'vlltiun that in .JUI·llbt.ic times enlargt',l
"New Zealand to continental dimcnsions. This movement resulted in rock-
foltiinp', and the main nwuntnin-t·angeH wel·e formed. The folding Wlib
atl~ul'intl'a with tIlt' il1tl"ll~i(ln IIf ig1l(()ll~ I"O( kH in 'l'":ll"iou . . loC'ulitie. .
ART. XXX\ 1. - TIll' AII.atOtlticaZ Structure (If the Nelr 7,ffl/fl1llf Piperact'at'.
By ANNE F. IRONSIDE, M.A.
IRma be/ore till .IIflJIIlllVlill PMltNJphir"l .'-Ioritl!l. "!flfll .lprll. 1911.J
ANATOl\I[('AL .I!'EATUR.I!lS.
8tpm.
The woody btem ot M(J,Crop~peJ' ucel9UIII show!! the double ring charac-
teristic of the woody Piperan6(Jl'. In a. t,J'ansvel'se section \\-1.' find on the
outRidt> periderm; beneath iE. cmilCltl tissue. A. young stem shows II
distinct epidl'rmis, the ceUIl of wll1('h urI' morl' or II'l:IS oblong in outlinr;
llDIDt>diately below are from 2 to :3 LOWS of celhl, showing tangential clivl-
tlions. It is from th('sc we have the periderm.
Below the epidermis, or periderm, is well-marked colleuchymu.. 6flpC-
f'iltlly thick behind the bundles. De Bary refers to a similar structu)'('
m Piper rllqOllIIJll: "The hypodermal Ill.y61 surrounds the whole stem
as a distinct closed, multiseriate (collenchymatollS) lay(.'r." The collen-
chymatous layer consists of from 8 to 10 layers; narrowing to less layeTfl
with slightly wider elements between the hunrllC's.
Immediately bent-ath this is a ditlCuntinuous rmg of IIcll.lrenchynllttouli
elements, one or two elements wide, abuttine, at places on the vascular
hundles: longitudinal Rt-C'tinnR Rhow these elementR to bt> branched oC(.'Il-
sionally.
Then comes the outer ring of bundles, each bundle separated by a wide
medull8J.'Y ray from its neighbour. The bundles of this ring show secondary
thickening to a marked degree. On the inside of this ring is a. sclerenchy-
matous band l:Ieverallayers thick. broken occasionally between two bundlrfl
(fig, 2).
In the centre of the stem iR medulla. in which is a ling of separate
bundles. In stems ~ in. to 1 in. bJ'oad there are about 10, but tht-
number varies in accol'dance with the size of the stem: 3 to 4 is common
in youug parts, e!!pecially seedlings.
On the outside of each bundlE' both of the outer and often or the inner
ring a1'e much-thickened sclerenchyme.tous elements. Then there is the
inner sclerenchymatous band. De Bary, remarking on the inner scleren-
chymatous band. says.•, The converse case, that a continuous layer (If
sclerenehyma supports the whole inner side or the ring of vascular bundles,
is rare in Dicotylt>dons. This is the case in the woody PilperaceaR-.Arta'flihe,
Cl,Q,vICQ sp:'
In some stems-Ranuflculus repel/s. tor instance-we .find IIClerenchy-
matous elements on both sides of the bundle, representing a sheath. It
is probable that in the Piperaceae these sclercnchymatous elements repre-
sent the bundle-sheath, whioh has become much thickened and extended
on the inner side for mechanical pUl'Poses. There is scierenchyma to the
inside of each 'hundle of the medullary ('ircle, but it does not form a COD-
tinuous l1ru!;; also at the base of the stem, and in young branches coming
from the base, &i! in :fie;. 6.6.. in seedlings the continuity of the ring disappears.
aU which lends to the view that the sclerenchymatous ring is an extended
sbf'80th.
The phloem consists of several layers showing deeply staining sieve-tubes.
Md companion cells; sieve-plates, where cut through, show very nne pits.
InObol'olDI: --.Anatomical Btl Llctllre of .:r .z. Plpt I .1l'('lIC. 34J
The cambium bItOWb ('r.peciall~' \\ell the td.ngentlal diviblom tor \\'ood
a.nd u.J,st formation. The cambml or merismatic legion IE. rontmuollS
r01lud the stem in tho ()utCI' 1m!.!., though it I'> onl~' whele the camblallnyer
pabHes throu~h thE' buudlf' that It give& risl:' TCI wood u.nd bar.t. Between
the huncUeH thf' tangenhal ilivi1'!10U'> incl'ea~e to the medullal)" ray. In no
Inbtance are secondal'Y bundleh found between the primary bundles,
The wood, as is seen. ill longItudinal sections. shows pitted and sodan-
turm vea&els and pittell wood-fiblf's, bebicleb annulal and spIral elements
1)£ the protoxylem. Some of the bundles show the pllmary elements of
the wood to be dtstinctlv ~catterecl: lonu,iiurliu.tl sections and ohliqup
a
b
c "
Young btl~lllb, the base of the btem, and to a le!;s extent an ok!",!' AtE'm, show
the centrol lllE'clulla and medull1.1'Y layt! thickly 8('1, Wltb htell'cll.
En~l('r lldh baiu that m MucroplJler heekeli(l. Piper C'hanira, tho huudlt's
ale in rin~h. tlt(' (lutor bundles joined. III ~~acroplp('r ('uelH1ml tho huntlll'lI
of the ontel IlIl,!, are I:lepa)'at<'u by very wldl' medullJI·.v IU)'h, ,11-1 lllll'udy
btll.ted. 'I'ull~l-'nl iul sectIons l'an also be ('ut ill whidl (he Illl'dullnl'Y l'IIy
If, contmuoUh Illo1,ltt throu~h. A pOJ·tlOn of the intcmodl' 01 II\(' \HlIIdy
stem ""lib bOIled tor SOIDP time in water, and the outol' t li'o,{lll'h 1I(l'lppt'ci off.
The bundlph '>.bov.E'd only an occasional join. t::Iom(' of Hw IllIlldleb could
be sepal'ated tlLloughout almost the whole length of ill<' 1'11-'('(', 2 in. 'l'lU're
is no netwCIT k oj anastomosing. A loin is e.ffp('«>u h,' .1 hlllllCh of VE'l'Y
FIG 6
Fig. O. Bundle from 6tem, Iihowing :!..yieul IICJ.ttol'l'c1: II, phloc'lll; II, lolombium , t, "yielD.
X 150. P. Efldlic1aeri.
Fist. 6 ••Joining among central b\lmllO'o in iowOIIt internode ..CI'dling (bee 1iA. U.\, II I.
X 44. AI. ,:reelBtI1lI.
Peperomia Elldlicllel'i.
Peperomiu Etldllchen is a l'ock-plant, less often an epiphyte. Its stenl
ir.. succulent, its leaves store water. A cross-Foection through the stem
Khows on the outside an cpidel'lDih of nal'rowiHh elements. 'I'h.. ('ortex ir..
'1ucculent, and shows oil-saeR. The pl'esenc(' of oil is hie;hly characteristi<,
of the Piperareae. Hele, ab in Pipnari'ae generally. the oil-sac consisttl
mprely of one of the cellb of the cortex. Strl:Lsburgel' and De Bary both
make reference to the ethereal oil found in Pip<'rfl; .Engler fIILyS mucilagE'
pafllile9 in the epidermis and cortex, but the walls of the paSMges do noi
diff"l from those of the cells arouud: the contents give a pungent odour
to the plant.
Scattered in the ground tissue of P. EJtdlirllen are bundlt''1 showing
RoD arrangement as in many Monocotyledonf>. But in l'epe'l'omia, as Engler
an others point out, the bundle!> hav.. a cambium. In the New Zealand
PepcTOfJlia. however, the activity of thp rambium iF! at a minimunl Thr
amount of lignified tisRue always remains small. There is no dennite
arrangement in this species in rings; whereas in 'lOme Peperomias there are
l'in~. In Peperomia gaUoides thpre are two rin~s of five.
IROl'oI>IDE.-..d.lInfofnlrnl 8fl urfu/f' ot .\' Z PlP!:!1 ilced.e 343
The petiole of P. Eltdlicherz &hoWI> one large central bundle and two
fllnaller ones, one on each side of the central one. The leaf shows an
epidermis consisting of several layers, as seen in many Peperomias. The
mesophyll-cellR are also large. contain but very little chlorophyll, and store
water.
Balle of MacroPIJ)er.
The base of M. ezoelsut/l 1S swollen. and stored with nutrItive material
(fig. 6A). This is I:Ihown even in young bpecimens. The nutritive material
chiefly takes the form of starch, which IS contained in the medulla and
medullary rays. and also in the corte..x.
In the young branch coming from the base of the stem the stem showb
two rIDgs of vascular bundles, as usual. But the sclerenchyma appear&
only on the Inside of each bundle of the outer ring; there is no contInuous
hand of sclerenchyma, though there are occasionally slight indication!> of
Cl, Base of Jf. eue18l1m (x "'Odolf> where fltema have been broken oft); b. young
'!l'edling, half lifp·size.
III region :~ the IUt,101l ot huuill(,b Cli the out('! !llIg \I It II medulhur
bllndle~ is tl aCf'd, but t 111' \\ holt, !14 ~rnrllllllly m.rlgl'd III tht> lI11t' ling t ypic,.J
ot the . . tructllle of t1u' "l1hbUh,tn- rout::. In lJJ #'11'(4'111111.
HII porolill
Thl' det,ttlb 01 tll.llbitlllll 111 the hY}lflcotyl III J/. 1'1('('/"'111 \\l'!I' not .11
all ligid. 'rhpll' me "'1'( plllmulul' traC('H, wluch Il.1t' .lIl'1Lllgt'd on (·it.!wJ'
a C d'
FIe 7
Fljl. 7. 'fra.n..verbe cotyledon, .M. ucelsulII, pan-.w.g throulJh midrib: II. upper: b. low(·r
epidermis; c:, xylem: d, phloeLU. ~'150.
FigA. 8 and 9. Hypocotyl below entry of ootyledon-trs.oes. X 150.
side of the axis in threes. These appear to fulle together, thl' xylt'm . . firKt
of all. A single cotyledonary trace entem from each cotyledon. On
its entry the phloem is found in two masses on either side of the x\ Il'm ;
the elements travel towards the centre,
and lea.ve the protoxylem exal'ch, as in b
Peperomia ampltWi/oZia, P. tithYlllaloides,
and P. mCU'uZosa. ThE' phloem masses
ultimately fuse with the plumular phloem;
as the xylem :£rom the plumular traces
moves also towards the centre a central
plate of xylem is formed: and we find a
typica.l diarch structure. But the plumu- F,r;
!ar xylem may be very small in amount;
1'10 that, as in Piper comiloUum, the central
plate is composed mainly of elemelltb
derived from the cotyledonary 1raCCH.
peliole. young pl.i.nl of
But in one seedling examined the plate '1'L,\UbVCl'8U
.J!. ucel8ttlli: b, dorsa.l; a. VMltraJ.
wall composed mainly of elements derived ....ide; r. bun(Ue. 44.
from the plumular traces, while the mes-
arch elements from the cotyledonary tracet:l decreased in amount. The
I plumular traces showed lignified tissne, and were not wholly meristematic.
as in Piper rorl!ifolium.
Root-Macropiper excelFlwu.
In M. ezrel8UJn th(' roots are adventitioUll. The primary root is very
early lost. On u}'rooting. the plant one finds it characteristically forked.
Very often a seedling bea.r~ only the two cotyledons has lost its primary
root.
lnoYI'oIDr. -Anafo'TIllcal Strllrt'lll'e of .Y Z. Pipel.t<'t'cl.f' 345
FIG II • FIG 12
a
b
'1. St'f!dling ..h'lD" ,,11(lWir'~ "h!'a.th lII.t'rl1oillg iJlLu ,1.1'111; b, J'_iD~ jntu lowl'St int.<'ruUtlt, (If
Sl't"lling-l ',Ii-tr<l(,"~ p~~NiI~ ill ttt unk'r rillg, bUIl(lll'lI }l.IM'Iino, in tl'uro 011i.1'l· rinlL
to Cl'utlY' (diagr,lmmllotic).
convergence liCI:I probably far back ill time, the habit ond IIt,mcture (If
Monocotyledons being adaptations from the dicotyledonous form to their
peculiar conditions of life. That Peperomia iK a reduced ganus, compared
with Xuoropiper: that the connection between Monocotyledons and Di-
cotyledons may perha.ps be shown by an order such at! the Piperaoeae, whert'
the one form, Peperomia, shows a reduction from the MacrO'pVper form, tht'
reduction being in the direction of Monocotyledons. That the Piperaoeae
are relath"ely an a.ncient family. They show a relation or resemblance to
Aroids, more particularly among Monocotyledons; the key to the connec-
tion between Monocotyledons and Dicotyledons ma.y perhaps be found
in a connection between the Piperaceae and .4.raceae.
Hill, who" is in full accord with this view, that Peperoma is a reduced
genus." suggests "that the determining factor which has brought about
TRO'lI'lIDI •. -.tllatumic(/1 Sfructure 01 .Y.Z. PiplJl"uceac. :H'i
the reductIOn may be found in the epiphytic habit of mallY of the forJru. .. :
and later says... Nothing haR been said regarding thc bearing of the
geophilus 'habit exhibited by certain species of Peperomia .,
Macropiper ea:ceZaml1 comell very near a geophilous condition in havlll~
a swollen base stored with food-mateJ.ial: the adventitious roots near th(>
bliHe and for some diHtance from it are also stored with staroh. The plaut
it! sometimes tall, sometimes short and more shrub-like. Peperolilia
EII.dlicheri ooours both as a lithophyte and ohasmophyte, less often as un
epiphyte. Under these conditions it has succulent stems and adventitious
_,/'\3-- (;1\ a
~ G --,
,I ~ r.\
to . rt!iiW \ . L;
~ D ~ <D
~ J
--....(;--_C)
.. _
...
_YJ' '
Lea/.
The lellf-wnation Been in .ill, uCeilJnltb somewhat l'Cl:Ielllblcs that t!eeD
ill such .Al'oids ail Zante4escllw, Arum. ProfetlSor Areschong ball J'emarked
that the lineal' leaves charactel-il:ltic of most bulbous Monocotyledons al"f'
better adapted to push upwardl:l through the l:Ioil than any dicotyledonous
type; and that the bulbous plant seem!! in many l'espects the most highly
specialized form of geophyte. itl:! squat axis and pointed leaves, with broad
!Sheathing base, being clearly adaptQ.tions to geophlIous life.
In M, er.oelsum the leaf is pointed, often Illlarply so, especially in very
young plants. There is always a "heatb to the petiole, which entirely
covert! the young bud or young leaf, und is clearly a protective orga.n. In
P. Enilliooni the leaves are more pointed in the young plant than in tIle
old: but here the epidermis is of several layers. and stores watm'.
Hy'}locotyt.
Although details of transition in the hypocotyl aJ'e not ligid in the
Piperaceae, they may still be of value. A similar type of structure has
been found by different investigators in the Ranllnoulacfae, generally
ack-nowledged to be primitive; in the Labiatae. ('entl'o8permae.. in all
e..'Camined Papavemceae, Oapparidaceae, Relledaceae, f'l'uoilerae.. in PinuG
maritima.. and ill many MonocotyledoDI:l. 'The Rame may perhapl:! in tinle
he shown for futther orders. Ls it not possible, than, that th~ bypo-
<,otyledonary structure Ulay be of phylogenetic valne in I:Ihowiull a line
of cOlluection, or it may ne COuunOll ancestry, for Mon(Jcotylt'uollll !ll.d
Dicotvledons ?
It would seeDl. thOll, th;~t Jla(,l'opi.}J~1' it! II. pl'imitiv(' fOl'D!; Pe]Jf."I'omia
an advance v.ith reduction; ,~hile MOllocotyledonl:l Ulay h:m~ nrillCn 3.8
modifications and reductioDs of the dicotyledonous ty]l(?, ILl:! more speeializ"d
forms, thou~h ell.l'lier in time, perhaps, than the Pipe-ra,cene. The ('O\11'!:!1'
of advl:Lllce i&, however, !!tilJ shown by the relation of Peperotllia to Pir1f'1',
~ hertl the reductions arising in the former in responl:!e to envlrOnmt'ut 0.1 f-
all in the direction of MonocotyledoIl.c!, the monocotylel101It1ulI type most,
closely resembled bein~ the A.rareae.
In the specialization and l'eduction of Peperomia we I*'e ten(lencies
which haye become firmly established in Monocotvledons.
COOKN.-ObIlUt,ationB Oil HnlicOlllin aUiotrulih. ;}4S1
INTROD U(,TION.
(1.) PLAXT-RXBITS.
From April to June underneath the plants of So Ullstl'aLi8 are 8eellll. large
number of Ileedlings in which the plumule has made no appearance. Seed-
liugs were only found where the pla.nt wall qrowiDJ,l in the sand. (For the
development of seedlings, see fig. 1.)
.It'IO. J~·Secdliug" of J:Jalieomiu I!;rU\\,1l ill lu.bur.ltlll~'. (The uUlD.lJllr~ 2, 3. cI!;('" IlpllIltt,·
thP a.gt" of thl.' plaut in wel.'Io.. The lUll' IlIlDlllcrl·d II haJ., thl't't' (~JtylccJllDl>.)
~owiDg on low rocks \\&hheu hv the lied <11(' lound on the tops of them, tlH'
action of the wavel:l helUg too btlllng i.or them to get a footing at the ballC
These plc:l.nts h.we a, h1l'11.,te .11)ppall111C'(', owing to the C'omplete dryin!.(-
up of old blllncheb, thl' !lew 0]1('1-> aU .~lll->lIIg hom the bal:lf.'. The roots tllU
C'losel.v pl't''>l->ecl agc:l.inllt th(' I()('k:-.. :ll1d al(> Ill,Ltted to~et]ler to collect all.,
pill ticles ot sou.
The stemb ot. lllan), pLIIl1h I.\lUwmg (Ill loose shingle are long, thick, and
\\oody, and ,11'1' contitlU('cl bun1(' du..tJ.nce uelow the surfuce, giving rise .It
the nodeb to thick, long. advenutiout! roottl.
A CUrlOUS leatUl"e is shown in bome stems. especially in those on which
,In inflorescence has bt'en bome. The internode nea.rest the stem IS
witherl'd and brown, several adjoim11p, It are green and succulent, then
again there are several bl'own and WIthered internodes, then eithel' the 8ur-
('ulent apex or the witheled ItlID.ll.1Il8 of the inflorescence (see fig. 9, a).
Plants growing on roC'b ..hnost in the sea become rcd, and wither vet)"
boon. Thl' more expost'd ancl dlier, both physically and physio]ogicall~·,
d. situation IS, the more IS the plant inclined to turn red.
ColoUl"ing always begin!. In the internode at the bast' of a branch, and
proceeds upwards to the apex. The internode does not lon~ remam
l'oloured, but soon tUI'Dt! brown: thus there arE' never more than two 01'
three internodes on the same branch coloured a.t the same time. It would
HeelD that the plant turning red is an indication tha.t the chlorenchymu
is about to disappear. This is further borne out by the fact that the coty-
ledons of the Ileedlings growing on a small salt meadow a.t Parematu turn
red before withering. That plants that die down early do not usually tw'n
red shows that the withering is not necessarily preceded by the plaut
turning red. The red colouring is dissolved in the cell-sap, which gives
an acid reaction. It is therefore probably anthocyanin. It is purple in
reflected and red in transmitted light.
At Oriental Bay S. australis grows on Cliffb to ft. IIobove thE' ht'a-Ievel.
and with it largl' quantities of MesenilwyOlfltllemlllll amtrale and Acip1l.ylla
squarrosa. It al&o grows at the edge of the water, but it! not immers('d
even at hi~h tide, althonAh in some other placl's it is.
At Napier the salt meado .... IlOvl'l'ed wit}I i{ (lustralls and plantl-> mell-
tioned a.bove extends t.or Keveral milell, 'l'hill il'l om' of the lew ext.ensi, f.'
..It meadows in Nt'w Zealand. MObt of the meadow is ill tb(' process of
dra.ining, 80 that each year it bl'C(lmf'H (hier, Th(' wilole meadow in ll.utunlll
has a reddillh tinge. The fiuel'>t l'Ipecimenll r {ound at the {'dAC of II lagoon
which r('ceives t'Bch day fre.e.h HupplirR of salt, and il4 t.helterl'd by a Hhingll'-
bank. The bl'9.nche& remain Rucculent. for twelve monfu, lind Hecondary
wood is well developed in th('m "'hile still Ku('culcnt. Tlu.',v are duller in
('olour owing to II coating oj wax IVhich st'I'\·es to oheC'k transpiration,
These plants contain much "alt. Hnd rt'main Kucrulcnt for II long time
when picked, showin!!. that transpiratioll is ~reat]y restricted, and that the
water ~ue has not ~ivf'n 111' its water to the' atmosphel'e. but to th('
green tissues.
At Plimmerton, along the l'ailwa,\' t'mbankment. A~alic:o}'ma fOlms II
thick mat se,era! feet widl' and extenclinll. for Romt' 51l vards, III some
places the plants are very shrubby. '
Salicorma al/lltmlis is a nutt'lICent m bhl'ubby perennial. The stems
are. ~pright or procumbent: the Ul:lua! height i~ 4-8 in. In a sbeltered
poIlltion at the edAe of n J.lgonn thC' plllntl'l R1'f' B-2 ft. in height, and the
bld.nches 11.1 e very 101l~ and thick. Sometimes \vJlen the btt>nlb ,ue ]110-
cllmb('ut they are ab much as 3 ft. long, and ~ive off adventi1hlub rootE.
lit the nodes. Sh0l1 procumbent bl anChtlEo often become very thick and
\\oody, being bometimt>s ovel ~ in. m dl8mettll. The young hranches are
cvlindticdl, green, and bucClllent, owing to the buccuJent leavefl and leat-
b,l..,es which sUl'l"ound them.
The bl'anching is opPobite, In 1I0me plant!:. the brdnches appeal' to bt'
!lIVen off from the main aXIS, jUtlt below the InSet'tlOn of a bl'aJlch Thefle
JlP, in the cases I examined, branches gJven off
[tom the 10weEot node of that branch, the node ri~
being almost indistinguishable. SometimeEo two '\,~
branches arise together, and are sUlTounded by
l('doves (with only slightly de'\'"eloped bases). in (t: ~,\ r •I I
d.
,~,,l:' iJ~\
the form of a "pira] (sep fi~. 9, r). f._~ .... ' 'I,' ,
(. )~:,....
Thomson, (Jheebeman. Laing and Bluck.....ell,
="I~~jj~'\7'
.... ~I. '~
and Miss CroliB, all follo,,,ing ::Iir JOEoeph Hooker. -~:II' ~
"f~~WI
descnbe. Salicorllia austra1IS as .' l eafes&,'
i ' hut I
find that as long as green tissue remains on the
Plant leaves persist, and are short and ('onnate
' "",'
"=r 1
x ' ~J I (/ I
I
!
-the frep portions like decussate scales. At the .. - : ~r~" \
apex they are only a few cells in thickneSEo, ,,}jt~ \
but lowel' down abundant water tissue is pl·esent. ~r-~:~~) )
The veins are not visible till a section is made. >-::5 .~~ I
It is the greatly developed leaf-base which forn1l'l .~=\"'=:lr;1 I
tht''' cortex" referred to by writers on Salicotllia. lY.m I
down the stem the mitotic nuclei are !:Iiill obsCl·...- ':_'\
able at the b~se. Thus this intel'cHlary ~rowing- ~: I
Tht> 'I".u!cui!n ~.\ litem It:- Wl'll i{('velop('rt I found ItO bundle-sheath
pleheut.
The followln!! reiclenceh "ho" thai tbe "tructure of tll(, so-raIled cortl'X
J~ consta.nt throughout tlle gellU!:!.-
Ga.nonp; (3, b) c1('s('ribc,", Salicot'nlU lIeliHlcea ab having .. II branchiul(,
hucru]cnt, practically leafle...s stem . . . posbessing .L compact stele
(with ('ortical system of bundletl replacing thooe of a.bandoned leaves), thic1-
w'J.ter-'4torillg cortex."
Warrui:nl/, (2) sa.ys that In 8alieomia ulllbiguQ the leavel-o stalld out lib
collarl:! round the old!'r parts of the brancht's. which lire sluivelled and
thin. and in structure it differs little from Saliromia herbacea, described by
De Bary.
De Bary (I, b) r.aYh SaZicornlfl herbacea haR ehloroph~Tn til!sue in pl:lh-
bad!' tOIn! in the C01-tex of the stem, and hah short !I('c11y leaveR arranged
in d('('ubsate Pl:Ilr".
Rea8(}1iS /01' ('alling IJt'lIt-ua,(· w/tat (tppeall!, 1(1 {I(' ,wd n((\ b"ell del'cribed IU
•. (10/ 11.'.":'
1. 'I'he vasculal-hullrlle '4ystelU in the cortex-lik~ portion resembles
that of a leaf. The leaf-tra.ce divide", the middle branch behaving nol'-
mally: the two iR.tel'Bl branche", are directed upwards for a short distance,
FtC' :1- -II, LoD.Aitudiml o.,cction of ~ro~inA-point: b. j', "'( lind y of fig. 2
und('r hip;ht'l' power.
and thell L'Wl downward!! in tht' aqueoWl tissue, lIJ.l8.HtomosIDg freely, a.nd
f~ a network. This position of the vascular bundles is comparabl.-
to that in the hasal portion of peltat.e leaves.
2. .Except {or the median bra.nch, the netwotk has no connection wit II
the stem.
3. The hundles end blindly in mesophyll.
4. There is no difil'l6Dce between the palisade-cells in the leaf and l&.lt-
base, and there .is no bl'eak iTl continuity, they being developed in bo1h
no. the ventral mde.
J. The "atN tlSf>Ue of the leat-ba~e (C'oltex) is JD u.ll paltb &ilIUm] to
tIll' mesoplwU of the lea.f, when thIs doef> llOt consist of palisade-celle..
6. Belm, what [ regard as the leaf-basc the f>tem loses its palisade
tJf>e.ue (bce Ii!!.. 3. c). 'l'hi~ il'l a VPlT &hOlt portion of earh internode, lust
the pal;; covered by the leave.. of the next. node below.
7. A lonwtudmal section throll!!;h the ~lOw1ng-p(Jint :"hows the It'olli-
h<l.&e of the Uflual kind, with a very extenf>lVe growlDg-point (sep fig. 2).
Cl'oss-f>ection of the mtemode before leaf-bae.e disappear!> &hoW's:-
(1.) Epide11Ul~, .1 "dllgle layer of cells. the oute!' waUf> of which hay!:'
duveloped a cuticlE'.
(2.) Palisarle palench~""Illa and bcatteH'tl tl.lcheides
(3.) Aqueoub tIf>l'>ue, the mternallimit of which If> the enclodelmi....
(4:.) Portiol1& of fiblO-vahcWal bundleR sca.ttered about In the aqlleOllS
tlKblie. Thebe 1 reg.ud at-. the vascular bunrUf''! of the leaf-base&
(5.) Centlal cylindel or stele with a well-ma.lked peli.cycle. Thlf> Ih
t!le only porn on ll1 this section which can be called ., stem." Tn thiEo al'e
embeddf'd the collateral fibro-va,sculaJ bundles, w]lich in thil> sf'ction ure
..,even in number.
Crosb-bBctwn ot stem below the leaf-hase showf>:-
(1.) A thin-wallf'i1 eridel111ll' of cells much Amaller than those of the
leaf.
(2.) HyPodt'l'Dlis. two cell'!! deep.
(3.) Cortex of thm-walled pal'enchymatouEo cells resembling those of
water tissue, only much smaller. Chloropla~ts are few in number in com-
pal1son with those of the chlorenchrma in the leaf and leaf-base.
(4.) Central cylinder.
Therp is no cuticle. no stomata, no palisade tissue. no scattered
tlacheidt'b. and no fibro-vatlcula]' hundleR e.."\:cept in cenbal cylinder.
'{toU/uta -~tomatd .m~ nUIJ1E"rou ... 011 lhe vell1ral "wial"P, LIllI .th.,,'nt
.Ill the dOIl, ...1 DevelopIDE")lt, as fal' itE> r J1,WC mcl(Je Ol1t. tnkes place ill
the u'mal way, ,md the filc,t slomllta. nre f01m<,d dbonl ihe Hit 1('.11 Irom
the a.pex. Tht' ~uil.1.(l-rell::, .lIe long, fl.lld UH' comp.tl·.ltivl'l:r 11.111()\\ ']'h!'il
w.tJl.., .nc thIck. anJ <It th<' lop nnrl hottom tll<' lIuf..'k,·lWd 1'0111011 oi t1u.'
\",111 bounding the pow ]l](lj('ct.., ill the lOllU of I tl!ig<'. Mirh\llY hl'tW('('lI
lhel!e lld~el! the w.llls .trO llot thwkl'nl'd. and Wll(,ll 11lll!,id jut ont into tltl'
pore, Jolld tllllS Iacilitate its clOl,inl!,. 'I'he guul(l-c(')ls ... 1\' hal! the ep'lh'rnlAl
{ells in heIght. and are sunk benea.th thE' eplll<'lll1lH, t Ite llllU'I' wu.llfoo beine;
In the same plane a., thE' inner walls of ihe ('pillel'm •• l ('('11M.
Warming (2) a.nd ;\Illss Cros., (7) (leQcllbe SlollmtcL as nol beiu~ bunk
lfound that Ghey \yeIE" ~ul1k in every cll'~e. eXC'E'pt m lht' cotvlpllhlll!. wllich
are not referred to bv PitllE"I' Qi thE'S!' Wl'ite1f'l.
FIG. 4. --tJ, Tra.nsvel'llll section 0{ btem 'Illrl'IIuncied by leat and lea.f-b_; b, ([ doDB-
verse section of tra.oheidm.; t. longitudinal section of IItoma at 9.30 a.m. ;
d, longitl1dina.1 section of stoma. at 6 p.m.; '. tl'llJlRVero.e _tion (If epidfrnlis
of cotyledon, llhowinp; oell~ dividine;,
:'1 Seedltings have numerous long delicate hA.u·l:I, the ouiliul' of which i,
often (,l'inkled and wavy. Tbe~' preRl'nt II. (,lU'iOIlR l'eMemb1ance ill f1.l11~1
b
- Roo/',t
FlO. 5.-a, Tra.nsverse lleotion o.f root of an old plant; b, trannverso bection of ;)"'Om1g
branch, near tip; e, transverse 'IeC1:ion of part of inttTllltl fltrnctl1re of rO(lt
of .. seedling.
hyphae, and portions of them. are uften swollen, especially the lip. ~ollle
of them are as much as 2 mm. in len{l,th, while the I'oot it! only 0·25 Dllll.
in diametex. They extend along the root. Irom just hehilld the W'owing-
point to the base o{ tho hypocotyl.
OotyZedot!s.-Stomata occur on the upper and ImH'1' t!lIl'IIlC!eh. liud lLl'l'
placed 1:1.8 in leaf and leaf-base, but are not sunk beneath till' opidermiH.
The guard-cells al'e short an(1 wide. so that ill t!urEnce view tlU' Iltomlltu
appear circular.
Epiderm,is.-Surface view of epidermal cellt! IIhows that t,hw arc wa.v,-
in outline. In IL cross-section the radial wall!! are I:!hOltl'J' thau the tan-
gential, a.nd the outer walls a.re flat. Seedling::! gl'own in a. greenhollt!e
t!howed. ehloropl~ts. few in numbet', in !!Ollle ot the epidel'mal cells.
In cotyledons palisa.de tissue is developed benE'ath the upper epidellllib
only.
SecOllda}'Y (howl],.
Stems of SaliCOrlMa .increase greatly in thicknest! owing to lIecondaJ"Y
growth. some of the older ones being ?i in. in diameter.
The cambium in the original collateral fibro-vascular bundles t!oon
becomes exha.usted, causing the secondru.'Y phloem. and xylem to have an
unusa.al origin. As a rule, with the 6.."l::ceptions noted below, interfascicular
(l8.mbinm. is not formed.
('UOK~:.-Oh'lel'1'(lII()II" Oil ~a]iCOl'llil\ .LI1btralh..
A ('J(.,,'.!-;.(>('tJ',ll ()f J. urall('h bL\: month!> old ,.,howtl jUtlt outl:lide the phloelU
.1 1.1,\"(;'1' of cell.., ,lividerl u ..uuUy br illll'./.('ntiul lInd o('clttllOnal1r b~7 L'adi.ll
\"dls. 'fIIUk tl, cOlllplett' t'.(tra-fl1'4Cicuhn' cambium l'iug j.., formed (&('e fig. 6).
Th ... h,u.; bpl;'ll l'l'('ul'rled by De B.u \' for SalicOl Ilia I/ell~((ce(l, (1. e). The
xdl'lll fOlllll\(l trlllll 1hitl CltmiJiulll ('on:-il>t" of thi('I~-\yrllIerl fibJ OU.M C('IlK,
11111.,11-1 ',\"IIii'll cll't' ~itl1l1t(:d. in i]1'c2ou1m' aill!~'" t]J(' \'e,,,,,~k with vI'ry large
h'llClli>. 'flw phluem ClIllbi"tk or tbin-',-allecl C(·ntl. (o1'ming. ttb usua.l, a
1·~'JJJldt'l· Olltbi,}t' thl\ c·,unbiulll. No l:Iicve-hlhc" ('ouM hI-' rletecierl IlmOll~
rb.,,,,, c(\ll~. which in ,'j,,1/wlllitt (I/lslrlll!h Uh\J)'b contdhl chlorophyll, O.IH1
lUI' delll:lel)' pachd wi1.11 "tawll-!.\laiuK. Kontp of thel-c celib dISintegrate.
r!.wse l'ellldillill~ ht'il'~ 1II1·,ln!l.Pt! ill laclinl J (1\",.,. hl't\\f'l'U whIch lire Jllr~e
Hlrel·('(·llu]U,l· '11<1('('''.
III u,(ltlitioll to the phloem cylinder. there are pilioem itilallll1:l tlCattel'eli
ubout ill the D.uroutoo cells of the xylem, ill each case lying just ollt!lide the
ll1r!.\e vel!IIeitl, from which they arc t!epsl'llted by onl~' a feW' fibl'OIDI cells.
A theory which might account for these phloem itll8.mls is thiJ:!: The
fo)·:mo.tion of the }at!!.c vesl:lels consumes time; while theJ:!e tLre fOIDlWp,.
the ceUs E'ach J:!ide of the I.;roup of vesselK, growing mOl'e quickly, urow oyer.
enclosing a J:!maJl patch of cambium. Thitl giveR rise to the phloem and
loIeveral small fibl'OUtl cells, the latter l:!cpal'8ting the phloem from the YeJ:!88ll:!.
The phloeul islands oonsitlt of thin-walled 01'111:1. which shU\\" great
Iw.ifol'lllity in length. When !.tllined. with Ha'ffranill they Ill'e easily
di ..tinQuisherl. RincE' they tUI'n an 01'ange colour. the Ct>IlR of the xylem
1',." n~"cti(1l1l1.
turniu!!. recL P,lI'tlC'lo!! in the CE'11H of the plJloOlO iblunr!.., l'xlJlhlt Bl'ov. niun
movelllcut!!. The }lhlol'm isluntl& "cre Itt til'~t thought to hE' "it hl'1' phloem
or xvlem plll'enrhrmn ('('UFo. ('ul'eful ill\'('stiglltioll t-lIOWl·t] that in lIomE'
cuseK 11l1tlouhtE'd ..,lr\'I'-t nlll'A are pJ'I:'Kent. UltllOU!!,11 llOIlP W('I'(' fOllllll in tilt'
phlocm <'yltndcl. 'I'hl' \\,,111":1 of OIl' Hieve-tLlbet:., 1111 llKUltl ILI'I' thin and
coloUl'leE.t-, und thl' KIl'\('-plllh'1! slightly ()blique. [coulll nol \\ith f'E'rtninly
obseJ've any pits in the sic\'e-plnH'II, or 111\)' ('a llu".
TIle Tt"aC'/leae.-In thiH llh11l1. 1111 usrutl, the' jll'otoxylt'llI \<t'HI'lI'IB al'e
spiral, Ow bpiral being here from light to lE'ft. The xylpDl of lh(' 1('lIf-traces
consistH entirely of Bpil aL vessels. In additioll to thelle. ilwl'f' al'l' the lal'gl'
vessel!! mentioned ahovc. In mam' cuseA th!',' are ObSE'l'YJ.IJie in tltl' COUrRl'
of fOlIDution. They usually orcur 'in groupb of
two 01' tru'(lI', but. thpre ma,\'
be as many as six. They are the only cellll of the xylem wh1Cll flo 110t
EndtJd.mrls
contain !!bll'ch. Theil' wnllH are greatly thickened and pitted, the 1111>:
differiug from the lIimple pitl:l of fibroull ('ells, althou~h in the walls of the
\"esseh hordering the fibrOl1.R cell!! they ure simple. The pits cliffer frOtU
ol·clinu.1'Y horclered pit!! iu that their Wallll, instead of being dome-shapt'd,
are perpendicular to the middlE' lamella (see :fig. 8. a, b).
Fibl'(lI~R Oells.-Iu this pla.nt the lleeoudl.uy wood COnsit!tR chiefly of
fibrous c('lls. These have gl'eatly thickened wall". and TE':.emble woody
fibres more or lest! closelY in iOl'Dl. There ill no sti'atificatioll 01' l:Itriation
observable. A few arc Septate. The fibrous eells alwavs contrun ahUI1,1-
a.nce of starch-grains, which u.re large und closely pcl.cked. In placeR life"
chloroplasts are present. The cells arc all about the sume lcngth.
The nuclei are large, thosp of the adjoining ('ells bt'ing ill a st!'!l.ight
line. showing there has beeu pl'actically no dispmcE'mcllt of tlt(' cellll due
toelr,lJI'!'"tIOIl. 'rhete all' nUDleroUf. pit!:!. !:!imple. Luth in I'lI.diallilld ohlique
wall" .
•\.1> tn" cr·lll'! alwltYb 1(·muin lh,in!!. rh£'J'I' ito 110 llitieleutlation lllto heart
,1I1(1 sap wood.
No I:!clerenchYlIldtOW! frlll'Ck Wille founu 11.11)"1;'111'1'<'.
Jletlllllfll"Y RfI/fs.-Primary medullary raYI:! are not continued throull,lI
the l"Iecoudlll·.v woor!. uud, Il'i u rule. no cleally di&tiuguished serond.try
meuullal'Y rayt. d.l'tl fOl'men j 1101' is tben' ~nr llecel:!hlty 101' thpm, seein~ tIlat
lIlost of the xylem-cellI:! ILle liying and lUI' connected by pita. Occatoiolhllly
I found a dibtinct medulla.ry 1'ay, Lho celb of which were elongated l'adiaJly.
being three times as long as they wtlre broad. a1l(1 nal'l'ower than the fihroUli
cells, There occur also It number ot bannb of cell... onf' OJ two wide, the
cellb having then radial diameterb blightly longel thd.lI the t ,\Up,entia!.
Thl'HC resemble the medullary rlLYH iu Manonia giveu h~' I-lchleiclPll, wllcl'e
they are very thick-walled, and scarcely to be (lihtill~lliHhe(l from thE'
fibl'OU8 cells of the wood. 111 Salicomiu they are packl·a with bhtl'ch-gutiI11:!.
The result of this slight development of medullul'~' "uyt:. ill that 1h<'y are
nut observab}<, in 10ngitudinlll section.
hrowlI llllrtioll tinalh- fa.lll'o off, 01". it the p1wt 110 ut the \\utel'h edgc, II->
hoon wmllied off, .mrl thc blnllrhes U,llPClll' l(l"rE'n again. 1'lu.'Y arc, hOWI"H'I',
much l:nmlllE'l', luwing IOl"lt ull palisuilc !\nd uqucouI-> tifIHllC, 1'h(' glPI'n
colour ib Ittl(! to c11l01'()plnRt~ in tllC 'P]lCllou('I'lll and tll(, plJlo(,nL cylindcl..
Ohloroplnbtl'o lIl'E' ali.;o pl'esent, althoug]1 to u. Ip'IK ('Ah't1f, 1n Ill(, ftJIl'Oli1l CI>lIt,
(If the wood und III the out(,I' portion of thl' medullu, The> de\'(.'lopm(·nt
IIf CbJOl'Opbyll CUJ'I'eto.poudK to thltt in lIevel'ni (lesert-plttntl-> UU'l1tiOIle>d h)"
Austin (Il) (Kee fig. 8, o).
This withel1ng of tissue is due to the £ormat.iull of cOl'k. The inner
["'yer of pel1cyele, which is now several cells thick, gives rille to pbellogeu.
Cork tisKue and phel10derm are formed in tho usual way. The phellodl.'lm
in tws plant even mote than usual shows great unifot'llrity in the length of
cells. (1hloroplasts are present, and &tal'Ch, wwell is nevel' found to he
t'xhaubted. The granules al'e large]' than those of the phloem cylinder.
FLOWIilREo,
J:jolil'Oruili (lII,~Ij'((li/l fillWCl"1I
from DeCplllllt'r tl) .Vlul'dl. The !iowel'K
al'e "jnd-puIlinnte>d. All the hranchp" JUay hE" fl'l'till'. and 01.'111' I'Imull
1;'10. 9.-11, Branch hho'\\ing witheretl inflorc..t'ellce, twO·lifthR natural HhE': b. Bower.
ing·brnnl'h, two.HUhs natural &izc: o. bud. X 3; d, lloweJ:~. ;{ U; e, tranRven.e
'!eCtion of tlowcr-(l) with one I!to.men, (2) "'ith two; /. perhiRtcnt perianth;
g. hooked ha.irs on tc&ta; 11. ])iNtil: t, pollen-grain: k, longitudinal section of
tlowering.brnnch taken in July, five mOlu.hs befOI.'e ftowcn. are mature.
insignificant flo"el's at their upex: thebe flow-ell:! are placed side by aide in
!he axilll of t~e leaves, Ilnd form all Illruollt complete ling. The leaf.balles
111 the flowerm!(-brancheR al'C de,eloped only to a comparatively small
extent.
The nUlnber of :6.o\"\"e1'p. in each a...'l:il, as 11 lwe, \'aries from five to t<'n ;
uooa.si.ouall.~· there are all man~· as sixt('en, lind then they fOl"1ll a double
row round the bl'8.nch.
('onr{TI. --OIN'l'I'flfwt/. 011 Ralicol'llin Itustl'alil!, 361
GYlloecium.
OlJlefY ilS sll}Jtn'illr, of one cal'}lel, a.nd ovoid, containing onl' ba.sal ana-
tropous ovule.
Sty"lea, two in numbel', are papillose. long and narrow, and tapering
to a point.
362 'l''I'onsactionlJ .
Fluwur I" prottl!l;\·l1ou .... ,Mill tht, loot rll''' h.mg out of toll(· lIlouth hefOll'
t h(:l stamellb.
nl'ic1r iR ovoit.l, ('nn~lhtlll!!, 01 11 thm loohe pt'ti('.up pndoRil'A t he Ref-d.
It iR i+Iot'lt hUl'l'OlllUlcd by tlll' Pt'l'lIibtf'ni IWI iUllt1l.
Tl'.hta il:l I>I·U\\1I. l'Ol'ill<:t'Oll~. lind cuvt.u·ecl \\ illl llOOk('d hJII·!. of differellt
shul'el:l (hel' fig. Il. (/). The innf'1' 1'08 t of titl' ~eed ifl f hill Hili! Inemhl'o nOllll.
Thore il:l no endot'lJlcl'm.
Embl'.II0 h:u' thick ~f'l:Ihy ('otylcd()n~. .lIld nil lI11'umhent, teretf' I'lloicle.
Blln.lO<'RAI'JI Y.
'l'ln~ In.1telil1l which tormb the subject (Jf th.i.s paper W.l~ ('ollC'(;ted hy tb.t'
expeililion 01 New Zealand naturalists which visited tlip KllllU<lfltll" 1<,land"
in Hl07 8 undeJ' the ~uidl.l.l1(:(· or lI6.!>blb. ']'om freu.tle nncl \Y. R. B.
Ohyel·. Fl')ru the labels we iudll,(' ihat it \Val:! l'()llected c·hiefl\, In :\lr.
W. L. \f,dlace. It WUI'> fUI'WUlChl bv 1\Ir. Uhvel' tu the hominion
Mllbelllll, Wellington. and the (Jurator of that iIlHtitutiol1, 1\h.•\.. Hamil-
ton hUb beeu kind ('nough to pillce it In QUI' bandll for nebcl iption.
rrh(' ..lIl&lloph01/a collected cumpl'il'>(' th.irteen species from frY!' bil'u Jlolltb,
iIollll on6 :-.p('ciel:! rrom a DlUm!rull. the introduced domel!tic !l,oat To these
we have added two spc('iPh taken by one of us from II IoIkin of Or8/l'elata
tlPgll'(·ta ~chl., forwlUded by )Ir. Oliver to Mr. A. F. BatN~t Hull, ot
Sytln~y. As lVIl'. Iredn!e'tI notes (lUl(') meutioll abuut thirty hpecif'1! of
biru'.! observed on the group, it il:! obvious that nothing hke a ('onlplete
collection of the Mallopllltgan fnuna wal! ohtained.
Of the Kixteen forml:! .redewl!d, we hnve uesClibed l!ix U1'o new I>pecies,
one a.s II. new variety of a known form, E'i~ht haye been al!Cl'iLJed to alreadv
Known Rpeeiefl. and consideration of one speri!"" haR been df'ff'l'J'ed.
LlOTUErrl.\E.
Menopon "p .
•\.moll.g the pal'uRiteh collected frolU tlu' mm'(ljiclUIl t-tnrliul!, (t;flll')l1ll1
I'lIl!fa";~ LinD.) are hpeC'illH'1U4 of It Menopon silUilnr to HOIll(, \\'1Ii('h W~ hllw
UUlI4t'I\"t'~ collt'ctetl l!'Om the flnnll' host iu NC'w HOUlh WHIcH, but whlt'h
do not agl'C(' with any- Mellopoll dC'h(llibed from tIll' hlul'ling thut Wl' C',111
truce. An it heelUll I!!omewbat improbable tllut whnt ill IlPPUI'ellllr II
common parasite o[ au extl'emely OODllllon !lird ~h()uld 1Iu \"I' f'luded O];M'I-
vation, we have de felTed cOllsidel'ation of this Hpecil'lI until we aJ'e deltllllg
with om Nl'w South Willes materiul, in mder to allow of II fUl'tJU'l' hl>lIl'('h.
Llpeurus languidus Kellogg and Kuwaua (1902, p. 475, pl. 29, fig. 8).
Two female.., and one male of this specIes were taken from Pelago·
droma maril&a La.th. The species wa.& originally described from Galapagos
Island, where it was taken from OCtanuBS ql'ncyili8 and ProceUaria tetk'IJ8
as weI) as on a nurobel' ot obhl'l' hO'lts to whi('h it had obviously
straggled.
Lipeurus exiguus Kellogg and. Kuw..IJla (1\j02, p. 479, pl. 80, fig. 2).
One female from PeZagodroma mal'ina Lath. The species was OrIgin.
d.11v described from Ocea1!ites gramlis flom Galapagos.
ban db. und in each lateral cw've i~ II. blllll.lJ tllbel'clf': In front of mandibleb.
lind intcmal to lateral bands, a more or IOMR oblonl.\ ullcoloured Mpace, con-
taining in it~ centrt' a dark oblong blotch w"ith serrated latm'al marginR:
tl'a J)eeull1c fanly Ktrong, colourcd, and extelldin!.\ to 2nd segID(,llt of antennae:
on dOl'Salllul'facl', at hlll:,e of trabecula.e, an acutely conical pI'oeess set in 0.
pltpilla..l'otlembling thoae found ill Gicbplia: l1utelllllilC Ilhol't, i:!li~htly bent
fl'om2nd segment, '"ith 1bt Sl'l.\llll'nt longebt. then 211<1. then 5th; 3rd and 4th
e<lual and I:Ihortel'; eye not pl'Ominent. with hJ1':ltlE'; occipita} uanru stron!!,.
wry I:Ilightly cOllvcl'!I,ing fl'om the anterio]' ,mgleb of the pl'othorux to the ba~e
of the manflibles; temples dark lUammilla.tE'd, with at lem,t 2 pm.tulated
hail'S about allgle~; occipital signatUl'e conical. the lIpex llleeting a I:!ee-ond
)argE'r cone. with its base along thE' Itlticuilltions of thE' md.lldihleR; bf'tweell
these and the occipitAl ballds. t!ubtl'ian~ulal' lI11c()lonJ'l-·d bpaces; hind
Jllar~ slightl~' sinuoUl-., pl'oiectin~ Ii Jittlc OlL to pt'pth')I'!i\': oel'lophll.~ea1
sciel'ittl and glaudb fail'ly conbpicuotlR (in cleared Hpecimeu): few" hail'fl
are ob!!cl'vahle; b(·yond the ptl!!tulatcd b.lirll alrE'udy mentioned only n
-mort huil' Itt trllbecular an~le and It I:Itout I:Ipinl" behind thE' eye cltn he mane
out, but the hair!! may eal:lily have heen uhrd.ded off the ,ipeeimen.
Pl'othol'ax JlIllf as long all head, IIJl(I hc\'ell-tent~ as wide: with
bJightly com'ex pOl:lteriol' IDllrll,in; II.n~Jell l'ouuded. Ilnd sicLps ronv-erging
nnteriol'ly; lateral IIl·eu.t! brown. with mediall uncoloured t!pace j apparently
1 bpine llond l I:IDlall plll!tulate<l hail' in 1,osterior lIugle. 1\Ietathorax about
half as long as prothurllx, and wider, with sides convexlv diverging from
III hont to prominent pOHtero-latcral anglel:l, theure curving to an almost
straiu,ht hind margin; a spine and a hail' in pOl:!tero-lateral angle; liud
IIopparently a couple of pustulated hall'!> on either I:Iide on :hind mar~ill:
latter with IL strong brown band 011 its free pul'ti(lll: ali, except It median
uncoloured line. of uniform hi'OWll colour: ~tel'llaJ ma.rkings of two pail'S
of VP1'y stl'ong intercoxlIl lines, the pObtel10r pair forming, T-shaped su'ue-
tures with lateral lines external to the (Ioxae.
Abdomeu of 9 segment~. orbicular, widest at 4th begment. and ,vith
las 1 segment Hmali and apparently bifid; the first 7 I'cgmenttl with
triangular lateral blotch~, elongated inwllrclB; blotches of Rt]1 Re~ment
quudl'ilatl'TlLl; 11th with only small I:!uffused hlotch; 1 to ~ pm.t ull.Lted
hairs III pOKterior angIeR of ~pl-,rmentR. and It few hnirs ~tl()ng hmd mn.r!l:illl:l.
Illl well ItS !lovel'al on thl' uncolourcd bp8c'e ill (,l'ntre of abdomen; hut no
eJ,CCUl'ato <ll'l:Icl'iptio11 'If ille !lumber a.nd clillpol'lilion of hairs is IlosHihle,
owing io the condition of the IIpecimen; on the ventrul I'III1'fuo(', hmall
blotches COI'l el!ponding in shape to tllOl:!n of ~E'j!.lUcnt 8; 110 genitalia oll,*,n"-
able.
Totullpugth, J·38 llllll.; brcadth 0·60 lllDl, ; head. 0,-1:2 mm. by 0·30 mm.
Lt is Ull£oi'turutte that we have not at our disposal better material of
this speoies, which we have included in the genlls PlIilopfel'tlB. hut whil'h
presents 1:I0lne Ililinitiea with Giebeliu,. Fl'om this genu!!. howe\"er, it iM
clearly ma.rked off by thE' lmiforro chitinizJ.tion of the dorsal t!Uliace of
the clypeus, and the ab~ence of the ventral memb1'8,nous :flap. When
mOl'S material ill available it ma~' be necest!lIry to make the speciel:! the type
of a new genus.
065 rom bl) th ..t b( ~1(.lI ... bem~ she,htl) lom,el It l~ plOp01 bOllateh
a.p1,...m ...i
much bLOadel Head me.l'lll"llllllt'> al(, o 7'1 !Urn h'i 0 7~ mm 1, a~.J.Ill'lt
060 mm b, (1")3 mm .\.llt(,ll ,I band.., lIL plodu('Po un, uti., turth"l
Illn. elld III I 10 umk cl l'0 ... tCIlOI
plOlon~ 1WOlI 1'h(' n UIO\\ b.mu
on the hm.l 1ll 11..,111 ... of tIll nT..,t
'>1'1. III III mOIl' ul.,hnct than
tho'le mdIc..J.tcd III Kello~~ ':> U.,LUL
of P jllill/~'IOH and mmeo\ ,,[
He olllv ,en uill1owl) mt.el
I uptec1 medtanh 1'he la.teral
IJl0ifhe., OD th" fin'll ,>pg;ment
of Ihl' ludom('n \II 'lot '>0 pl0-
nonn('ed, 101 Ib 1h(' abuomell so
h ... tlllctl) tm om II \ d 'l'hPl e 'tIl'
,1"'0 a fc'I'I mmOl ilifillPU(,(,'1 HI
the nnmh('J IUd Ul lqull('nt of
hJl1'
We hI H b..,uwJ. we UbpO...1
1'1 b 11(111 01thl c1utlllOtlC, m ukm"..
PI 110) ft
on thl' 'l'nh'll ~t1lh(.c ot tIll'
I If' I [,h,fol terri' ,wll" I
VI ntl"ll lbdomcll 'I" ,~ell tc, tIle ~eDlt.J.1
'tppalltub whIch 1<, e\.tJaOldl
n<l.uh I) .,( .tlld wmple"\., dl d (outmut''> thtoue,ll 'U\. l.bodmtnal ">Cgment.,
.a.r. In P /U'if/IJIObltb
One male from Oha1ad Ui' donun/(1lb l\Iull We h'].,e named th\
l'peCles III complIment to Ml '" L Wil.llacp to whom 11'1 dl'l<..OvelY 18 due
Il:l. 9
Ph,lope" u, arnult'lH
III 7
1'/11101 It, '" fit mat,,,
\\ ht..lC the ((,uvelgm6 &ld~ meet thc '>tlc),lghi. m..ucln d.le ty;O lIttle iolds
In the rhlhuoub bOldm ~ "'lllgle helu In the angle A fcl.uly strong
htt'lell bcl.lld {'\.tenlim..1. along mOll' than one-thud of lund lll.llgm :Meta-
thola"t WUH'\']V dlVelgent to a truncated Po&telo-Iatelal dongle, WIth 3
"bong hcl.l1... hmd Illdl~ln obtu&ely lounded, btlonglv plO]ectmg on to
d.bdomen VI lth do "Ellie" ot a.hout 20 pustulated haul> ill addJ.tlOn to the
b d.lrea.dy mentulIlt'u, \\Ith ..bOlt dalk a.ntelo-latelal band'.!
7'/ '11/"11 f/o/I'
,md nSllO\H'I, thl' prothol'ax diitt'lent in bh ... pe. ann there !lll.' utJlel mmor
cliffe' ence....
De.W·I. }l1/()/I oj Fl'mnle.-Hearl lungllr thJD hl'oad, with elonAate (u1l-
cuvP-biderl clypeub, evenly lOunrlecl tempoldl lobE"'. itnd slightly ronCd'"
hmd lDc:Ll~m, with yelY ",mall mew.lll IOllnded }llOjection on to pl'othorax :
markinL,\s lllllC]l tilt> ..,ame a'l 111 tIll' lnst bpf'cic.." bcl.\"e tlmt the buture is mudl
more dh,tt'lrt. th(· acnmina.te POillt of the g;gnatllre ib fOlked antelioIl\·,
the I1lllt'l .1utt-uUJ,1 bands do Jlot reach ..0 tal to! wllld, and the occipital
band.., olIO \\ell defined: prothor.tx Wlth ..ide., converglllg al1teriorly, and
hind maL!~ill ..,bL\htly convex; llll'tdthorax with pm,tero-latClal anglel:l n()t
t:unc.l1ed, l111t acute. and With deeper and m.)1e J,D!!ulated projection on
to ahdom!'ll: hinrl Dltll!!,in With about 20 pUbtulatecl 1lairb not 26.
AbdoIDf'll mOLl' loundly ovitl, wltll Jabt fooOgment& llOt 100 much plO-
clucen. tI'!tnWClhl' bandh not so nil:ltllll.!tl) angulated at apex. and
I'loduc p c1 hutl1el I11WcLld... le.tYlIw, a lDuch CJmaller clear "'pare; the
l'\1b1 ulat\!c1 h.lil'':> of the ",eu,ment., geut'rall~' Ic>" 1'1 in numher. rJ,ugillg
Jrom 6 on '>t'!!,Ulent 1, throU~]1 from 12 to 11- on the in1pl'vemng
"'1!~mel1t". to 1- on begment i: 1 to 3 haiI'Il ill the pObteriol angle..,
pxcept th(' first, which if, without hail'S: glmital blotch closely l'ef>emblin",
that or the Jnllt bpecleb, and more eatilly compated on ft"ference to the
fio,uleK.
l,plIgt[1 1'~17 mIll.; 1)) eadth, o·~n mm.: head, 0·66 Illlll by 0·60 Dllll.
TRICHODECTIDAE.
Trichodectes climax Nitzsch, in Giebel (1874, p. 58, pI. 20, fill. 2).
M.n.n~· s})ecimeDR of both bexf'S were obtained from the il1troduc~d
clome':ltic I!.oat (OUPIQ liil'CIIR Linn.).
The typeb of the new species a.nd v.ll'let)· del:lcribed ill thit. paper aN
cleposited in the Dominion Museum, Wellington. New Zealanrl. Wherf'
there has b('en sufficient material, cotypes have been retained ifl Oul OWll
(·ollection.
All the ti.l;uI·es ill th~ Rl1.icle hnvt" been du\'l<u with a CUm('I',1 lucida •
•II1J have b('('n equally magnified,
LITERATURE LlbT.
1812. DenllY, .. MOllO!!.l'u.phia Atiollhu01UDl Blitalllllae." .Londoll.
11l10. hednle. c. Bird Lif(.> 011 11l!' Ke11l1ullec Ismuns." c'The Emu."
vol. lO, pp. 2-16.
1896. Kl'Ull~l!.. "New J\'IallophakJ.;' vol 1. (Cuntl'ibutions to Biology
from the Hopkins Seaside Lnb01atory, iv.) Palo .Alto, Oal.
18~H. Kcllol!,~ nnn ClIRpman. ··"N(.>w Mallopha~Il;' vol. 3. (Oolltl'ibutio:n.e.
to Biolog,v from the Hopkins Seabidp Labol'atory." xix). SlIn Fraurisco.
1092. Kellogg aud Kuwan..t. "lJul1ophaQ,a from Birdl> of Galapagos."
Pl'oc. "'usb. Acad Sci., vol. .J., p. 137.
1!)08. Kellogq. Mallophuga ill Wvtsman'" •• Genera I~ccol'1lm."
Brul'Mlb.
18;4. Nitz..ch. In Giebel. "IIlI:Iecta Epizoa." Leillzig,
1880. Piag!'i. .. Let.. Pkliculines, Essai MOllographique." Leyden
1885. - - - - - - Supplement. Leydell
1874. W(,'1i wool]. .. Thesaurus entomologicus oxoniellsir.:'
374- Tl'aIlBactioll~.
b. a }ll'eVlOu... l).t}n' I ('L\!tlll:l. N.Z. hll:lt., vul. 43, l!lll, 1" ::i~2) I tll>Kl'rlbed
the general ..Il1.::t1omr of HlpltOlIai'ia ouliqualll. the common limpot·likl>
marine Gastropon. WhllSO n.flinitios II.I'C still l1ud('('icled; to)', \vhile !:Iomc'
8uthoJ'itieK inclinf' to tlll' view that it i!:l II. modifien OpisthobJ'anch, oth('r!:l,
and pel'hapl:! tIlt' llliljol'it~·, regard il all a Pulmonate. [no\\ sUJlplement
that .lrticlc hy !.\iyill!.\ ll1'l't' an ac('ount flf the \'nl'l('ulal' bVHtell1. which 'Yak
omitted purpol'>t'l~- trom thll t paper.
The adnpt:1 tioJ1 (It l'IlpllOlIOI'ia to Ii life along thu I:It'u·hhul·C, wIler£.' lit
timel:! it in lll'lolY ",,'«"'. u1 orlll'!' timcK nbovp. iN w('11 illu",tratell by iib
.. clipnoan ., t'llcll·II(·tCl', hJ.villl.\. thl:lt is, both a hmg and II gill, each with
11s OW!l illd~p(,lld~'llt Lluod.'1upply. If, ttb J:o(!tIlllb Lho mOH' I,>l'Oh,lhll', it ib
II PIJmOn.lte wllich h:tb left itl:! terrel:!tl'iul hOnl(' and wn1ur"d buck iuto
tIle ol'iAIDui hahitat of the (hu,tropod molluscs, we must look Clll the ~ill
,IS a nt-VI htl'udllll', anulogoub to thol:!~ of Kuch fOl'lllt! all the Proxobl'ltllob,
Patella (limp('tll), mid thC' OJJi~tllOhru.lIch, Plellropllyl1idi(l, wlii('h hall uriseJI
III relation to thih lH'W mudl' (If life--tbat is, it is ttll ,. auaptive" gill, as
In these fOl'ms. (ts structLU,(, it! not tbllt of II typical ('tellidium, as 1
pointed out ill my pl'tlvioUK ..tl'tiC'It'. thollp,h it hilS SOUlO Tel:!emhllLnce til
('ertain gills amuu9,st the Opit.thohrall('hK. We already know a ffl\V Pul-
monates \vhich have taken til It1l aquatic life in whieh ,I ~ill is pl'el:!ent.
Thus, in Isidora. Pl'otal1cylllll. lIud utlt!"]1! thtll'o ib II ll111g1~·f()]dt'd IUIJlma;
but in SipllOlZaria. Itt! I have del:!('ribcu Jt in my }ll't>viOlll' ul'tid... (p. uR5).
there ill a lIenell of ill11epl"ndcnt laminll!.' a l'ulllged III U hl'llIil'iH'lt' 1'0111111 t h(,
ruantlll·eauty, ('t1.cll lamina bl>al'lng h('comicll',Y Iumillut', xu t hut it il'> mudl
lIlorll ('omplex thun tht> gill in thc IIhov(' PUImOIld.teK.
I,uug Te~~tl'dh thelle pulmona te ~illR elK lJl'Ohably t·tt'liidiu, t hough Pel·
l:Ieneel' and ()thel'l" hola them to he " uduptiyc .. 1/,i1lH. Ou the> ulllt'r Mnd.
HUppO&ng Ibtlt SlpliOllaritl is Illl Opil:ltlwbl'lIl\('h .....ltil·h hUH dllnc uuIt,
1
too remain out uf WtltOl' fo), a consid(,!'llhh' IKllli()1l (If t'1H'h day. lind lu
. hl'ea.the nil' tiut-ill!/, this pl'l'iod, \\'1" lnl1lot IIUPP0!>I.' blllJl it iH Iht, hwg thlll
il:! the new tIliug; and Illthougb, "') fal ali 1 lUll mVUI'U, no OpiHtllObran('h
has beun found to have u lung, yet aUHIIl!!,Kt the PrOl-l(Jlmmcht! Bovcl'l\1
genera, belonging to diffcl'ent families, 1lI1\'l' cle\,elolwd II lll'tw()I'k or blood-
vessels on the mll.ntll'·l'ooI; hO that thtl l'lwil y al'ts lUI II lltug. allll the
genrru. cun be 81'l'all~!.'d in beriel:!. in which toIle ctt'luclilllU 1l1:lIdulI]]Y Jem'eases
in siz(' and imPOltalll'C till it ramallis as 1\ mCle vest4lc (aK ill Oeritkidea).
Hence, wbatevel' \;ew ill taken l:III to tIle affinities of SipllOllaria, it
:remains n.n extl'cmely interesting form to biologillts, hllviug either II.n
adapti'\"e IUDI/, 01" all adaptive gill.
THE HEART.
The heart liel:! ill the periCI.Lrdium on the left !:Iide of tIle clol't!al SurntCIl
of the body, just ill front of the middle or the length. The position of
the heart, as seen in the animal after the shell has been removed, is shown
in fig. 7 of plate 28 of my former article. The walls of the perica.rdhlm
l!o·r~·Hm,I .. -T'(l8rlll(l1' S!lstl'1n of Kiph(lllll.llU "bliqullta ::n:,
al'o furmed by II Lhin tou!/,h Lrdll"'p~rellt mC'lllbl'ckne. The jwurr, (lr. IlI!1lJI.
i., two-rhambol'P(l, tb(' ventride bein!.( situ:t:ed to the left 01' the nuricle,
iLb ttpex poillting to the leU unu rlownw,Il'U'I, "'hile the auride lies
almost horizolltd.ll}' and tl',IlIhVl'lboly to t1l!~ hotly. TIle wall., of the
,LIIl'icle :Lrc thill, h',LIlbL'Ul'Ont, Mid extreulC'ly dt:'[ie,lle. \\'hell this chu.mber
il'l di'ltf'llri,·tl wiill hlolld rhp hluiHh tin!!!' of the lultel' mllv be lloticE'd.
:FIl...l.
Disseot.ion of thE' a.nima.l, showilUJ the chief arteries; ,:2. .i)orsa.l W.lU of body (mantll':
with heart and porioardium turned to the left, exposing the ventralsurfaoe of tho'
hoa.rt. II., anulI: ao., o.orta; b.lt .. superior buooal a.rtBl'Y; c.a., cephalie aorta :
d.y., tligestive gland: t., edge of loot; f.b.c., :floor of body·oa.vity and upper
II1lrfaoe of foot; gn.II., g~ita.l artery:: I!,a., ~a.stric artery ~ g.b.a., ga.stro-buc~1l1
artery; h.y., herma.phrodite gla.llU: t.. mttlStino; p.a., penbl. a.:rtery: po, penlI':
pro., prostate; o.n., viscer,d o,')rta; !'.. velltriclll; RIlL., spermothOO8; at., stoma!'h
r .. rectum: IL'.n., artery to body·wall; lII.t•• mantI!' turned aside.
\'entricle is ot more definite and constant slMpe HHm tho UUl"ic'll': f1'olt!
above it is broud at the- fimicllial' end and tttp<'11> to j,1I1:' oth('l' c'xtlel1litr,
"here it ends in s hJUllL poillt. .\.ftc'l· 1'c111O"ul of th(' lluri<"lC' til(' vuh-('I>
C'I1U be been PUI fly 0P('ll. lea.villA tIl(' Kilt-Iii\{' Ol'ifif"I' tlrroll!.\h \\ hidl the
blood paRRe!' to tIl(' v(,lIt,dde. Wlu'lI tlu' v(>lltri\'l(' C'lIl1tl"Urt~, .JII' lip" 01
tlles<' valves ('Ollll' tOW'ilwr .lIId r1()I>I' til(' PaHfI.I!!,I'. 'I'hill IlC'tioll "1111 b(,
I\ell seen hy filling til(' ('huIllIlI'I' with \\utC'I' uncI !!,l:'lItly "(llIc'('zing It. 'rill'
walls of tho Yt'lltri('il' ,1I'C' IItout tlll(imUHCnhll", Illlcl Imlll tho (;'XtC11(11' ('UII Ill'
seen the multitude oI IIIuKculuI' fibl't'K l'lmlll1l4 in VIll·.,rillg dil'ect imlh III
them. Most of tho :6.hrC'H ("onve1'ge 011 <1, I'tout l'in&/, COI'1I1e-d I'oulld the' eXit
of the Iloria: their otllel' (;'ndR. often 1)I"I111chl'o., Ill'!, nttn('heo. to the "nil" .
•\.RTERl.l!lS.
The aorta leave!:! the vellt1'icll' 11ell1: the miudll' of ils vellt rill si(lc'. ,mel,
piercing the pericardi.:tl wall. tu which it iK fuml)' uttlt<'hed. tllkeR II C'OIll'HI'
.. Imost directly transvolsely towltl'dl! the
l'i~ht lI.ud Ii little backwaru,." J:lIiKSill~ ho-
tween the anterior lobes of the rligestiw r--=
~Jand till it ]'eachetl thc intestin('. At tlliK oe ---I-
point it bifurcatel'l, giving rise to the cephu-
li(' aorta (fig. I, c.a.), supplying the ante-riOl
I bn
I'egiou of the hody,ou.nd the visceral aOltu
(t'.a,), supplying the posterior Ol'g!ll1/:!. Thill
PIiSSes backwards among the lobes ()f the
di~estive gland, and divides into three main
IJranches, which canoy blood to that or~an,
to the hermaphrodite glaud, intestine, und
pClsteri,Ol end of the stomach. For the pur-
110Be of studying the distribution of the
arteries I injected coloured •. starch injec-
till]l" into the ventlicle, and was successful
in obtaining a beautiful prepara~iou showiug
t'\'eu the smaller branche!!, but could not
II liord time. nOl' c'l.i.d I considel' it worth
while, to wOl'k out the distribution of the
\'iHCeral a01ia in ful'thf'f demil.
The cephalic aOl'hl paNKes to tho right
m-er tlle dorsul IUll'face of the viRI'ela. Itnd
1'0meR in rontSt'! with the body-wall (i.e.,
thl! fiool' of 1he pullial chamllm') jUl'lt IJeJow
the kidney. ('untilluillg itK ('()Urse to thC'
light, it curves forwlLluH Lill it l'euc1Jel:l tho b'w.2.
1'Il1ermotliecal dllut, in nu.nt of wllich it .\rtl'ricb ()II tho .\l'ntral ~i,ll' c,f
puseH downwal-d.!! and fOl'walUs on to the Ktomach; A 4. b.II., I'oIlpcriur
floor of the hody-cadty, Just before reach- buccal artary; g.b.n.. ga.stru-
buccal ariel'Y ; g.(/., SlloIItric
ing this point it givel:l off the genital 8.1'tCry artery; oe., oesophagus: bl.,
(un.a.), which, paldhillg backwards, lIupported htllma('h.
by a thin membl'anc, gives rise to the vessels
of the spermotheca and genital duct. The cephalic aorta now rUJll:l fOJ'Wal-ds
for a short distance on the floor of the body-cavity, and after giving off
11 vessel (w.a.) which disappears into the muscles of the light body-wall
a.nd foot it turns to the left and reaches the nCl'Ve-collo.r, through which
it passes belo\\" the oesophagus.
('O'IUI'I! 1H,1,.-f'({'('llflll S!l~felll of Hiplion:11'ill ohliqulltn. 377
.\. UtI!' ht'ior(' l'E:'t1chim.!, tIlls )lomt the cephalic aorta' gives rille to a
1,I1!1,t' hr,tl1Ch--t hc !ldtltl'o-buccal al'telY (g.b.a.), which paSlle8 to the lcft
,tiun!! tl)(' flf)(ll' of the> h()d~--('aYlt\-. reaching the stomach on the ventJ'etl
~;a 'lulface a littlo behind the point wherl'
the oesophal.\us (fig. 2) entel'll. Here
It an onCl' bifUlcll.h~b. one branch-
the superior buccal al1iery (b,a.)-going
forward!! to the buccal maRS. and the
other-the gal\tric artery (g,a.)-bttck-
wal'diI along the ventral wall of tile
t-tomach.
The l:lllperior buccal artery pa&':Ietl
forwardtl al> a large ve!.!Rel alon!!, tho
left latero-ventral l:Iide of the oesCl-
phagus, and turns up "ith the iattcJ:
on to the dorsal surface of the buccal
matis, giving off small vessell:! to thl'
j"II" a.
()E"sophagul.'l 011 the way. Olll'eachiug
Al'totieb "ullplyin~ the uUI'Ml burio.('P lit
hu('elw lila",,; " ahclIlt 4. A, right a point a little behind the buccal
lJranch of bU('cal .lrtl'j'Y, whieh JlM''''~~ ganglia it lJifllrcatct! into right and
helow the OClAoph.t.gIlN; B, left branch left In'anelell, ..1 and B respcctivel~'
of bu('ca.l artery: b.g" bu('('al ganu;lioll: (fi~. 3). Branch·..:\. passes under the
/1.11., I:IUperior bUl'calartl'lY; r.N., ('nu ot
rBlhtlar foI1('; lW .. right hIllivary artery: oesophagUtl to the right side, and gives
•• , tI&livnry o.iaucl; 01'., Ol'",uflhaAll~. • off the artery which supplies the
1:!I!.1ivary gland, It then passes on to
the latero-Uol'lld tlurface of thE" C1el:!ophagus and sends out a branch to the
right bucco.l ganglion. Tlles(' Aall~lia me well I!tlppli~d with blood-vessels,
and Wh~ll the latter It1'l' well injectprt appeal' to be encased in an envelopc
of tlmaU arteriel:!. An al1;el'Y
o.lso fUns on to thc COlll.mi.S-
I:!ure connectill~ thel:!e ganglia.
Beyond thi8 point bl'ILnch A
paasel.'l to the IInterior end oj
oesophagus. and filJ.ally dipt.
into the antel'iOl' lllUKcleK ot
the bu('c'lll ])lttK~. Bl'Ullrh B
of the blllleriol' llUccul urt.el'Y f
lUl.K a liimilal' diKtrlhlltion (Ill I'
tIll' It'fl I:!.i(ic.
ThUll thiK JmUlch o[ till' 1-
~allJj]'o - buccnl Itl'tt'),~'. which
I have ctlllt>d tiLl' buperiol' Artt"lil'l:! un th(' fl(lor IIf the hUlly hU]Jpl~-inu the hNtl
buccal, to di"tinguit!ll it from elml fout., 0.9 seen wht'n tho buccal malll\ ih ~eve:rerl:
another \'('1'1111.'1 (the inferior ,2. a. 8. anti ('. the three main arteri.el. supply·
buceal artel'~'), suppIiel:! t ht' i~ingl'lIt the heae!: B hi th(' infPriur bucl'al artt'l'Y. and
bhurt; r.R., eephali{' aOlt.a. (which is much
oesophagul!, bHlivll.l'Y glauds. ...wullt'll at the point whM'~ it branchp!! as it passeq
buccal go.ngliu, u.nd Kome of betwepn the pedal ganglia): (J,b.a., gastro-buccal
the dOl'l!al mUflcleK of the artery: ,!," month (hucco.l mal!b beinl! remcT'etl):
b •1 p.I/.. ]Ienial Ill'tny; pil.n., pedal Rrtl"ry: piJ.g.,
ucca masH. flE'dal ganglion.
The gastric artery (fig. 2)
ta.ketl II. backward course 011 the ventral side of till' tltolllach. L,ri.ving off
branches too the ventral surface of the stoma«:h; th~ extrt'me posterior of
tbil:! organ, however, is supplipcl by a branch of 1bf' yisceraillorta.
P.etUl'nmg to The cc:phalic .lOrtct. whwh VI (" tl'llced .l hon' Ill' fnr I'~
the llerve-c~IlHr, we find t11ut tlb it PUHflCIi 1)(·twt'l'n tIlt', fJcchLI gllnglia It
b..~ktl up mto It num]Jt'L' (If lnl'ge Vl'bI>Cj~, 1>01110 of wllb'u 1lI11 fO]\\"I",I"
(111 tIle 1100J' 01 the uody-ul\'it)", bUPJllyill~ illt! O1'!!,.IIIH .1Ilt! wallH of 1111'1
1e~ioll, lind "OIlle palos h.wkwul'w" Hllpplyill!.\ I h,' fpol (fi!\, I).
RUlInuJ!!, 101'Wul'clb we have thr('(' IIH1Il1 mh·l'J('h-.\, B, l' (Ii!!" I),
'l'he :til lot 01 tlwtle, .\, l:ll1pplieH thl' Vl'ut nl wall nf 1114' 1ll'lIci. )Il'llIh, :1111 I
musules round the mouth. It would UPP('" I' from l"lpt,11I1 1I111t0l1'fI li!!.III'C'
that the pellial art("ry Willi milltaken by hilll fol' the VII,.. (It·fC·I'I'IIH, ...
S, which 1 tt'llll the infCLior buec'al II rtI'I'Y, a1'iRNI to t hI' It'ft 01 .\,
goas directly iuto tbe bucclIl nlflSI:I 011 itl:! V('lltl'HI mdo, and Hupphl:'t-.
alm<...t the whole of this orgall. To the l{'ft of B u~ain if! th{' bra.nch
C, which il:l the largel:!i of the tlu'{'t': very ll{"U' itK origin it bifurcate!>
into right and left branches, th(' Inttel COl'l'(,KpOlldhl~ to .\ of the righ1
tUde. The right lm!.nch rUllfi forwfl]'ds lor ,~ flhort clillt,allce, where It
bifurCJ.tcs, the rami ent{'ring thE' n!lu'Icular vl'ntrol WIiU of t,he heau,
Runuing backwards from the cephaliC' aorto Itt thiK point :Lre ~o
lal'ge arterie!'!, right om1 It-ft, ",hi{'lt rliflOPPC'lIl' Imll)Jl\\ thp TIlnfl('hm of th('
foot (pd.a.),
VJtJINI:!.
The distrilmtiol1 uf till' '>'!linK iii illnsi,lll.ted ill my ptevious ILrticle
('I'ra11l1. N.Z. !net., vol. 43. pI. 28, fig, j, and pI. 29, fig. 2).
[n the foot and body-wnlh'l al'e llUmel"tHllI large bloorl-\>f-'sf!els 01'
spl,c'es, and into these, all far all I have been able to make out, most of
the blood eventually makes its wily. Part oI thill Mood il:l collected by
a large \-ei11 which l'llDe vertically up in the left body-wall nelu' the end
of the gill; on reaching the dorsa.l surface it bifurcates, one branch
-posterior renal vein-being distributed to the kidney, and the other-
the afferent branchial vein-running round the pOl'ltel'ioJ' border of the
gill, and distributing blood to the giU-lam{'Uae.
Near the rellpiratory ormC'e it Aivel:! off u. large brandl-the anterior
renal vein- which runs between the gill-lamellae on to the kidney doBl'
to the renal papilla, a proceSij of which l:Iurroullds the VE'lIStll betwel'll it!,
origin and the kidney. Th.il:l vessel has been ligul'C'd ligMl}", 1\1:1 it lie!o
deeper than the other vessels, and itA reference-lino lutS IIE'en mispla('cd
In the figttre (vol. 4.3. p). 28, :fig. 7). (lo1Ulected with Ute' afiorcnt
braJl('hial vein donA itl! whole length that'(' ILlt' a lo.rgp uumht·l' of palliul
veS.8E:'ls. The blood pal:!Ses from thiK "ein throulI,ll tlw gill, wIlel'c it i"
cl.tll·u,ted, into tIll' efier('nt l)l'anchiuJ. vein, which runK l'oUll(l the Imtl'rior
margin of the gill. This vein retul'US blood to the aUl'icle> purtly hy u
\'ein, leaving it half-way along the gill and crossing the kirln('y. fJ'om
which it receives several sIlUIll vessell:l, lIud partly by the efiel'Cut
pulmc)o.ary vein, which it joins at the right end of the gill. 'J'bis latter
\"61:11:1('1 receives blood from the efferent '\"'essels of the lung and entertl the
uUlicle together with the efferent vessel crol!l:lin~ the kidney.
Another large vein receiving blood .from the body generally is the
afferent pulmonary vein, which emerges from the body-wall just in front
of the pericardium, a.nd l'UDS round the anterior margin of the lung, giving
rise to the a1ferent veSl:lem of the lung. The blood passes throup,h these.
IITld reaches the t!fferent vessels, which oarry it to the efferent pulmo~'
• Tranll. N.Z. [mit" vol. IS, pI. 17. fill. "8. I'.
CO'l"I'IU'.',I,. - rasclIlor 8!flltnll 01 I'lIphonlU'iu nbli(plata. :}7H
IN the folh",illg liilt the llUllleb Hnd 11U1I1OO18 (315i-:116::1) of new species
uf B,lJrrllidflf' I'CCU1·tlccl ill Bullctin Xu, 2 of the '~ew Zl'uland Institute
U1'I' prelh.etl sn UK to Sl1ccccrl tIl!' Inst number in Bulletill No, 1. Tbis
\,'as l1eceHI!al'~' to mnlu' till' 1I11111hcl'ing con Sl'('ut ivc. ana to prevent tht!~
I!PCC'it'H bping IIverlookc',1.
Within the llr~l.4ellt real' (l!HO) descriptiollb of hevcn lIew genera and
] 89 "pecics of ~ l'" Zenlllll d buutles have been IJl'epured. 'fo these are
alltlt'd. in theil' jll'Opl'l' plllC'l;'<';, eight spccies or belaphi(/(lt' published in
the nerlllan langung(' h~' HUIT Hl'ittcl', of Yienna.
rrhi" lllwxpcufl'rUr hll'ge ll!ltlition, to 0. great extent, il:! the result
01' UlI.plOl'ati0l11:l of rliJTpl'Plit Ilt'lLkK of the Turlll'ua Range by MesSl'l:I,
A, O'ConllOi Ilwl n. W. RilllllllllHlh, (If Welling-tull, and of portions "f
tilt' H(lutlwl'll AlpJo. h,\ .\It" II. Humiltoll. ulso 11 1'<!hident of that city.
MI'. W, L. W1111 II 1'<.'. of 'l'illll}l'1t, cluring the lud'Il"V"lllll'uhle part 01 tlle
('4111C'(,till~ lit'aMOn lIlallll!!",l to ,C't'lIlt' h!'\"('l'Ill 111'" blwcit'h Oil the Kaikollra
Hang-e. Yal'i011R locnlitielo neal' tIll' clevated Wailllllrino Plateau. owing
chil'lh' to til(' 1I1ll'-iHtllll<.'C 1'('111101'('11 b,' .\11', W, J, Guinne",I'I, ;'ielded about
a tlti'rcl or til(' total 1I1IIIIhcI' ('"llectetl ,)m ing the year, • In all OIlRes
crel1it if, givcn. ill the Mtlcl'ipti;-C' ]lIl.I't of til iN paper. to everY' indiviiiunl
who hclpt.>c1 to }Jroiiuce the gtJllel'al rehult,
'Phl, fm·t'l!.lIing r01l1al kh incolltC'.,i IIbl~- prove thut our knowledge of tL.e
iTu'lCl't £1l111111 of t1lt' hi£!'ht'I' ultitucleR ill ver~' impcrfect. notwithstanding
the fuC't that ~,~60 "pecies of ('oleopfl'l'(l have been fountl in New Zea-
1,lDn, It may also he :.tatl:'d thut I) cOllllide1'll.bll· }ll'nportioll of thes.e
alpine beetleR are exponents of rlistillct gt'llE'l'U. nnll. a~ a rule, art' finCl'
or morc inter('Sting than thORt' of c01'1'e'l]Jonl1in~ grOll}ls procllrf'iI 011 the
lowlanfiR,
Of Stewart Ir.lo.no. \\e know Ilcl1.rC<.'lr an,nhing entomologically. only
one species. so fal' nil I ('(Ill remcmber, having bren described from tha.t
region, which. if cDrpfully RC'ul'cheii. will probnbly yield r.ome forms more
or IE''''! alliod to those obtain<.'ii h~' the memben of the recent e~pedition
to th,. ... lIbnntarctic islandF.,
~o 'i'rflll ~nct io 'I~.
(ll II' ('" ... ,,,\lU\L Hrolll L \\UIII 111- till/tin 1It1},
:n~33. P('IltllrthrulU illlpre...... um
/JIOU71. :1:1-17. l'lolll,lllrlm po ... liMli ... Ri 01111 ,
)13M. {c-nphl'o,UUl I:IIlIW. !I:J4l). {"lI'li~nl" IIrollll.
:I:~4!J. .. plllL\ui ... HUlIlII.
Ilrnul' A'TIlIIfHIIl \1 :1:1,;0. '1'('II'Orl',1 mlll'lIl,lln /11<11111.
:1335~ r~lIu.nHi"~ll"" tlll'lU.~I·i }JUJU" I :13;;1. H~ bul,l'Iill' ('u{.IiI'ndu, IJIOI"'.
:1:J;;2. tUl"itJC)]II~ 1l,0UII.
3:136. .. ...ylvnnu, I~rollll.
3337. An{)ll~hllb '·01·lm{c-Uu ... BIOIlI1. :13:i:t rIllli"ollit-iJIOUII.
a:li:l8. lovinl·ll~i ... Ihulin.
a:l3!1. obhl'lIru~ 8H}/W.
(holl" ~lll11l)Ll'f1l \ ro.
:1:140. wai,i'·('lI<.i~ IJrm,". 3:1,)4. )'i!aC'ola....pi ... 1Il11.\1lwtUh IJ/()ulI.
I :1:l,.Ii. l,ltip(·lmi... Ilr(mll.
Orollp ('ElHlllIl'lID.\Jol.
:~341. J)iclnlll)(',mthn media BroIHl. (~roup OALIIl.lW('IJ) \E.
:J34-2.· oc-dC-IDC-fa 8,01111. :13;)11. J,upel'uH Hilnmoluihi BroUlI.
334."1. tU"'('J('olli... nl'OIi II , :13,>7. fuvcil\CrUb Brolili.
:13;)8. o'c-un,lori IJloUli.
Orull II L \ \lII1H I., :13;;!l, ntrir~nniq Brollll.
:1344-, I'lnmatic1in thor,~(!icll BI'OUII.
33411. Iludularia Brollll. Group ~]I!.OTYLlI)\I!l.
:134(1. pi">('oidl'll UHlIIII. 3:U;O. ('rYJlt()d&~nt· (l('lliln"I HrouI&.
'I'llt' IIi.lgnll::'IK hhUI'" cll',lrI,\' clIClllgh Ihnt thil. CHIlI,llt VC1'y well he
t'flJlfOllllth'II \\ it la pI'c\'i()ml~' IIet.cI·ibcII Rppci('~.
!j!. Lt'ngt h, :I::i ItlTIl, ; bl'l'lIdtll, 11 nUll.
1,~viJl, 111",11' Wl'l1illgtoll. .\ biJlglu rem ,til'. 'l'lll~ hl'mt. t1.l' lI,tIllC 1)1'
its di,,(lllVI'Il"', _'fl, .\. O'('Ollllnl', ,,]w hUh l'l'('elltly III ()lIght to li(.\"ht many
1lltl'l'I,,,tillg 1I('l't11'''' froltl tilt' '1"11',111111 Haltg'I' allIl olllt I Im'aliti<".,
,Il'l'pel !.lUll Ill(l1'U ... t lOng ly aUlI dlll!l:liy, yct 110t VCI,\ eoal'bt,ly, pUlIctureu;
thOHl' hf'come COllV(" gcnt, Imd do Hot rench the bllhO; tIl\' :ll'd and 5th
illtt'I,.,til'CII aI',· II 1ittl,· blllllc]l'I' thllll thp utllCl'S, tht· 6th at HOllie distullee
1'1'0111 tllt! 11I1I~" ill toplit up by a fillul~ pum'tllrcd filtJ·in., the 7th ib 1111llClri-
(Jull(!tnte; till' hllw01 It ,.,puce ulong each ~ide i~ I'Ilthcl' oroad and convex;
tIl!' lIuLrginnl PUllctlll'I'h 111'0 lillian; the Hrulptlll'O 111':\1 the ttpex becomes
('"urlll·lv l'ulIotirlJllll 1111'1 it'I'l'gullll. hut til!' IIIUI'giu, though fin(', ix Iluite
.lildi net tlll'l'(',
Lege. 1'l\thl'l Ihil'k; thl' l')o.il'I'Il.ll allgll' nt the UPI'X or tIll' fl'ollt tibi;w
ih hardly at lin pI'ollliUCI't, thnt ()f the illtul'lIlediutc pail' ill lllo!lel'atel~'
lI11gnlnte, Antl'lIlIlIe" Hh till' balml 1 joiut!> sllIooth und gll1hl'oUS, thl'
l.thel'lI linl'l~' lI!IIl dOlold,' Ill1llrLlte, hut. in HI," hlwci lIlCIl, 0111," ...cantily
pllhCl!ccnt,
'rh", ...11:11'0 of the thorull ill 1I01l11;1"l1l1t "imilar to that uf .II, laBI',ceps
hDd At, l!ognntllm, hut tIl!' dyh'nl Hculptul"c, l'ather thick legs, and
I'eduction of labl'ul llUlll'tun'H 111'1' good distinguishing featurefl,
~, Longth, 25 rom" bl'(·IIIIth. tI mill,
Mount (~uoiu, 'l'UI'UI'lIlt HILIlgl' j Oll:!VRtillll, 3,900 ft. My llpecimen,
IIOmcwlu~t oanutgl'fl, \\1111 foulln by Mr, A, O'Connor, of Wellington, who
fleut it lllountcII Ull (·Iu'llholll'cl.
about ,I third lnoaclci Hum the thm'II'l, "jll! luthcr broad I'ouudetl
~hoI1111(:'1~,
111<'.1 lin' Yl'I·.\" ,li~tillctl~' ILlId l·el.1.lIl.ll·l~ ..,h·into, hut t.he PUllll-
tilt 10 II or tlte I>triul' i" mtlicl' tim'; "al'll hnK II "hOl·t ij(mtdllli' ~trjl~j tholl'
ito. no ~lIb.ll'il·1\1 cm'illa, tJ.CI·C' i... II Hillglt' 1111 IIct II 1"1'. plul'l'd bl'lol'c the
IIlillclk'. Oil th(' :ll'cl iuh'rlll iccl:i.
llll'll'n.iclc 1-\101>".' piceollh, 1'0\,11' Il,c!,li"h, tltl' ["hit'l illl tl'ochulItl'rto
tl'~tIlCCOU'" ~ll'toO ... tl'llIll1l1 lllf'lltUlly COIlVC\ hut 1I0t CIlllUlil'111lltU, itH fhlllk"
PUlll'tutl'. ,\ 1,,1011 Il'II i IIIJI III 1l'1 uh', hut " it It ",mit' "lmIl"" Illtt,t'81 i1ll-
Pl'C!!l-liOIlIl, '
'I he 1'1I..,tt'l·illl t .11 "i .11 (' "illl pit" hut til(' 4.th joint of th", otltel' pai I'll
ito c1iHtjncil~' longer th,1I1 til(' IhiIC1, "ilh IIl'll-clcvl'lopocJ l'atllN' eolongnttl
lobeH. III thc Illah- tlll' hatonl fOUl joilltx 111'1' Rlightl," (lilatod, Hnd of ILbout
(:'quIIl "idth; the ~llc! II1HI :Ircl 111'(' ('ol'lliroI"IU, but the hClollal disparity
i ... llll,l'llly perceptiull'.
Val', J', l"hrat/s. - Lnlll'll111 II ith 1IIi II II 11'1." uOI'ilLl'I'OIl~ toCHlptlll'I', hut
,lititout tltl' lell"t tmCl' or Ket ig<ll'oUN IH11lclnrel! in front,
1t1.'1Ic.lily Ilistiuguit-.hllll1e h," the Hubl'l)I'(lirOllll, reglllul'ly rotJiILtcc.l hiuu-
hocl~', with ullipmll'taflo illl!'I'Hticeto, but lacking tho IlHunl cariuae, 'fherl'
it-. olllY 1)111' lIl'ullll' "ptn, 111111 thu1 (If tltl' thm·,n. iM Ioituuted behind the
l1licldl~ of eILch Hill(·,
Length, 6~-7 llllll,; b1'Ollelth, ~i-:1 Will,
Huul'imu, I HCl'1Il'ec.l. five R}lccillll'Uh fl'om lIIulol' logH in JUlluary,
UHO, and Cuptain H, R Wllitehol'u, oj' Ihe (lcologiclll Hurvey Depa1't-
llIent, d\lrill~ .M arch, colit·ctl'lI IIUe! 1'01'\\ ll1'dl'l] U PUl'l'l:l} or Iureyt lell.vell
out of which J Jlicked II llotl 1(' I' . 'l'heHc el1'en~ ('11 INLVC'Ioo \\'el"o glltll('l'ed Jleal'
the ht:lld (If the Ul'tllrllkt· HiVI'l', IWlll' the 1'(:'CI'Utly RU,'vl'yed CllulfiE'lcl,
."rel, t.he tlltH'hCic ,otal' ,\1"o o.Ih'l'lIt, thOll!{h the tlpcC'illlent! III'e qUito II cNh
atl(l nninjul'l'd.
Length, (; nllll ; bl'ellilth, :i mIll.
Erua, at an clovatioll or alull1t :l,rIOO II. 'l'hl'l'c 'H'll' [ollud l>.} 1Il,\'helf
ill ,TUUUll.l"-, 1910, aUlI two hI" Ml'. W .•J. 011illlll'8H, r.tih'IW stutioll-
'uabt.Jl", in'Mal'cll, ull 1I1110nw~t ',It'cttying 11'lWl'h till till' U:I'OU1lrl: Auotlll'l'
WitH fonnrl nt'or the 1Jm)(' of Mount NgtLIlluiL(1l' by 1'..... Guinlless; IUIII
uno taken pl'OVil)lUd~' h,Y 1ll.Vl:!eli' ut Rlntl'illl\l, t'lt'vlltioTI I.!IOtl I't., has mote
htrongl~' IIcnlptul'ed elyt I'll tlUlll thl' othE"l h.
lHoHtI~-, ILcneo-nigel', clyt l'ul lUll! ~inh !lull IIpiCl'l', tht· legh, ltlltell11 IlC,
IIntI palpi telltlLceonli, lrlandibicli pitchy l'l)(1.
Head tlmooth. l'('rldish ill front. intol'ocular ilupresslollll elungate, the
l!l'u.ce between each of thelie anll the IlLttlml groove somewhat elevllt",d.
EYeI:I lal'gtl, not pr(lruinent. Hndy rnccted. Thorux a fourth hl'outlet
tl~l\n lung, widest nelll' the IIIid<1lc, itl! Hi,1"'11 rliHtinctly margined, lUode-
t'ately rounued, !!lightly SillU()ut!ly 1lI11'I'O\\tld behind the mi,lrlle, p()stel'iol'
anglcK l'ectallg"uI11l'; th(' 1L1Ittll'ior, though not prominent, are mlll'e dis-
tinct than Il~lIUI, owing to thtl lllight dilatation of the latel'al llHl.rgiuz"
there; disc model'ately convex, IllU()Oth, itll centnl HlIlcUIi dOetl not uttain
the apex, the baStl hus duplioate fOHsae nelLl' oaclt Hide, aud the (luter une
is separated h'om the lateral mar!tin by I~ oA.rina. Scutellum distinot,
I'ather broad. Elytl'l. ovif()I'JU. u fifth Inngol' than hroud, widest near
the middle. of the Hllme width aM thorux at tho baSt', witlt 1'",HextHl mar-
gins and rather broad chaunels; they are moderatel~- PUllctute-Htl'iate,
the striae, however, uo not I'tlach the buse, and, except tho sutural pair.
become indefinite towards the C:'xtl'cmity, and those bey-onll the 4th are
inrlilltinct nenl' tho sidel!; :ird intel'stices lllightly broader thltn tilt' con-
tiguous onCH IIJ1I1 I'ntller coarsely triplillotllte, the IIpical l'lIJ'iutLC Ill'(> well
developed.
Certain peculiarities are observable ill thiH Hp()ciuH, 'rIle Inhl'll111 iN
rt!'markn.ble, for in place of being Rubquadl'ate, and widely cruarginate
in ft-ont. it is quite conioal, and beal's Itn t,'xtltll'te<l tentral RCtl~ Illld 1\
pair of finel' ones at the sitlCR. The maxilllu',Y palpi 111'0 6n('I;I' HtltOIK',
The inner emargination of the front tibiae is lesil lIiMtillct than usual. RO
also is the basal margin of tbt!' elytl'a. The surfaco of the frollt tarsi i"
mure puhcHcent. There is onl~' II single oculltr ~t'ta, Ilnd that of tile
thorax IIPPf'UI'S til bl! abscnt.
Length. 4i rom.; brellclth. 2~ UI1l1,
Greymouth. Describcd fl'oUl II fE"llllllt' sent b~y Mr, .T, H, l,t'wiM, wboKe
na.me has been givpn to it,
~ltllatctl bel\\ccn tli(' nli,Mlc allU lh(, t-it1('h, I1nd, owiug pllln,' it) tl.l;IllIlIIl:-
I ,It illll, aro lIul VI'I,r (It'finitl', 1~I,ytJ-n wicl, ...,t 1)('1'(\J'o tlw lIIidrlk" II goml clenl
,ltt'mnHh'll pOh{('1 iol'l~', with tIll' latel'ulllllll'~inll well dev('loped to withhl
,I ~Iltll t 11i~tallt'(· of tho 11))ic('H; thl'Y 1I1'l', I'('intively, IItl'ongly loel'inte-
p11lll'latl'; 11(',11' Ill.. HUltll',·, 1111\\l'VCI', tlw)' 111'0 Htl'iatl' III III lulller IUOIt·
lilll'h- 1III 11(' f II I ('11 ; the ullicnl clI_I'illlll' 111'(' 110! Vl'l'.'" Idllll'}I1;. ell'vu(('u, IUHI
11. .. '1flac(· h('t\\oell till' ouh'!' HOI'II'N of }llIl1l'hU('H Hnd thu Hcnlvtm'pd mlll'-
I.! i 1I,t! l'lullIllt'l 1)11 I'lll'h pl,v! 1'11]1 iM HlllOnt I._
trndel'Hine Rhilling hluck; nl"lollll'll 1I1ll'\'('II, wIth hl'UlI I11,1 dl'Pl""Nl·d
'!utUll·t! het \\'1'1'11 itll H(·A'Jll(·ntf.., lIlt· tel'llJinll1 biplIlIctate at euch !oHle o[ thc
middl(' at the apex,
The mthol' ('longat(· ('on I0111', longiHh thorns, I\nd the- postel'iori;o,'
attt·lllwtt·t1 ('ol'tliform hilltl-hotl~' 111'(' go,1(1 (1iRcrimilllltin~ dllll'ucte-I'N.
Lell1!th, 6 111m,; bl'('at1th, 2~ 10m,
i\fount Quuin, 'rlll'III'1I1I HI! II go". ,.\ '11111111' 1',·llln!t·, 1"'0111 MI', H, W,
Simmonek
(lI'OII}, Poe,ONIDAI!l
3174. Oopterus laevigatus sp. nllv. OWpfl'rUB Guarin, MaD. N.Z,
Coleopt., p. 54.
Subublong, "lightI~· COllVl'~, niticl; clIHtlineollll, femor", and elytl'"l
mal'gins fulv('hcl'nt; thc tibiae, flu'si, MIIl·lllllle. !lilt! lllbnllll more or leRN
"ufo-custIULCOllh,
Hend smooth. till' t'I'ollilll illl)lI'('Hlliollll wall 1Il11l'kl'(1 lIlltl oxtelldin~
l'l'Olll the intl'l'lIl1it'IIT1111 ImtUl't· to tho lllitldic of the tl,"CH, which are largl'
'11111 llwclorately convex. 'rhoI'll" 1\ Ht'v(mth hl'ouell'r lluLn long, widest at
the middle, gently cUI'vodl~' nlll'I'()W(·tl nnt,·l'iOl'l~·. gl'lIt1nnlly, hut I4l'lll'cel,\'
NinuoulIIly, cOlltrllcte<l behiud, with I'cct ILngul II I' hut not acute ptlRteriol'
angleH, hal:!e tl'ulloah', apt·~ "'lightly iIlCllI'Vt"tl, lah·rlll 1IIlll'ginl:! I'llthl'l'
thin j baMal fllllimt! brond Itucl well mal'kl.'ti, oach I'it'plll'lllacl from the side
by It distinct carillO" the I:('ntl'al gl'oove hlll'(Uy uttllillH the IL}ltlX, itl!
w]lOltl Hul'i'nee illl}lllJ1otllte, hilt with a i'ow filll' Ntl'illl' 111'111' tIle midrllp of
the haKe. K('utl'11um Muhelll'tllltt', Rmooth, EI,,,h'lI oblong-Ilvnl, IL littlt·
wi tIel' than thurltx at th(' bUK(l, not 'luitt" thl'ic(' ifH h'lIl!th, with hroad
Iotel',ll mltl'ginH, whit·h, howl'vcr, bl'eOllle Vl'!','· nlll'I'OW 11('111' the bl'()lldl~'
rOUlJlINI IIpit'('K; 011 ('Itoh (·Iytron thl're' IIJ'C .( Idiglltl,' illl]lI'I'''u'If'll, illl}1un('-
tute Htl'iUt·; !III'!oIc UI'I' ohHol(,te' nl'ur tIll' hUhl.', till' Nuhn:!l II10n~ l'l'lll'llCM
till' lI]lt''< j th(' apicnl pIi(ln iK eliMlitlt'l, tIll' hllMI' ill tt'ipulIt·tlt'Ln hut tilt'
:h'rl intel'Htiel' iM ollly V('l'~' fl't'hly HH, lito Hidl' iN (l"u(ll'ipll1e·tllte JIt'ur th(·
hhoul!l('!", t hI' pm.! I'l'ior Plll1ctlll"'11 111'(' 1('HM (It.fill it(', t h('I'(' 111'(' 111m, lit 1111
T I'llI' tlu,' mi.ldl(·,
'I'jhin<> MII'night, IllI' rl'(1111 puir 11('111'1;.' glnlllllllN, 1I1I'l1lh('I'1I Ihl(·ly h(Jt()FIt',
TlIl'Hi wi'li ,,,,,110\\ N(·tU(' IIhove-; till' nni('l'iol' \I ith tIw baNul joint <IiJILt('d
anrl ohl01lg, 1111' III'~t (lol'eli r"I'IlI, nlMCI elilnf(·tl. ,\"(·t l'III'(U,V aM hl'tlltci, the
inn(>!' n.ngk of hoth Hlight1;V prolongl·d j 3nl ancI 4:th cordate, evidently
smllll(,I' thun tht" 2nd, .lLnll eOIlRNllI('lIt1,\' 11111'<111 lit 1111 (·xpanded.
In 0, TJ(lfIf11l~ (1141» th<o thol'l\x is wic1E'F.t hl'fore tne midrlle, and
tbere-for(' Auhcol'dnt(>, ann its scnlphn'e is rliffel'l'nt; the elytra are 1!11Ol"tor
anrI more oval, with fine ;vet quite perccptihly pnnctul'cd striae, the 31'0
intert'ti~R arl' tl'ipllnri'nt.f', IlnO tIll' I'OIOl'utioll i'l ll1toJr('tht'r illll'kel',
&. I.ellgth, G! 11Il1l.; bl'l'ntlt1I, 2~ IIUlI.
Habtw('ll. Ileal' Nupier . •\ Mingle millo, f01U1I1 by All', n, Sl1tel', 'L'his
genul!I 1'.11 ... fllm'I',1 I,,' LlIl"lI'IllIiJ'" ill tlU' ('I//'U/fICfllltllid'llJ, h~' Blltl'" ill the
I'oflilnirltll' , '
l.·/'allsa('fIOI/~ ,
390
(h'oul' L·'mwllomAj~,
3175. Trichostemus wallacei I:Ip. llO,,", l'l'ich()S/l'mus Chlluuoil', Man, N,Z.
Colt·opt .. p. 31.
Uhll'ug. hlighH,\' 1·(lllV....\, Ilwtlel·lItcl.\' lilt id; hhwk, cl,\ flul Hlltl gillo
l.tintl.\ 'l'illlh'",,·clIt. Ipl-l" awl II IItl'll II ,\l' Illl'o-l'icl'III1'" lip ... or )'alpi 1111'1'''-
relIt,
Hcn.! hllliloth, 1'1'11111 al 'I'ov('ue OIUlIl-\lItl', \\ jth ::.! HC! il-\l'l (lI1K PllllcttlLCI>
a!ong.,icll· Ullch eye I1l1cl the FUlIllC II 1llUUc I' Oil tltl' !'Oll'hl'lul; 1111>1'11111 tl'nlll,'
vel'~C, fet'hly l'!lllll'ginnte, I<~~'eto. p1'omillout. 'l'hol'll \ huL,ll1l1el rat(.·, II
tldl'el III ofldul' thnll lOllg, tlll.l ape:\. Willdy illellI'Ve(l, b,lhP 1I1l'Ilililly 1.'111111-
gill!lte, Intcl'lll Illlll'gim, IllOulll'utuly thiel;:; it ito. n little hl'oaclpr hcfOl'l'
the mieltlll' (in Millie C\lllup!t'!I h'\l'cll.r lit all) tl~all t.']t..CWhl'~·I" gelltlJ
l'lIUllIll,!l tcmul'ch, thc front, IlclLd~' I'Itl'aight bclJlllci, pOHtt.'l·WI' Itllgles
u.:mctl~' I'el·tnngulal'; thc IIwclillll hUlCllh elot'M not I'l'ueh till' ,11'1':". allU ito.
hliglltl~ l'\p:nlllell hl'ilincl; thl' hmml fOlootoll.Lt.', to.itllutl·11 hd \\1'(,11 till' mioelll'
IInel hilll'H, Hl't' llll'gt', Ioo"lIlt'what IJl'olollgl'l] llltel'ull.\, 110 tIlllt lhe '-)JII ('l'
nClIl' t'llt'lt IIngll' Iool'I'IlI!! flath'Hed, K('lIt ..lllllll hh·illll' at llll"C, 1j}lyLru \dtll
dentiflll'llJ ~1t()llltlcl"fI, ,im,t l'L'I'ccptihly hl'()Iulcr tha'l thlll'lll nt the hal!le,
theil' billclI Klig'htl~' eU1'vntt', with well-elevl·loped 111 11 l'!dllH, llIoell'l'utl'ly
sinunte-llllgllRtnte nent' tll!' P:..h·l·lllity; their Htl'inl' .11'" wl'll III II I'krll, und
become elt'l'pt>I' nnll wifl,'1' tuwlIl'lIH tit!' Hiol'!! Ilnll Upit.'I'Ioo, hilt 111'(' I'uther
il1Cldinit~I~' p1l1IetUl'c<1; inter~tiC('<I plnne Oil thc lIlilldle, ClllIVl'~ IICII1' the
I'Xtl'(,lIIit~, IIncl hille'l, the :Jrll IIml :ith gcnel'II11~' hlightl.\' hl'ouclt'I' tllllJl
thohC 1lL'IIl' tl)(' !lutnl'c, the fOl'mel' tripunctllte, the itlt \\ Ith !'i punu!llI'es,
!<'CIlIIII'1i IllOell'l'lltcl~' Mlcnoer; intCl'lIll'clillte tibial' \\ itlt hliglttl~, prll-
mint'llt hiIlIl lI11gle ... ; tlll'Hi l'athl I' Hll'IUlcl'; tltl' \.);111111 1'011\' jnilltH of tIll,'
IInteriol' onl~' Illodel'lltoly dililted,
lTndpl'Kiuc gluMsy blnck, uhllost Iluite blllooth; ttl(' I1l't'\ IIf till' tCl'lllinlll
ventl'l~1 Hegment qUlldl'il'Ullctllh.' ill hotll b(.'XCH, •
Frm(de.-Elyh·ul toItl'iUl' vel'.\' liltl,ly hut l'utlll'l' IIIIII'C Ili~t illl't]~· Pllll<'-
tatl" than in thtl lIlale,
I have ~eull II C]lI?.oll Hl'l'eiltll'llH; hOllll'tillll'" therl' IlI'C' ~ 1'1l1lctUI'CI. 011
tho 3rt! intcr!ltiCl'H nllel 1 cm tlll' !i1'0, Iml ot hl'l'\\ iioil;' tlw~: IU'(' hCllIICl~eneollloo
Ilnd vllry hnt littll' ill Hi1.l', 1 h,' 0111.\' hiudj,II' ~pl'l'il'i:ol i" '/', //(lIlltlt'l'rnllix
(26:H).
Length, :21-2:J 111111.; hl'l'aclth, H~k 111111,
Wflirit'i, Ht'l\wlIl'el Kllikolll'Uh, 11 iH "itlt 1'1",,,111"(' thllt I 11111111' tlliH
hPlOCit'h uftCl' itH 11iHCtlVI·l'l'I·. :\Tr. W, L. Wnllul'c.
3176. Pterostichus hamiltoni sp. 1I0Y. PlnoBfir/lUs BOllPUi, Man. N.Z.
c.toleopt.. p. 31,
l-:!ubohlong, IIUI'l'owetl Illotlinlly, ouly Illightly uonvex, hhiuillg; hlncl"
the tibiuc, tllI'lli, 11Iltl antennae picl!(l-I'ufuUH, pnlpi pal(·I'.
Hend, incluoing the pruminent e.\'es. J'Uthel' lHU'l'O\\Ol' thull I'I'Ullt uf
thllrll.X, nlmOtlt qnite smooth, \\ ith cloIlgate Cl'Ouhd impl'eHHiOlIH, 'rllllrux
willeRt Ilt the miclrlle, regulttl'ly rounded frUID the obtUHO nnterinl' IInglf's
to be~'llnJ thc middle, ILnd fl'om thence consiclcl'nhly, and KOlllcwhllt
~i~\l(lUH!y, 1l1ll'l:0\\"eel townrus the wen-define,l roc~lIllglllul' l'o!ltel'iOl' nugleh;
It lb ,llUltc .n tllll"ll hronclel' thIUl,lcll\g; tLll' aI'ux. IN elulll,l.\', the hllKe Rlightly
Illechllll;\', lllom"\'oc1; tho blll!lIl fotoltolue m'o e1el'V, I'ntllel" elollg'htt.', elistinctl~·
RPpol"uted froUl tlll' Hi(lcs lind IHllI't' widely fl'om tIll' lIIilleUl" the llI~llilil
groClve dOeb not I·tlllch til(' apt'x, tlu.' IJUl'fuctl iH neu I'ly slIIo(lth, Hcutcllum
Hh'tlugl:v w.;ildl' at the haSt', Elytl·n. morc thlln ctollhll' the le1lgth of
thorax, u. gooel Clenl, yet gl'lldunll.,·, I1I1l'r()\\'ed tnWIIl·tlK till' ],IIAe, tllC
3111
ILlllUCI'.ll HI.I1'gUl", rhickNlL'tl hut Jlot ]lltl,jl't,tillg, 111111 ]lut ('\cccclillg tllP
thUl.lcic uJll.IlctI in width; till',' III'l' \"el,) p\"itll'lItl," ~illlillh' hehilld, till'
apicl'~, ncvcl'thcll'bl>, ,up '1Uitl' 111"1111 aIHI mil) :-.lil.\htl,\· I'otlllllc,!l; th~y aI'l'
tillcl~' :lml l'l'glll.II'l.v )lIlIU:tate·,..,tI i.lh·, llltl'l'lo,ticl'''' pIalit' 011 th~ diRt·, bllt
hCCUllIill!.!, II litHI' t'oIlVI''\ IlPar the ~iclc'", ancl c'xtlf'lIlity; the :II'cl. :J til ,
II lit I 7th III (' ,.,li~htl,1 IlI'O.lIlt'I' than tht' othc'I'''', tIlt' :11'ci III (. t l'ipmwtlltC',
all.\ till' ilh hllvl' ahout G rlltllel' Illl'At' pl1lH'hll'C'''',
Uu.\t'l'l'Iiclc' ... hillillg hIlII'll; the tt'l'Illillnl Wilt I al "l'gllll'lIt hm, II bl'ollcl
obliqur tlCPI'l'loolooioll l'\tl'llIlilll,( i'IOJlI tho hUM' to tIlt' llIiclcllt', lit l'ncll Kille;
the .lpiC.II !Judioll i... II iii til' c1"pl,C'".,ccl anrl cli"tilletl~· longitmlinall,\'
... t1'igu~C', ulIcl i'l 11l}l1l11etah' at (,lIch Hidl' of tIll' llIiatllc. ,It the c'xtr(,lIlitv,
'I'hen' HI'C' ~ "'l,ti~cl'ou;" pUllet 111 ('101 Ill',ll' (.'uch t'~'(', tIl(' Hllme nlllllbei, (Ill
1111' fOlC'Iwa,l; till' It' i!. clTle bt'1"Ol'(' thc millllle> at ('(I('h ~illl' 01' th(' thOl'.l.'I::
,mel nnotht'l' ,II till' hincl Illlg'le,
'I'hi ... h ,lone, ... to thl' hl,t,tioJl 1'lllll}lO'i·tl of TI'II'II(J,I("II'I~-lik(' "Iwelch (,(,t'
p !)HCi. l£a1l ~ ,Z, l'oit'ol't.), hilt al'il'l' 1'C11111HIl'i",01l with itA lIllml'I'OUI->
mCllllwlh 1 I'ail to find an,\" l·\.:lC(\,\' like it, 'rhr llllllHllal h('u1ptul'c of the:'
In~t aliololltillnl M'gllWlIt iH a g'ood glliclr to IliH('I'illlillntion.
~, J.ollgtll, 2;, 11I1lL,; bl'('lIdth. H~ 111111,
Bold P('ak, \YUl.lltipU; I'll'vatioll, Ilbullt Ii,OOO ft. A "inglo iC1I11I1c.
1I,lI11l'II ill hOllOIII' of JIll', II. 1I1Lllliltoll. itH tinclel', wILII!!(' l'ec('nt coll('{'tin~
to 11 1'101 hnv.' IIcl(lt'(l cOIl!.icIl','ahly to 0111' klllmlC'clg'l' Ill' till' ('01('nptE'l'llUH fnulla
"I' the 1Il1lUlItnill1111h l'I'~i01I'1 IIf tl'r HOllth J1411111('I
3177, Allocinopus smithi HI', IIUV, .IUoci'floPU8 13"OUIl, .'\IUI, Mng, Nat.
Hillt" HI'I', 7. \'01. 11, p, 1107,
Oblong, Hhghtl,1 ('IIII\'('\.. IlIllfleratt'i.1' lIititl; I'1Ifu-piceOllH, lawl'ul IIIIU"
ginN (If tho!"1\ ,llIe1 el,rtr'u "'01'(' 1'l1ff'I'I('Pllt, lahl'l1ll1 IInel Illlll](libll'~ dhl'l,
recl; tht· leg" 1111(1 unh'mllLe l'l1H('O-I'Ul'oll!.. hut 1\ itll th(' hUHIII h\1l joint!,
nl' thl'hl:' lILttel' lIncl the }lllipi l'ulve"cPII1.
Hent! HhOit uml lmmd. n't l'IItht'l' 1111 I'I'OIH' I' th:lll the tlWl'ILI" 1!1IIonth,
"ith It pail' 01' ... hullo\\' ill'!t'I'lllllctllIul 1'0Vf'llt' j l'l'iHiomo IOllgitlloillall~
'itt'illtti. IIml, IWIII' (,at'll 1'1'0111 UII~J('. willt II h(·tigt'I'OU)o, IlIlllr.tUI"C; there ill
anothcr OlJ(' 11('111' til(' 1IlIl,k fir ('Ul'l! (lfl', 'l'hOl'lI \ ah(ml II thit'(l hroadl'I'
tlmll long. iill uPI')' \\ icil'l,\ hut 11(11, (kC'pl,\' i 1I('IIl'V('II. (0,(1 t hut till' 111I~II'h
IIPl'l'H1' ,,[ightl," PI'Olllilll'lIl, hUh(' "U"It'lIlIl'Utl' IIl1cl Ihll'l\' 1I1111'g'iuocl; it i ...
II littll' \I icil'l' "l'fol'l' till' lllicltllC' th.111 I'I~I'\\ hl'I'(' 11IOC!c'I'ufl'h' 1'01111111'0 t1U'I'I'
lIIuI mthl'l' ~I'.llllIall,v 1I111'l'O\\l'ci tU\\III'c1h !lll' '1·(,(,tuuglllu'" hut 1101 Ilontt:
}lUHt('I'iOl' angll'H; the la1l·rllllll.II'giIlH lilt' \11'11 clC'vl'lopl'ti, ultll lhl' dULlIlwl...
inHill,' tll!'h(', thl'Olll.l1t 1I11l'I'CI\\, .11'(' II lit til' \\'icll'lI('c\ in rl'oul; the ""Hnl
fllHMIC', Hitnatl'c1 hl'hlf'l'lI till' mid"l" lIucI Hi I1('101. Ill'!, '1hul1ll\\ allll iur1('tillitl',
lln(l Cluthilil' 111l'H(' til{' 101111'1'11(1(' iH !.li~ht1,\ IIl1ttl'lIl'ti. till' ll11'cliall gl'Oovo
extt'lulH ft'UlIl 1111' bllfo.ul 11111 r'!I; it I hilt (10('101 1I0t n'llI'lI till' 1I11l')', lh('I'(' iH IlII
otht'l' lliRtinct "ell1 }llIlI'l', 1:<~I,\'II'1I m'u 1'1,\' t w i('to 11101 lclllg aN hrol\(1, 1'lLt lIul'
",ioe>I' than thlll'lIli. ul til(' 1)11101(', tlwil' HicltlK di'ltillctl,v Illlll'g'incr\ IIJ1(lll('llrl~'
HtrlLight, hut wi(ld,\' Kit1l11l11'c1 lIucl cOIII~illt'llIh1,1 11 a 1'1'11\\'('0 tOWlIl'llK till'
f:'(h'l'mitr; thril' illll'undat..· Ktl'ill<', with til!' l'A.I'('l'lioll or tIll' tl\O Imtul'nl,
whiel. (10 JltJt I'(!nch tIl<' ha:-.aI Illlll'~ill, 111'(' \\1'11 111111')(\·,1 tIll'IllIghllllt. tho
!lhOl,t I4cl1tl'l111l' Htl'illl' al ...o HI'(' 11 il.l illl,t ; illt!'I'Mtit'PH Hilll}!I!· 1111(1 1lt'1I1'1y
pllllll" t\tC'llltl'l'nl PlilIl'i il'm'lII illll"'('KHioIlH al'(' nllt C'IIIII'H(' m' Ill'l'P,
LegH "tollt, thl:' tihill(' I'r it I!l;C'!1 illl1 urcll,\ \\ itll till!' H('tll!', till' illtl'l'-
IllclliulC' willi '1pillirfll'lll .. C'IIIl·l'dl'llIlIn~ ; 11I1'",i filll'l,1 "I'tl)~l' nhoVl'
3112
('wlel'hillt, hlllllillg, ttll' hl',Hl Ilwl hll·nh( !,ltdl,\ l'l·II •. LhIIOIll(\lI lllglll-
"ireoll', l\Iet.IHtl'l'llIl11l inllh,tinrtly glllll,nlatt., ,Tel'~lIillell wntl:nl ~\gm(:llt
uhIicplI'ly nlU"l'u\wc] (0\\111'11100 thc' C'\(I'I'lmt~', ,\lIIICII 110. hul,lU('lIIlll1lull', \\Ith
,I l:Iill!dl' Hctj!!~'I'(ln ... [JlIJl('t 111'\' tlll'I'C' .It ('al'" hlllc· of tIll' III 1111(11'. 'llll' CI'ollt
ILno Hlicll11r tm'Ri hl'al' olollg'lIll' ,d.lI' alo1lg tlll'il' 100011,'" 111111 1\ IIpill i1'l)I'l 'I
ollC' lit till' ...id" of tll(\ IntRal rUIlI' jllintH,
11,,1, ,-PrIlRtol'nnl IIl'oro'!H II ith 11111111'l'oIlH I'Il'tnc' lit, till' tip, M(·ta-
hll'l'lIlll11 hhol't, with lIlinuh' gl'a1l1l1~ 111111 hcbll' ill tiJI' miflrllt· BalleLl
youtl'Ul"'CglllCllt ut the ImflC,. ill tIll' lI1icll11l', Ihlel,\ Hculptul'l'c1 IUlIl minutely
h~tOl'>tJ. thc' tC),llliJlIII ClIII' flilo.tinctl.v toinllllte IIUIII' thl' l, ...tl'UlIIit~', ulIglllnte
nt thl' COlllllllllCOll1l'Ut IIf fhe ililll1molitit·~. tIl(' ap(\'\: it hl'1 I' nllt lUII'll IIl1mdl'o
:lllc1 \\'ith 1\ }Jllil' I,f l>Ieti~cl'OIlIol }ltlllctnl'c'!o\ Ih('l·l·.
~. T,clIg-th, X1Il1ll. ; hl'Pllclth. :1111111,
Ratapihipiili 1"01'1'10,1. Tal'ulIllki, A IJHil' IIf 1IlIltihtt~·cl "'IK'ClIlIUllb wen'
tUlIlllI 1)\ :\11', \y, \Y. Smith. \\lto,,' 1IIIIIIe htlh 111'('11 attllclll'~l to thiio
"pl'(·il". "I'IIC' c1i",covul'~ of thu 1I1ulc ito iIl11)(lI'tllllt. aM witll(lIlt it I hllcl
hf'en unul,h' to Ilt'tL'l'lIline tIll' g'CllUH to \\ hich othm' I.Jlccic.·1! bl'lcmgl·11.
o,,~ -'rhil-o anrl tIlo following Kpeciull will IIl1t, 1 think. 1'('lIlaill POI'-
llIulwntlr \vitlI AllorillopuR, 11101 tlU'y cxhibit cCI'tnill <Tiffpl,(!11(·c.... III buth
,c'xel> thl' PI'Otot(,1'1Iu1 Pl'OC(,H!! iH HPtigel'lluM. The- t('I'lIIinul 1I1'ticnlntiCllIH of
t1ltl pa.Ip! HE' :Ioute at tllt, (\xtl'l·mit.\· i IItotll'ull of hoi ng ohhll-oU or l'mhtrun-
c,ltl'; the lllIIXillltJ'Y uro till ely 1II0tOllt', IllI(I the pUllultilllllh' joint ilol shot,ter
than thp tl'l'minal. 'rh(' l'igid -;t'tut.' uttllche!l to til\' IIH'Utlllll Ill't.' very
I·lollg'utt'. hO aI",u Ill't· tllC piliI' lit tho apex or tIlf' ligonlu, Th(l 80]e8 01' the
mule tllI'Hi Ul']lNU' ttl he without cU ...tinct HPOlI!?,l·-likc V('htitm'E', hut 1Il~
specimen having Inl.t 011(' foot. nnd 110101 thr otll('l' iH 1I0t l'l'l'f('rtl~' fl'po from
f;appr llllltt(,l', J no not f('e1 A1I1'1' on thiR point,
31 i~. A11ocinopus castaneus I>Ip. nov.
Oblollg'. IlII1(lCl'atl'l~' Hhilliug; hl:luu nUll thol'H~ clt~rk CIIHf.IIlWlIlIh;
lIutt·nnnl·, Il'gH. und 1'1~·tt·1I J'ufu-rustalu.'Ou!.. tl1081' laHl Il1d01' ulong thc
hicll'~ 1}1:1I1' (It,· IlJ,il·(·... ; Inhl'11I11 lIucl muucliblt'H pi(·clI-l'llfnllH. pulpi "111-
Vt'llCl'nt.
III Jllllt:lt l'UNllt·l·h. "Iiwilnl til ..t, Mllif/li, 'j'llt' tip ... or tilt' tl!llUiuII]
juilltlol of tI)(' ]llIlpi Ul'(:' 111111'0 poilltt·C!, und tltc :"hlu joiut or th.. 11IuxillnJ'Y
11I00·t· c.' II I'\"ll II', '!'11l' flll'plll'lIll ito It'KH uviduntly "It l'iatt·. Tltt' IIntel'ior
1U\~luH Cli' tlH' tllol'llx. m'l'iug' 10 tllo VC.·l'." Hlighi Itllicul illl'I11'\'ut 111'1', lire
IeHI'> IIl'dlllilll'lIt. ith latcl'al 1II111'l-\'illll III'\! thilllll'l'. uncI tltt' hmllll fUHKIJ.e
I'Iltlwl II III I'C' oIi ...tillC't lIilcl I'longllft·, '1'111' "lyh'lI 111'(' HOIIIC'\\lmt 1ll0rA
ovifol'lll awl 1,·...... IIcl1l1liuutc' PllloIlt'l iol'1,\'. 'L'hl' Hl'tn lit ulIuh Hic]1' oj' thll)'l1\
i... plul'l'cl ht·flll'l· tIl(' lIIi(II11('. UI-o ill tIlt' pn·ccclill!-\, uml fnllnwillg H)lI'tJieto.
(T litil'lhiclt· wholly 1l1(·l·CH'l1fIlIlH. PI'oMtOl'nlll Rt·t 110 lIU1l1l'I'IIUH, hilt 1)1)t
l'mll·Hl·. '1't.'I'llIiuul '\"1'1It1'111 loot'gllICIII \\iclel.\' Hillllnh.· tllwut·d ... the l'xtt'(,lIIity.
Iyhich ill hl'CI;UU,:I I'ClullIlt·ll. in~tc.·llrl elf llf'iug' ~ulJ1\(,l1ll1iunh'. ml ill the
TII'c.'Ct'dinl-\' RIIl'cies. it iH Hilllilo1'ly hiHt·to'!!!.
~. Lt·ugth. 7-'l~ lUlU.; lll'tlutlth. 2i-2! 111m,
:\rllk~tll. HUllUIl UmlJ..·t·. Both HpecillwlIM 111'(' I(,llllllt:'h. 111111 '\Olt' 1'0111111
111I.,ut twent~' ~'l':U'K '''!;O, :It 10111( intcl'vo.lK. cltning clIll('ctillg'-tollrR.
:\179. Allocinopus angustulus tip. nov,
UaHiIlll!:lU ),U£011:-. shillillg; tlItl Iegll. }Iulpi, uull hUHnl al'ticuiutiollH oi
an~nnlle fulvebCtlut; l'tHIlh.il~illg jllill~ of thl'Xl' hl!!t dud,.t·r unrl opaque j
eplBt01l1e. labrum, aUIl lIIundlhlc'lI l'odtllsh. thE'sl' lliRt. lIowevl"', uro nip:I'UH-
cent nt the extremity.
Hl'IUI nt'nil) tluitt, ~lIlt1otll, 'l'hol'll\ .lbout II ,'''n!'tll bruac}t'l th.11I
IOllg. lIot pell'ectl~· t.lllooth, hltving HOHI(' oht.tllt·t<, longiturlillul lilll·.Il'
illl}lll'~~iull" ill I'lout Hllll 1I'11llt.V('I'1I0 0111'101 IIlI thl.' tIi ...e; til(' 1I1l'l>inJ 1'1111('111>
IWUI'J,\' IIttain" th,· .11'1'\, lh,'l'l' :ll'l' llO 11:\11111] fO'~lIl'. ollly II hmlltl ol,litJlw
illlPI('h,ioll or ('adl looi(lt, m'al' till' nnglt·; ih, I'oj.l,,1'0 .11'1.' l'Utllt'l' hlightl,\,
nltllllHt I('g ulu I 1,1 101l1l.l('II, rt'l l'utlU'1' mOl't' htr.lightl,r JlUI'I'Ollt'.1 to\\ul,(IK
the ~nhl,(,tJtllllgll1.11 11.1,,111 IIl1l!,lt'h, "~I~ II',L \\Jih \\('ll-IlHu'h.cd illlIIUllCtutc
.milll' Illlil "hod "l'"h,l1II' gl(lOl"t'h. Ih(" .111' 1':llll' I \\illcl Ihnn thoi'lI\ lit
the 1111"1', 111111 II gOlld tlt'al, tlullIglJ gl.lIlllUll,I, 1I.IIIOIIl'II Iwitilul the jJI"-
tC'l'iol' rellIOI',I, II ith 0111," n'I',I' "JiL\hl hUh.lpictll "lllllOl'oiticl-l,
"'ht'll l'OIlII,IIt'tl \lill, .1. '"l1tll/' ihi ... ih '(,,'II to 1)(' nlll'I(mCI' nlt,l
tI;~ttel , ')'h" rl'Ollt or thl' 11 III l'lI \ iK ,I liltlo 111011' i IIl'UI 'l"C'U, loti that it"
ungloH "'l'l'1I1 I,IOI'C Pl'Olllillt'Ui, hilt Hle pOhll'1 illl' ,'"gll'''' .11 l' 11101'1:,' ohhlM',
nno the hllM' ,1il.l'htl~' l'IlUII'L\illll!c 1I11'.liHll~·, The 1'1,\1111 appl'lll ro h..
ILIloI'1'OWel' U11tl long-N, IIn(l 111'(' lIot lit :111 ablUjltIy Illll'l'O\\ud 11C'.1I' tILl'
I'l.tl'tllllity; thc (lOI'HllIll i~ 111',111,' 1I.lt, hilt the _ntul'al II'!£ioll i, ,I littk
('levntod b..hilHl.
!j!. Lc'ngtlt, H UIIII.; hl'l'atltll, ~~ 111111.
I!'ol't,v-mill' llw;;h, 1)('111' Nll.l'it·)', A ~illglt' 1'C'IIIUll', nll"(,1 Ull' 1111111])('1'
:152, from MI', II. Rntl'I', hail to hc Hot Uhilll' "01' IIllIn,\' ,1(:>111'101 IIlttil tlu'
I'('cent IlihCIlVPl'Y or tILt' 1111111' <,r .I, ,11/ iflti
:URII, Zabrollothus major HI'. nov. Z(/bl'()rlfllhu,~ B1111 I II, ~lall, ~ ,z,
(loll'Op1.. ", 1:127,
t Iblllllg·ovul, gl'ndllnll," 11l1l'l'IlWed IIl1tl'I'iol'l,Y, Klightl,\ I'flll"(,\", lI,odl'-
rntely mining', pk'OO-llig'l'I'; It'g~, IIULllllil']('R, and lulll''''" pitl'h,I I'ell, till'
antennae, palpi, and tnrsi pll.l\!l',
Hend narl'IlII't'I' thllll thOI"II,(, HIll!JOth. rl'(lIltn.J. imlll'ChKioll" 1[llito Obh"-
leto. 'rhOl'ax illum'vu(l lit npex, with obhlHO (tngloll, tIll' willth nt till'
blll:le nearly It third mOl'l' thun the ]('Ilg'th ill the llliulUo; itK Rinell tliH-
tinctly lllurgilletl, nenl'l~' Hh'ltight lor two-tbil'c1l:1 of tho ll'ugth. being
only very Hlig-hUy 111111 !,II uclually lllLl'l'OWe(1 IInt.t'I·iOl'l~·, hut llelU' Ih,'
fl:Ollt l'athe'I' llIf1I'P l'ul'wdl\' ('Il"It'IU'II'll j hUM' medially I'IllIII'~illllt(' nil"
)'cRting till tIll' C'1~·It·Il, witlt uhlm4l'ly 1'('otllllg-UJ.Il' nng'IC'II; tlll' tnullinl gl'om'('
~tu.l'ts f),om thl' hntllll IIlIIl'gill hut dlll'N lIot 1'''al'll thn apt", LI~lIltl iIllPJ'I'H-
lIi(JUII froblo amI C'lllIig'lIft', 1'1I1t'l'" b"IIII'('1l till' Illiddlt, .tnt! Ki(h'K, HCllh'l-
lullt tl'ir\T1~nllll" Il~lrtl':I M'III'I'd,I' lilly bl'onlll'I' thUll till' HUll'lll: at til('
hilKe, 1111 1uI.'I'I11 nll~lt'" tltil,I'('III'(l IIlld pl'ojecting U1tLWIII'(Il~', ,,"ly \'1'1','
1I1ighUy, ilUWI'Vl'I'; till',' HI'I' II litill' Witlt'l' nt, alltl hC'ltilld IIII~ 1IIilltll!'. alll1
l'onsidel'uhly thongh IIf11 ulll'IIptly UIII'I'U\\'('II lWUl' tltl' l'dl'Ulllity; tboil'
IItl'iuc nl'C' Kimplc, 11111'1'011, IIllt! Hhltl'}ll~' 1III11'I(I,t! tlll'uughollt, nD(l J.lCCOllll'
deeper 1>ehiutl j illt('I'KLit~l'" hrollcl lIIul [1111111' j 1I1I1I'gi1H1111111ld1U'('1o. all11o"t
u.bHenl Ilelll' thl' IIIjll11k,
There il:l a singlo ij('t igl'l'olUI jl1lncturc 1Lt. l'lIdl Hillo of tho thlll'IIX 1>(:>(o)'c
till' 1.liddle, HII(I IIllothel' at I'uch hiwl IIl1g-ll', u piliI' nl(lngsido ellch oye
.1IIt! llIl the l·dgl' or the 11l1'I.!IWllIl, and Iwil'c' tl1111 munlx·1' at tI,t' Ox1I"'llIit,"
t.f the IIl.Ht. vCllil'nllltlglllont.
Aut('unlle thickly c()v('red with yellow pllhI.'Hl.'t'llCC', nnel tI fl.'l\ A}f'1lI1el'
11011\(:, il'()m til(' lilt joint (lnwnrdl'l, t]l(' bllKnl thl'l't' g-lllhl'ous;' thl',Y o:.::tonl1
IlIIcklVQ.)'cls til tlu' Hhnll1clel'H. l.uhruUl tl'IIIlHVl'I'Sl', Plilpi with aCllminll.tl;'
tel'mint~l 3I'ticulnt.iullii. l~yl'''I mQ(lort~tel,v !tn'gl', but rIOt }'Il'lIInincnt, r.tll~1!I
I'ObUAt; tihi II I' ,vi'lh Kpillifnr'll1 '4C'tlll' (,,.tt'rlwlI,' , tho p()Ktl'ritll' t1('X1Wl1l1,
394 'I' I'(/ IIMII'tioll~.
3182. Aphytopus porosus sp. nov, Aph!/fopus Sharp, Man. N,Z. Coleopt.,
p. 1024,
Elongate. moderutely nitid; pui>cl'lccJ)oe gl'oyiKh, flue but Ililltinot,
thioker on the elytl'a than it is elsewhere; heMl and thot'ox palo fusco-
J'ufous. elytrn light brown, nbdomen durk CUHCUUR, l<'gR Imn basltl fiVl'
joints of antennae teHtaoeous,
Heud rather broad. not perceptibly narrower than thorax, finely ~'et
cl.iHtinctly but not closel~' punctate. Eyes large, not prominent, Thorax
of equal length and brea.dth, about a third nUTl'OWel' than the elytra, itt.
baRe rounded, the Rides neal'ly straight i it ill mod~rately olosely :Lnll
distinotly punctured, but without other impreRSions, Elytl'll. neal'l~'
b'ice the length of thorax. their sllOulrlcl'O! J'(lUndell. the pUllctation II
little coarser but more rlilltant than that of the thorax, Hind-both"
elong~te, rather fine1r and distantl~' Hculptnrecl.. 15th RCglllellt I'ItnOllth
the uudd1e,
on
.Antennae distinotly puhescent; 2nd joint clongtlte hut IItont; 3rd
!nt'I'e thnn hnlf the- length of 2nd. with It 1I1!'nd!'1' hOll1l1 Htnlk; jointFl 4-R
'4ligl.t.tl~ Ililttteu; !Jth and lOth ]11l"g(.... I·, tl'llllhvl'I'",e; 11th (lvil'unll, not IlIII
long Itll the pl'~cc(ling two cOlllhiuoll.
Evident]." Himilal' to the t~'l'icI11 H},ceil's (lMan), tllll'kt!J', till' helul llOt
llihlinctly 11II1'I'OW01' tlum till' thlll'lIX, whi('h ill not anll, 111111 hll~ 110 lliM-
('oil1111 i1;lpreHlliollll. 'I'hc tlll'loUll HtI"lll'tIll'U if:! llJ'ccillcly lIlikt! ill ho1h Hpt'clCH.
I,cug-th, lIC'lt1'ly 2 1I1ll1.; lll'lJllclth, ~ 111111.
ITUll1la IhlIl!.!l', lIl'III' 1)1'111'.1', I 1'II1IIId 011(' IIIll(lJIglit tll'I'II,dllg Il'IIVI'K
oil j hi' g)'OIInl1.
• Hilt.. nat. los fitaphylulitio, dl' l'AIl~t.ra.lio ot. dom 'PolV'n~il', 11l7!I, p.118,
398 l'I'({fI,Mu,tiO/i ~,
Ill'Ollll KT.\l'lIYI.INIJ)AE.
3189. Quedius eruensis sp. nov. QUl'liius StephenK, IJucord. HiKt., cit'H
Ins. Coleopt., tom. 2, p. 84. .
Xal'l'uw, elollgatt·. h~acll\nd th01'nx Hlliniug black unll Hlightly bronzeli.
elytl'R !lnll hill!l-hl)d~' fUIiCOUFI, the latter uiten blllckiHIt nnd il'idt'SCeut;
the flllllOJ'n, ltntel'iol' tihin.e, nn,l bn81tl thl"l'C jointll I)f antennae fusc()-
l'UIOI1t1 I)l' flllvctlCent; l'~mniniJlg jllilltll dull fUIIClnlH; munclibles rufouH,
Head oval. rathel' nat'rower than thl)l'llx, bipunctate lx!hiud l\nd olao
in line with thc innor margin of the cyoo, and with a HlIlallcr botigOl'oUS
puncturt' neal' the ft'ont of each eye, Clypeul! mcmbl'nnOl1K, not l)allid.
IJabrum deepl~' emal'ginate in frout. Eye!! large, nut prominent. MaJI-
dibles short. Thorax ra.ther broader than 101lg, gently un1'I'owod 0.11-
teriorly, poste1'ior angles broallly rounded; with R morc 01' leas diRtinct
punctures before the middle. and a like number close to tlle baHe, but
.only lluipnnctate nt tIl(, sidE-H, R('utellum triangulal'. Elytrn short,
llnCllK,-.r(II' (/11111(/ ((1111 SII('('II~ oj ('olC!ClptClu, 39H
ILt!lldy twice Ith bl'olltl IIh 101lg, 111I·i I apil'cb uhlil[l1e towlll'lll\ til(' RutUl'e;
they are elohtll,v RculptUl't'tl ILIIII oovel'('11 witil y('llowiRh PUbt'Hl'ellce, Hind-
hodv ('Iongllte, 6th bogult'lll lUCladl,\' l'IHllIrlCll buliil1t1, lcl'llIinu) RtvlCR piceo-
I'1lfous. thC! ]llllletlLtioll anti vOHtiilll'e I't'FlC'mblillg thollC oC tho w{ug-caseH,
UnderHid(' 11 i1i cl, find," }lUbl·KCCll1. 1Il1l1 PUllof.ll1.e, ILl)doniell varyillg
r'I'OIll 'l"iolllc~oUH to fUHCOIlH, Auterior tnl'Hi or tho mnlo I\tJ'ougl,Y dilated,
Anlenll,lC:l IlenHcl~' 11l1hl'Hcl'nt 1'1'11111 their -ttll joint Ilm\'lll'dH, 211rl lIh
long llll :lrd, IOth Hliglltl~' ohl(l11g,
I,ike Q, Ill'lIfil'l'lIfl';/l (;J(I!IIi), 111I1·kt!l'. the hilHl-borl~' OHptlcially, and
llever with allY bl'allsy luFlll'e,
Length, 6-7111m.; breadth, <Iuite l nllll,
Eruo., Found b~' lll;l'Ht'lf ill .Jnnum','·' uno; also UllloUght !Ieud leaveN
('ulleete<1 by llr, W, .r. Guilln('''IR in MIIl'('II nUll A.pril.
3191. Lit'
~l.l
erect, in.
a.nd thClI'a~
}lulpi sOl1lowh;.
Hend ovif0J1.
brou!1, tIll' fOJ'eholLCL
unteuual tubel'elCH;
\'l'l'r dnA!:'; in fl'ont
400 1'1'1II/lnl,tionlo,
-1'11!.1'I_~~ndOl"
t ...q, Ib'
~
elo;lgate, I!lightly tl'ansverl!ely convex, nitid; l'ufellCent, leg"
d-II,rg tht! antennae, pnlpi, JIllllllliblcl:I, and tarsi fll.lVl'IICent,
~i i!. i- 6ea'l.lul'gC:!, HOll1ewho.t rounded, broader than thOI'SX, abl'Uptly con-
o , I l l ,*t€tted ~'6\thind, fin('l;v n.nd llilltllont1y punctured, Thul'u.1 nenrly twiCK'
~ l';JlIj~A loUl_~r!H broad, \\'ideAt Ileal' tho front, lUod('>l ntely I'tnmcicd ther(',
; ?:~. radl~..J'IU'l'n;\Ou ho.ukwal'lllI, baRI:! and apex Bubtrullcaw; ita Burfu('e
-.! ~,~ den~~Ib«<~illlltely Boulptul'e,l, with II. few diHtillet I'ClllUtl:! pund 111"'11
.....: Iln~~ j.9#lfjto central {OVOII, Elytl'a very Ahort, widl.'Ht hdlinll, milch
•.bl1/., l~ fOWOd towards the haKe, which iM lIn hl'M<Il'I' thall thut of
Bt!.'(, eil' lijIic('K oUlilJucly tl'unoate tOWat'I1M th(' Hllhu',); they llI't'
;ve~ Olll'Re~ bllt not clofICly punctate, !\lui h('II1' K()lIl(, Iloprl'KHC'll
, lilliI'M. "Rin<l-hllfl,v clongntc, plll'l\Il01, Iplite hulf of th(' whole
'>o.sol segtueut Il.II long nil the ulytl'l.I., eaoh of the following iour
nl)l'rowen tIlWII.:f' the bllAe, the 6th sef:,l'Jllent tranavel'sely quad-
oboonioal; ·th~,ubesooncc it! pale-yellowish, and tlle punctation
Iv coarse, bilt not close,
shining, l'UfOolcaAtaneous, moderately coarHely punctured,
;~tinct yellowish ho.il'M, Prosternum with its flanks dis-
~ by oblique sutures, the coxae situated at its base.
'$han metni!!temum,
, occupying quite half of the whole Aides of the
".n~t, with coarse facets, Mandibles as long as
'\ ,. very elongate median tooth on the inner
~;pi not very muoh ~horter than the antennae,
10ngate, the tel1m nal hll'~e, elongate-ovate,
Rlf.OUN.-Ne:1O Gell~l'a ami 8pecit:~ of Coleo:pt~lu. 4.01
J.ntennae im.ertcd on the siue!:! (If the fOl'ehelLd at bowe uibtance frolU
the eyell, lO-nl·ticulat~; hU~.l1 joint .. tout Illld nearly the length of the
following two combineu, 2nu ",lightly IIhorter than 3nl, 6th a. little
larger than 5th, 7th monilifOlID ILnd rather bmaller than adjoining
one!'.. 9t.h a.bl"llptl~' enlargcrl, bl1b'llladrnte, 10t111011ger, conioal.
LegM olongatl'. tibiae Imal'lIll'II; t.li"t.i klt'IHIol'. pcmt.llIlUl·Ollb.
Lungth, 2 min.; breauth, lnun.
Uetarlll~c, nom ErulL. .F'our ilHlividudlll of tIm. l·\.tlemely alend<'1
but interetltlng beetlo "ore found ulllongbt le.lf-lllUul,1 collect!:!11 for me
in :\1 arch, 11-110, by Captnin H. S. Whitehorn. ill "hll"'e lUIllIIUl' it lU18
ba>n nam!:!u.
Ob~.-i\l!lny ~'oal'tI ngo I bont II. specimen of the Dlbt species 1 boll
founu to M. .Albert ~" •.lUvcl, or Colen. OliO of tht; greatest EUl ope.Lll
<lUthol'itiel:l on 8taph!lllllld"e, who nu,llllld it jJimt:/"IIH brouni. It. evi-
dently reprtllOentcd a new genul., but 1 have been unable to obtaill ih.
Joscription. 'rhe llallle lJimeruB has now been useu lor one of tlw
PRelaphidae by ~'ioJ"i, and Il. copy of the diagllobis publitlhed ill At tl
:'100. dei Naturnlisti Iii Modena, 1899, vol. :12, p. 103. ill no\\ in lily
llossossion, having been written for me by Mr. ('. 0 Wntl'l'houFIe ftt thf'
Rl itish Mur.eulll.
Hroup OtiOltllD,AI'l.
Group OXY'l'ELIDAE.
3194. Bledius bidentifrons sp. nov. Bl6iliw IJt'"ch. TJIt.('ord. Rist. <i4"R
Tnll. Coleopt .• tom. 2. p. 114..
Elongate, narrow, nigl'tlbCent; heau. thorax, and elytra ra.ther dull,
with greyish setae, those on the elytra KUbe!'ect and much shorter than
the others; hind-l.lOll,v gloBlly black. nnd bea.ring numerous e1o~ate,
4:02 '1' f(/ nxartiollll.
Group PSET.APHIDAFl,
8195. Sagola monticola sp. nov. Sagola Sharp, Ma.n. N,Z. Coleopt .•
p. 134:.
Hubdepl'eHlleil, elollgnte, nitid j pubescence yellowish, slender, and
alongate, intenuingled with longer erect hail's on the elytra lind hinrl-
hod~' j hend a11l1 thol'tlX red; el;vtl'l1, leglol, II.ntcnnne, nnd pnlpi fulveH·
cent; hinrl-bod~' J'UfO-clllltllueouB; tm'lIi yellow
Head trigonal, dilated and prominent laterally behind so as to be
Ill! broarl there nA the middle of thol'ax, the median channel extends frolll
the antennal tubercles to the back, where jt becomes Ihl0a1', and there is
& narrow basal fovea n.t eaeh side of it j there n.re no distinct punctur6l'l.
BUOl S,- XI'''' 0,./11'111 IIl/d Sllt'I';" III I '0]('111'11'1 a, .t.()3
EYeR ovi!lentl~' 11l'lllllilll'llt. 'l'hol'lI\ Ill' ahout 1''1l1nl l(,ll~th ;tlld III'C III lth ,
"illel!t 111111 Hil'lIll~l~' ,'ul11lClc,tl jUKt 1wfol'l' thc miclrllf'; uh1il[ucl,\' lllLrrowed
ill fl'ont, \\ hlrc it iK ollI,\' hull' the width of the hoarl, \1 ith a ueel' rovel!
tit ellch Kiclo c:..toncling frolH hl'llind the miclcllc to tho basnl 1Il1u'gin, tJIit-,
part tlt.,l'cfOl'O i .. llJuch COllil'.lClccl j tho clOl'Hltl fu\'on behilld the eellt)'u,
though llll'ge, ill not h'lIllMVCI'bC, thcl'c ill II hlll-1II1 JIll actUl'e at "lLrh Hicle
of it. l~lrtra m'II)'I,\' twicc til., long-til of ihe tllol'lI \, gl'luluaUy nnrrowod
tu\\'o.l'lIR yet uI'oucll'l' at tho baKl' HUIlI tJll' tllol'ux, hlightly OIl1'VO(U~'
1111 l' row(>cl hl'hiucl j Imtul'ltl tltl iue wdl llial kod; the intl'uhll11l(ll'lli
imprestiionH couKiHt I'tu'h 01 a bmmll'1l1wlm'c' IIlIel IL lllOl',' elllllgu(l'd fovea.
Hiud-ho(ly l\ tllircl Iongcl' Chilli IIIl' ('I,vt I'll, illCliqthctl,l' 1'1llll'tatc; the
bllHo.l three Rogll1cntt:l bl'olldl~' llllll'ginod anc1 uf about cCllHll leugth; the
tcrminal oueH, eomhincd, ohoollicul ancl MllIlIiullte, Itwl much Vlllel',
Lcgtl ('longute, Kimplo; 211c1 btl''!lIl joiut cliHtilll'tl~' pl'ulolll.\'ccl llIult'l-
nenth,
AntenlllLc elongato; i.>uHltl juiut l'ocldiHh, e,l'lillcldc, ILbllOHt Ul> lOllg lI.~
tho following t\\O together; 31'cllclllgol' thun bl'ullcl, hut clilltilletly nlt)'l'lI\Hll'
than the IIcljoilling CIIWH; .(.th l111cl ;jth cCl'ml, longel' than broud; 6th
Hlig'htl~' h1l01'tOI'; ith IIIHI Htlt tJ'lIllClth' lit the [lIIHe, It little 1l1Ll'J'o\\cd
ILpir(tlly; Hth lIud 10th tl'UllKVt'I'IWly 4 11 ILl lmtl' ; (Itll c(lllimi IIlId IlCllllli-
illite,
Whou lllllc't'cl Illollg~icll' 8, 1'1111111'1111 (~i~+) thi~ hPl'cit'h iH HC'('J] to be
Jell.. I'OhUHt, the IWIlt! iK 111)t c(uih' tt'lll1Cllh' ,Li the 1)l1I0il', itb hillel Ilngle"
1I1l' lOMI! PI'Olilllg't'Cl olliWltl'Clly 1I11cl 111'0 Hlightly cl('flt'wcl, tho hllhUl foveac'
lUI:! smallo!', but tIll' <',n'H III'C WOl'C })I'o1llilH'nt; the thOI'll" is lIIU')'OWOI'
IInc! the elytl'IL 10l1gCl', IIUd, 1110 I'co\"p I', tlu' clillltl'ci hiu(l 11Ilf.!ll'" oj' th(,
h.'ne{ arc not concuve tUldel'Ilenth,
Female,--Genuo lIcnl'l~' Htrllight 1,t'llilld the l'."\'h. with ubtul!t!, but not
Ilt nIl dilated, pOF!tel'iol' nngleH; lightcl' ill OOlOUI', nnd with lllUl C pro-
lIIilll'ut oyeq tlum th(' HUlllt' h(':" of 2724; tIll' ~icl(,H of thc thOI'II:" II 1'0 leRio
,lilatccl before' the micleUl'. aut! tIll' It'gll 111'1' 1I1OI'C Ml('IH{CI', On till' 1l1lc1l'l'-
or
..ir1!' thE.' h<'lHl th.'!'p iH It W(On-llUII'kccl tl'IlUHV('l'MC rlt'I)J'(,F.Hion.
IJClIhrth, 2! !lllll, ; bl'eaclth, ~ Tlnll,
:\![ount NI-\'11111'uhoc, .\ toIing-h' lIIall' Imcl hl() felllul"H CuulJcl ,llllllllg'Mt
,1"c',I;Vl'rllullveK which \\'I'l'~' gathm'c'cl fill' lilt' b~' Mr, W, .J. nUinnCI!H,
0"11,--2724 WIlK tliHO()V(lI'l'(l :It 'J'ILI'ukcngll. IWIlI' HotOI'UII, nncl I fount!
III II' lIlulc' lit Waillllll'ilio ill .TllII1IIlI'r, l!lml ,
:11 {IS. Euglyptus foveicollis Hp. n()v. atlg/tnlltJ.R Broun, Mall, N.Z.
(Joleopt., p. 1,111.
Kluudur, Klightly ('(mVI')., Kltilliug; I'UrCHl~; l11ytl'a, leg", IUHI IUltOlllUII.!
fulvcscl'lIt; pnlpi, tarKi, Itllli tOl'lIIillul juill! oC IIl1tenUllt' HIIV'I'IolWlIt ; pUIK'l4-
('t'1I0l' g'I'l',dKh. HuhCl'eot, l'utht'l' IIC8.nt,V,
Hend (lhliCJIU'l~r lUu'1'llwc!1 htlhiucl, whol'(' it iK Hlig-htl,r hl'(),IIIl'J' tht\1I
tho IIpC:" of th,' tllm'u'(; illelllcliug the largt' ~Y'~I!I, '1lUdi \\'idl:'r; it iH.
Pl'opUI,tionltll,\', lIIoclt'l'uh'ly COU.I'Hely }lum'tute, with 1l.11 illCliHtiIlOi mecliul\
,hia b£.hinc1, tilt' P"()llIill('Jlt Illltelllwl tulwl'cleF! HopllJ'nted h~' Ii diMtinct
hut not widc 011ll1l1l1'1. Thol'lIx slightly longer than l)1'ollel, \\'icleflt amI
obtll~ly 1l1'()lIliuent lit the middle, llUlI'(:' narrClweil ill fr(lut thal1 behind;
a. trnnsVel'!4C illl(ll'eMMi()n nUll!' the bll.t:Ie O()nncctK the clung-atl! lo.il!l'al foveae j
i,he ilillouiclnl fovell ill clozl~nte, but doeA nut l'ench tIle bllRII.l impl'eSsion;
it!! punet.ition iF! iJl(1iRtinet, Elytl'lL ncarl", twico the length ()f thorll)',
v~r~' little hronc1(,I' thun it iH Itt th(! lulse, wit1t'l' 11(,1\1' the micldle, t11(oi1'
4-04- 7'nWRQcfIOWI.
tlldeh gently J"lllllldecl; thl'.' lIJ"e not l>l'I'cel't ibl~ PUllet utt'; the' IIUtUJ" II 1
striae 8re rather fine, but are foveiform and deep at the baRe; the intra-
humet'"l impl'e~sion on enoh ill alMO foveiform, so thllt the Khoulder nurl
~he interval between the illlpl'ehHion Imd the:> sutural fovea nPP('III' elevate!!.
Hind-body flhorter than the elytl'n. nAI'I'o\Vod ann ne.fl.cxed pUbtoriorly;
the lRt dOl'Ral Regmcnt horizontal, Witll n Ollrvnte hll'llll dtl}lI'c~Rion; it ill
I'llightIy IOllgel' than E'itlu'I' the 211d or :Jro.
Antertnll.e elcmg,dl', hlLbal joint thioker but only a little IOllb"el' than
the 2nd, joiutH :3-5 tiuite oblong, 6-H hardly IlK long 0.11 bl'uud, 9th quite
douhk the Foize of tIll' 8th; 10th cvidently broa(1el' than 9th, laxly artiCll-
lclotecl at the bahe, but elo'll·ly ILpplied to the] lth, wlliell is large; conical,
and acuminate; th~ 3 terminal al'tiouilitionl.l 00111' numerous slender
but elongate hairM. a.nd form It well-mllrkerl cluh
'fhc thoracic Roulpturc and abbl'l'viutl'cl Illllllel'al impl'eKSi()nll nl'c von
clifterent frum thOR(! of E, efeg(lIl~ (2460), Ou the othl'" hund, it Il~
not agree very well with the type lIf tho neal'ly Illlied genus lIIirrot'!lruR
(24:61). 'lhe position assigned to it Ilncl the following IIpeoics illl there-
fore between thpse genera, ThC'l'C ill no othel' "~'st(,Jnlltic place lot, them.
Louj..rth. If mm.; hl'('l\dth, i 111111,
Retal'uko, neill' El'll11, Describell fJ'om a sillglo IIpeoimen !,il'l"ctl (lut
of decayeo. leavelil collected fo], me in March, 1910, by CaptlLin H, S
Whitehorn, of the Geological S11l'VllY Depnrtlllen1.
Heul! cVlIleuth' ",,"unel thll.n thlll'u,.. rmmril'el hchincl, .It't hloaril"l' thore
t.han tho thOI'.IOIC apt·x; bu~al £01111111' illlli~tilld. the bma<l IIIl'eliun depr('I>-
~ioll (·"{tend" flll""Ul'elz.. J)('tw('cn the' }Il'Ollli 11(,lIt Illltclllllll tnJ)(,l'clt'K; the
~culptllre inclcfinite, C()IIRh,tillg lL}lPUH·Utl.\ 01" ,,1IuIll gl'allllle'~ wit II .1 Illillllte
punctul'(' in l·arli. I·~yc.,., Tlloell'I'ILtl'I,r 1al'l,!,t' a1111 PI'OllliJwlIt. \\ltl1 ('(,,11''1('
[neets. 'l'hol'.L't z..nhuviform, oi' almut !lrlll.II Il'Jlgth .lUd hll'uelth, l'athel
widcr hl,fnl'l' 1I1l' 1Ilirlrlle thnn l·Ih(·'\!ll'It', l'mJnrll'r! I1U,'I(', .1 good c}onJ.
n.'ll'!'o\\ed .1IItCI iCll'I,'; tho mCbillllongiludillul "'1110u... ill \\('11 1Ill\1'kcd, allel
ends in till' bllHltl i'Ohl'4ol, which iH joilleel to tIll' l·}Ollgltt!· fOV(lU at oach
,ide hy di~tillot gl'()OVCK; itR Houlptlll'(' h, 1i!,(' thut oj' till' h(lad, EI~tl'a
II third longer thun tlulI'ax, 1'Ilth('·I' hl'oltelol' th'1I1 It ib at tho bar.e, with
r()undcd Rllouldol'h j upiocs Klightl,r ublique tOWal'(lIil lill' butu1O, "itll
minutE' cli.it,tnt b('Ulptule; buhll'ul .. triao elc'ap, iutJ'J.hulIIl·I',tl imprl'RSilllU,
hlLol t, {'acil Cl~·tl'Oll tl'i}'UllOt.ttl· ,It the baM', tIll' punctlll'ch, ho\\t'vcJ', ,11'(' III
tho stl·iae. llill!l-hody oIDll11l'l' than the (,'1,\ tra, it,., bat~lll (hl'u(l SCJ.\IUOlltb
nClu'ly t'quu.l, tIll' I'lt "lili a rll'l'P tt'lmllvel bO b.IK.ll iIllPJ'C:'R"ioll, ",ith ('ILI'I-
nate sides, tho 211d ",ith It Ailllillll IJut morc lillCfll' illlPl'Ohl,lioll,
LegK elongatC:!j ,lutel'iol' J'OIlUH'1L thickel' th,.I1l the (lthel'll; tlbul.l,· UII-
'lrmed; nIl lIlClfiel',ltl'ly IIrehed out II IlI'cllJ, i ho h out puil' with JllIll'(' CIIII-
RpiouOUIl pubehcelll'e noll.l' till' l!,xtll'llllty .
•\utUllllUU '1tout, bellring mtller ('lulignto pulx·t!CL'll('Oj tlu'il' ~ud joint
ullnoKt UVILI, ,Iz.. thick and long ILK till' 1l1l<'COVl'l'l'cl POl'tiull of the bll.llal j
3rd nearlr aK IOllg &.II bl'oad, benel-lik(·; joillh. 4-6 tJ'UUbVCl'M>, diifcl'illg
but little; 7th and 8th abl'uptly bl'uallol', Utlhynullotl'ical, thinnel' ut thc.'
innel' than Itt tho outel' ilidetil; !lth and 10th tl'U.UhVCl'KO, htill 1ll0l'U ell-
larged, but uot double the breadth of the precoding pail', uoth broad a.t
the batle but muoh nu.l'l'()w('d in front; llih conioal, IlII hl'QtHl 1114 the 10th,
but not 8.11 long UK the 9th u.nd 10th cOlllbint,!fl; nt itK hURe. elII tho inRi<lo,
there ill an obtuse tubercle.
1hel'e is no neceRRity fol' compul'iI'IOn with otli(' I' Kpecies, 1111 tho very
remal'kable nnt('nllat" IIl"C Ilhlllldantly distinctivE'
a, Length, 2111111,; bl'E'....rlth, ~ mm,
Mount Nglloul'uhC)e, Om' inclivieluul. piokl·tl Hut oj' II hug-fill Clf IJoou.y-
hl~ lenveR hent to me by ?II', \\', .T, Hninllel::llil in l\fnl'ch, HllO,
.\ni\!'ulI.lll "jth hh'llth'l I'Uhllhlle1l(lC, tIu'it" ohlong lind joiut not 'lulie .LI>
"tout hilt ahout nb long u.. th", h,t; 3rd oboonical, slightl) lOllg(,l' thaIJ
hlOnd; jointR 4-6 becHI-lik(" lIud of about th(' flame length nnll hl'E'adtb. j
7th ano. Rth flliAhtl~' lIll'gol'I, 1I1Il1'owen t(l\\ ,lrllR tb.e n}l("x; Hth amI
10th rathel laI'At'I', huth 1lI111C1\H'11 IIJlictlll~'; 11th lI~T'A(>Ht. ('011 It'a] , sub-
IlCIIlIlinnt(".
lInderH"idc "hilling \\ith Huboi oct gl'cyish pl100SCell('o. lIt'llel IItudded
"Ith minute' lmt quih' (lcfinitl' gl'fUmltlR, nm1 hI'll.l'illg NO('t ... It'wlol' l!t.>illtl.
PrCll:lturnmu llIcdinlly cal'illnto. .\.ntCl'i.11 ftllll!ll'1l hroadly gl',)(Iv,'(I, nIl dis-
tinctly pub!'Hceut. Ml,tuHtel'l1 1III 1 I1Il illl}ll'l·...HOd. .\ htlOllll'n t'lclllgute i bllS.!l
segment hut littl!) ex})oRI:'II, fl'ingeo behind; 2nd and :JI'd about ,..qunl,
tllloCh longer tliltn 4th OJ' rith i 6th linclr amI tli~hintl)' pllllctnreJ, in the
llliddle nCcLrl~' ollllblt, thc longth of tho lith, with 1I Rlennel' basnl mlll'gin,
lind, ILt the apl'X, with 1\ mtldian semicircular cal'ina.
At emee <'l'pnrllhll' 1'1'0111 R, llt,trmrtlll'lIlI b~· ih, 211el .11ltellllul jui1lt bt,jng
thinnel' than the IHt.
IJclIgth, l~ Illlll.; brl:'lLeltlt, eplitt, ~ 111l1l.
~rlla. 'I'hc tYl'iclll Hpecimcn Will! found by me in January, 1910, and
h\o 01' three otllt'nt W(;'1'(, pieT,eel out ur leaf-mould cIIllected two month ..
llfWl'wftrllF, h~T Mr. W, J, ({uinne-h. In two of the.lle Hpecimens the 9th
IIlld 10th jn1JltH or tho antennae nro Ulore tl'amvel'Hru IllId not dilltinctl~'
1UII'I·UWt·O npil'II11,\·-T)clIs~ihl~· II 1'01''(11111 eliNl'nrity nlll~·.
belor(' the middle, mOIl' 1I.II'I.,\\ocl ill C.'out thun behind; tIll' lIle(lilLfl
groove is dcep lind extendR into the bu~nl fOI>Ha, which iR unitl-'d to the
elongate lateral im}JrcRHionfi by h'ltnhV('1"HO Rtriac, itH sculpture seems to
bl' fim'ly SUhglllll·\l.II', E1~'i 1'1l, ut lho hUHo, hl'(llLdl'r thall thol'llx, II thit'd
Il)n~,'l': without pel'ceptibl(' pUllctntioJl, Hlltlll.11 "h'iae deep; liol'Ral im-
pl'cl>HionK deep nt ihe ba~o. I'llther broull, but hocoming indiRtinct bE'fore
the lIliddle Uintl-bon.~T Ill,tI('l.ed pORtoriOlly. mudl Rborter than elytra,
itR firRt vil,;ihle Hegment 1l()rizoutal, with a tram,Vel'hE) basal impreRHion.
Legs ver~' eiongttte •• the t1"outnl pair~ plll,ticular1:v; fl'mora rather
Mendor, anterior tibiae moclerately curved externally,
.Antennae elongate, with Klelldel' vn1x'hcence, hahul joint cylindric,
"tout, red, distiuotly punctate, m'o 1'1y 110uhle the length 01 the oblong
2nd; joints 3-8 oblong. 3-5 slightly lal'g('l' thllu tIlt.' thl't.'e which follo\v;
9th and 10th ohllnt ,.'qunl, as long all they m'o bl'mlcl. nUl'rowed apically,
(listinctly, ~'et not very much. Immller tlum tlw lith; tcrminltl I'longnh'.
llonical. IIcuminn.te, nnd about as long as the 9th Ilnd 10th combined,
The l'emlLrknhl~- elongntl:'. B,lIroJ'iB-like k'gK, l'tLtht'l' ",hllrt de1lelo.ed
hind-body, Iluberect elytral vCl.titul'e, and the peculiar sCIlI]lture of the
head and haHal joint of the ILnh'nnac m'l' hldfiduntlr dil!ltillctive,
Leugth, :.Ii 111m,; breadth. f mIn.
GJ'eymlluth, A hinglc inllivi,lulIl hOIll }.oIl', .J. H, Lewis,
Byraxis Reitter, BeitragI.'" ';Ilr Kllfl'r-f8UU8 VOll N,Z, Vt'rh, Nat. ver.
Brunn" vol. 20.
Near lIubgenus Reiclumbachia. Body ra.ther shurt and convex, An-
tennae lO-ltl'ticlllate, robust. distnnt ut belHe, almust biclavate. Maxil-
lllry palpi Iluadriartioulate, u'l'millal joint fUfliflll'Dl, narrow, Basal
joint of POlltorior tarsi short, the following two lnnch elongated, with
Rillglc cllLwiI, Head obsoletely bifovc()lnt,·, Thoru::\. tltnlloth. not .tt aU
foveoillte, El~·tJolI without 110rslll IItriac.
3214, Byraxis monstrosa Reitter. Verh. d. Naturf. ver, Brunn" vol. 20,
p. 197.
Itufu-ralltIlDc(Jnb, Hhining, IIllllloth; extl'l'mity of 1l1lh'lIllu,c piccous, legs
I'ell.
Relit! llt'IlI'lv qUMlt'nu', 1I11ll0Mt piUllO, with 2 HhlLllow fl'ontal foveae,
Tho1'8.x about I~h lllug IIh hl'unll, c()J"Ilifol'm, "mooth. Elytl'a HIDooth, with-
lIut dorsni grollvCK, l'Iuturlll Ht !'iIL(.' vel'y litlc.
Mf,zll,-Anhmmw l'olmHt; Imlllll joiut stout, oylindrio; 2nd quadrate,
lIlLl'rowel' thnn lilt; jointH :l-H i1'lInHvl'I'Klll, '7th ILIIII Rth aplll'CCiahl,"
widenerl; 9th and lOth 11l1'gE', llnovt'lI, a.nd pubeHcent, the former acutely
produced outwlU'llly, the lattl.'l' very thick, nearl.v suhlunute inwardly,
140 thut th~ extl"·lllit~· IIr th(> 9th joint enn he I'ccciVt'll ill the hollow,
PCllllltillmtO vl'ntI'lLI lIegmcnt (looply impl'E'HHtld. th(' basal st.roDlCly
foveolate lUedially. OI'~Btl'd aud biilll:lciclIlate latel'all~"
Ll'lIl.,>1h. 1'5 ltlnl,
O"l'\'lrUHltlt, M I', II, Hcllllfl,
lI~ad lle:u'ly UI> lUI g{! Uh 11101'11:'" oill'lol{!tel,' biJ'ovt'lIlule ill fl·out. Eye,.
hlightl,· prolllim'nt. \\ ith ('oltl'I>e 1'1I0('t". 'fhol'!LX of IIhout t'1111111 Itmgth
lind 1,j·....vlth. wideHt jUHt h('fol'l' tho llliddlt·. finely !lulI'gmoc1 and feebly
lJiHiullotl.! lit till' 1111hO, \\ itilnllt illlpl·l·..."i"lIh. l~l,YtI'u 1Il'IIl'ly thrioo tIll!
ll'llgth of thOl'll\, .dightl,' h"OIICII')' at tilt' ha II{) , tlll'il' Hiell'H n liltlt, 1'I1IlIHlod
Ill'I\" tilt' hillll thi~hl>. with IiUI' ImtUl'II1I'lll·illt'. Hillel-hIl11," cll'tit, \(\(1 , \,ith-
lI11t \WU-IHIlI'kl'tl HCulptul'e. I.egl:! Hlew1t'I·.
(huh'I'siele uitid, l'UrOUh, \\ ith HUInI' lim.' g'1'u,ritdl PUhI'HI'('IIOl" Motu-
btt'l'JlUm hl'OIL(1h' modially del l1'l'jo,Hed , tho Hidl'R 01' tilt' clcpl'cMMioll 1I0ml'-
whnt eioviLtol1 IJuukwnl'dH, BaRll1 velltral segment hLl'gl', fhLttcnecl behind,
with a dillt inct tuhl'l'('ll' ('IOHC III l'IU'" 01' til<' ~()l.:ll·; Hl'l.\lIll'lItH 2-4 vel"
short in the Illiclclle, the lith with II 11l1·g'1.' fovoll. .\Jttt.'l'iOl' trochanterl\
spined .
• \ntt'TlIlIW LO-lIl'ticnllltl'; hllhlll joiut )'ItOllt, 1'~'lillc11'iI'; ::hul HillLillll',
bllt shortol'; al'll l'athl'I' HllllLllel' than ~lH1, and l1ILl'r()w~II towards tht'
hase; 4th 1'I011l('whut lIloniliforlll, un(l Hhort\'I' than Ileljoinillg ones; Gth
Kubqllad1'sto; thellt' 1I1'tiolliatioujo, hell I' ~IOlld(·l' pubI'Hc\'Il('(' ()nI~'; th~ 6th
and 7th conlCK(l(' un (I 11I'l' Iml'dl.,· (liHtinguiHhnhk' from Mcll other, except
that the luth·t' IIUH IIhClI't ,n't clil<tillt'i ),I'II"H," HI'bll' Imel ill Hlightl~' obtul:!ely
hulgiup: lit OllC Hille; Rth oxtremely Hhol'l, l:IometimeH overlapped by itl:!
pl·elleot·HHOI'. 1111 1I11'l'C I'IIthf'I' hl'01l1101' thllll the lith; 9th very large,
stnight inwnrlllJ, longel' ollli gl'lululIlly becoming broader outwardly,
itll frunt llllg'lo Rlightly cl1rved Itlul olaspitlg tho bUH4) lit the 10th, itK
I\pex ohliqlll' lind HliA'htl~' l'(llIl'U"I,.'; 1Oth ('onionl wlll'Jl l'xtlel'tccl, Imh-
rotunclutl' ",11('11 cl()HCl~' Ill'plic(1 to the 1ll'II11lt illlllte, Ill'arl,\' 1111 hroad all it
is; hoth of these bellI' finN' Metlle than the 7th and III 0 finf'l~' out quih'
11efinitcly granulatc.
:Malformation of jointH 6-R of the antennae rendeI'll an accurate dl'-
Kcription of one male inapplicllhle to tlt(' (Ither. It mUl'lt h<' placed next
to 1645,
Female.-.Antcnnat) ll-ILl'ticulotl', jointl:! l-!l like those of the male.
their 6th joint of about the I'Ilimc Flize ItS the 4th, joints 7-9 transverse,
10th brollcjcr than Hth, 11th llenrl~' twiCl' al'l long all hr08n..
T..ength. 11 mm.; breadtll, ~ nl1ll.
l[ClIllIt Pil'ongill, 1 r0l11l(1 two mull'H an(1 tlll'l,(> fl'lllllll'H in Decembel',
190fl.
Hl'onp Hxr,I'JJ1DAH.
3216. Choleva caeca )'Ip. llOV. ('holt'1.1(l T_utl'lIillf'. MIUl, N.Z. Colt-opt."
p. 151.
Oblong-oval, nitid, pnl>l'iiccIlC~ tll\Jll'I:IHHecl. Hlonder, pII.I~ flavescent;
fURco-eRNhmeouR, heael tLnll IC~H l'IIfl!OOCllt; tursi, palpi, l\1ul hllsa! three
jointll of antenno(' tCRtIlC("()UH, I'~mllining joints :Cuscous,
Head small, widl.'Ht II 11 el HOl1lewhllt angulate ht'hinu thc middle,
nal'rowl!,l alltel'i()rl~', lllotlcl'atel~' COal'I\el~' but not clol!el~' puncturel].
'rhol'ax large, the bl'l.'lIrlth noorly double tilt' length, Inuch curved IIn-
tel'iol'l~'. bltl!!e tl'Ullcate but ''I''ith its angles I!!lightl~' ovel'lapping the
shoulderH; it ill lUuch mOl'e finely pUllctate than the head, indistinctly
on the middle. Scutellum tl'iangulal'. EI;vtra nowhel'e broader than the
th~rax. Ilnd quite tw~ce itl:! l~llgth, ~l'lullllLllr nan'owed posteriorly, the
apices not lit all aU1llumatc; with finl.' Imturnll'ltl'ilLl', and 1II0ciernteh' 01011('.
tl'lI11IlVCI'Rely Ri:l'igoRe sculpture. •
.\ntennl~e ILII long Itfl head and tlwl'u.x, g'l'lI.tluull:r inol'assate, Dae1y
l'ubl'llt"Cut; 2nil joint rathor longer than ~h'ci; 4th a'nd 5th longer than
bro,"I; 6th l'I1lhqluuhll.te amI n littlo Hltllllll'1' thu,n itll, buth I'Ihghtl~
nnrl'o\\eLl to\\lll'dlol the biLHo; Rth vc!'y Hhul't lind tl'Ul1HVtlr~e, but .11'> hroat1
'1M the 9th, \\ hioh, m, \\,,·ll 111'1 ill(' 111th, iH h'1lII HVl'l'Hl'1," Ilulttlmte; llth
conical, hwgcl' thun thc p!'ec,'ding,
Tibinc set INC. the intel'ulol1inte om'vo.tl'. the I)O~tl:ll'ior distinetly bi-
HpinuKC Itt tiLl.) I.'dl'l'llIit~" "'III'Hi C'longnte, thickl~' pnheRcent, tho antl:lriol
'wt Llillltecl
The e~'cH IU'l' not t1iHCl'l'lliblc nhovu, ~[axillltr,v pulpi "ith till' PUll-
nIt imntc urt iClllati 1111 uht'Ull i"al nnt 1 I'll th"I' llll'gc. 1Ill' t"l'llIi 11801 IllllaU IInll
ItCnmilluto.
Mo~t llellrh' 1'C'F.t'lllhh·H 2i:i4 IIwi :Ai:iC;; hoth 01' thCHt·. hOWCVl'l', ClIlI he
t'u"il,\' dillting{liH1wd 1))" thei!' pl'oTHhwnt e~-eH,
~, Length, 2k mm, ; bl'elldth, Ih lllm.
Mount Ngaul'uhoo. My Kpecilllen T owe to tiLl:l kindnosK of ~ll'. W.•1.
UUinnel:lR. ",hI) coU('t'il'tl tho loaf-lIllllllll it \\ at-. fonnd in,
321A. Camiarus estriatus lip. nov. flmninrUB Hllllrp, MilD, N.Z. Ooleopt ••
p.148,
Knhobloug'. Hlightly 1..~IIIVI·l., niticl. lIigl't'KCC'Tlt. leg" 1\1111 IUltcllnoc
OhHCUl'C I'll fmlM, )l1l1){'HC('X1CC ItiHtincl,
Hl'ad tllIIlI11, HlUonth, with a fl'w VI'!''' "lender g'l'oyiHh l"til'H. ~yeK
pr()minent. Thlll'ax II thil'III)I'()I\tlt'I' thlL;! Inug, wi.it·"t nOlll' t1l(, l1lillc\IE"
well l'tHllI,letl IIncl Hll1t'll 11111'1'0"'('11 111It,'I'illl{", H1ight1r t-.illllut .... llllgulltlltt'
~Hi
towardb tilt' l'ud,lIlgulul hlll<i nuglt',,; thl.' cll.'up n.ll'diulJ l,h,11 1Ill'l, v.hiuh
is expanded behiuu, cl.tc,llId~ Uh1lOb!, 01' quiLo. £ru1II bUbl.l to U}.JO.ll.; at
I:laeh hicll> 01' it, ()n tlll' rui(lclle. thenl ill a \'\oll-ulIIl'keu P"llctUl'e. alld 0.
"maUl'l om' IIl'l\!' tho alll'\; hahnl COSMII.' doop unn Ulodorutely ('longate,
wIth ,I t l',lIll-oV'I.')'1!O sorioH of 6 punctul'oll behH'l'n them; along elLch side
fr(lll1 tho pOHiP)'iur lingle to l)l'~'ond tho lIlidcUe thm'e 1.1.10 Hl'veml small
}IlUHltUJ'O"; it hours JlUlIlOl'OI1S elongate ILl:lhy IlIIil'S, ~~lytl'lL oblong-oval,
with C:llrvo(U~' llUl'l'Owetl "houlclel's, so that thl' hUHC' h, (lIlly a little broader
thau thut of the tlwrllx. which ih hal'clly hulf tlrd I' length; thoil' !>onlp-
ture ill irl'l'gullu'. COIlHifliting of, on each. 6 d(lrsal rO,\,\,H of vel'y unequal
punctlll'l..'M, fIIollll' Ul't' elongllto tn' oblong lLml othel'H vel'y dilltnnt from
each oth,,}, jllst at the base lIomo almost fUl'lIl strill.El; they al'p clotheu
with subl'l'oot, very elongate, Hll'lIder dnel'oolill hairH, and many COIl-
Rpicll()Uh whittl nnE'H Ul't' intcl'mingled,
AntennJ.c I!tout, finely Hetoso; the blLlIlLl hb, jointll al'c subcyliudl'ic,
and cliilel' hut littlo. the lKt. however. ill thickt'l', 111111 tho lith mtllel'
lhinner than the contiguous ones; 7th )'lltJler broader than 6th; !oIt1t
~'vid(,lltly tho smallest; joints 9-11 about alii bl'oad as the 7th.
Male,-1'llorsi ank'rio)', with tho haHal tIm'!: nl'ticulntiOlIK uila.ted, tJle
1st lurgost, intf'l'l1Iediutt, piliI' l-oilllplo, :ith Vl'lItl'III Hl'~"Il('llt illl'llrvocl at th('
npex.
Rather bl.l'gel· thun {/. "wI,tIl'ira (~70), which, h(l\\,O,"OI', OIUI be tlllllily
I'eoogniz~,d h," tIll' VPl'~' rl'll'ltllll'l,l' Ktrill1.t'-punctatc plytrn anl1 well-Illlll'ked
interlltices,
1 }.JOSSCIIIl eight specimens of the pl'l'l\('nt Al'l'ciell; two were ,*,cured b~
ulyseU, till' others were iound in 1('l\I-1I1011111 M'nt to Ole by Mr, \y, .J.
r,t1inneRR rIming March ann April. 1910, The:v Ul'e ver~' homo~('nlJO\1",
Length. 4~ llUll.; breadtll. nearly 211l1U.
Flrul1, lICllr Waimul'ino; altitude, 2,50(1 ft.
:i:HH. Sllphotelus obliquus sp. 1I0V. NltpltotelWl HI'OUll, A.JlIl. Ma~. Nat.
Hist., 8E'1'. 6, vol. 15, p. 83.
HlOI:lKY, oblong-oval, Hlightl.'· l'Clllvex. h(,lIrillg only 1\ Co\\' minute Ol'toot
groy setnt'; flll!cumq .Hiclel of thCII'a \ IIl1d the' KllOuldel'" (Ie It pulor and
morc rufeRcl'ni hm.!; lhl' Mi(l('M of (,h'll'll hl·hinel the mitltllt' iul'lcc)-
teMttU'l'Ol1h, '
Hend slightly bl'otulel' tiulIl Il}lex of thOl"U.. vcr,l' l'vidl'lltl~ tlll(1 brClucU,"
clepreAsed b('tWt!Oll tht, Vt'l'WX IUld the ('yell, withuu1 viHibl(' sculpturo.
E~'l'll nearly flllt. occupying more thun huH nf l'l\llh Hidt' of the head.
jUht free IJ'nDl tho thorax, tl'uncl\te }J('hind, llIu'I'oWE>d unteriorly, witlt
moual'ate ftLcetR, Thorax h'n.IlllvcrRe, the sidt'll very (lilltinctly ml1rgined,
gently ourvedly no,l'J'owed towul'ds the Hubtruncl.I.te apex; the bnse closely
ILdallten to the c1ytrll. feebly me(linll,\' curved and sinllate tOWIll'ds the
!lidell, itR ungk'S JURt 1't'L1n.ng'ular; itH RCulptlll'e vcry fine anll htlrdly
discern.ible. Scutt-lluIn large, cUl'vililllJarly tJ'iallg'lllul', }i~lyt1'lL oblong.
just n little bron(lel' than thorax lit the bUKe, (Iuite twice itH h)ugth.
oblique townl'ul:I the obtuRe apices, the latel's.l DlIl1'gins much thinner than
those of thl.' thol'ax.; their pllnctation irregular, rather nne and Ahallew,
nowhere close. the Ruture Khal'ply defined, with obsolete striae,
Antennae inserted at tlle Aides quite cleRI' of the eyes; basal joint
l'ylindric, mightly longer tha1l 2nd, both fnsco-tcstaceous; 3rd 11101'e
Hlender than 2nd, and neall: as long !loll it ill; joints 4-7 longer thun
il1'ond. and nl)()ut equal; Rth ~lip:htl~· bronrlel' thnn 7th. not abbreviaterl;
RUUl"N,-.Yeu' (Jelltrn, anti SpeC1't8 of C.:rI!Olltel'U, 417
lOth trllnbVt.'I be, "hOl'ter Ilud IHClllllcl thlln !lth j tUl'luinul largest, bub-
l'otunu'lte,
Dmh' \\ ingell. Pn~idilUll covered, TlLl'hi seemingly only 4-jointed,
but I think thl!Y Ill'e pentnillerom., with till! trUtl bUF.al al'ticulo.tion SU10Jl
tlnd intliRtin(.Jtl~' IIIU I'ked off; nlc l)ll~ul three joinbl of the front pah'fl
lU e slightly dilated,
The ]aJ'ge ~entellulIl nn.l ohlit(ue pOlltel'ior pOJ'tion of the elytra dis-
tinguish it f,'olll l!757, thc trpical species,
Lungth, q n11l1,; bJ'cndth, llllUl,
Gl'eytllouth, A e.ingle intlivillunl, iJ'om Mr, J, H, Lewis, October,
Hl09,
(klllll' VULYDUllAll,
3220, Syncalus explanatus Sop, nov, Sy~u:al'U8 Sharp, Man. N.Z. Ooleopt.,
p.200.
Convex, obluug,ovlll, bul.J0ll1~(l11C; fm,ou-piOl'OUt!, the front and side!.
of thorax, Il.h "ell Ull the ll'gt., ni>bo1ll'ely rllfuRCent j tarsi and antennae
of It pllltli I'otl; thc hl'tuU ~'l'U()wihh, ruther tillc, el'cet along the e.ides, 011
the after pill t I)f the bOll,'" HIltl lIll the tihille j on the elytl'a, near the
blltuJ'e, tiley Itrc (]epI'l'Ml:Il'tl und fillt', but bl!YCllltl al'u il'l'ugularly concell-
trated, without, hO\\l'VUl', furllling rlil:ltint..'t tuftb.
Hond with fonb~I'nn111nr l:Ioull'tIll'e. l'ui!l'mbling shOl't rugae behind.
't'}1(II'ax nelLI'Ir twicc Ilfo l)1'luul Ilt. long, diKe h'lllllolvcl'Hely oonvex, the sides
(':I,pIannto 01' tinttl'llt!tl, ~lightl," I'!>uncled nnd more nal'l'owed in frollt
than behind; tho UUh'1'i1l1' Illll.\lch l'l.h.'l1l1 ab fill' us thc fl'out of the eYeI:I,
thu postcl'illl' al'c nenl'l~' l'eotllugllitu'; diMe It littlo uneven, Witll in'agulal'
&.olllpture omlHiMtillg (If IUl Ilumixturo of KHlltll flattcned gl'll.nules and
short rugae. EI~·tl'll (If thE" hl\lIlC width nfl tlWl'llX at the base, twice
its length; their flcIlII,tUl'l' BtllIlc\\'llll.t iIl-llefined; when examined frolll
behind it hllClllh ttl OOllflie.t of Rcrics of 11lllllerutul;l' conl'se punctures, whioh.
when lool'utillil!ud HiclowaYH, nppcal' 111'1 if tIIC;I' WOI'C tJ'l1.llHfol'med into
gl'lLnuloR, II l'coulilll'it~, which iN also Ilppul'cnt ill Romo Hpeci6fl of COfllel1fH.
Antl!nnllC Rlltll'Mcly puhlJllccnt. the CX}lOHCrt pnrt of the 1st joint not
longer than the :hlCl, :hll twire ILM lung UM hl'()Iltl, 4th Mlig'htly longer
thlln ::ith, Rth 111111111 nud il'1l11HVCrMl'. (,Iub uhlong, nb,'uptly enlarged,
Ullh,"llllUctl'ionl, hpi ug ILttuchod to tho Hth joint uutHide tho llliddle, its
hasal joint nlllltlKt UM bl'olld 1110 till' intl'I'metlilltl'. the tel'millullnrge Ilnd
l'otuunlLte.
It 1U11.~' 11(, diHtiuglliHII('c1 t'I'(1ll1 the oth,',' I'Ccol'l{ed I'Ipeciefi by tho
fi.lttelleti hidl"! IIf till' thlll'lI '\ 111111 l,,'culil\\' IIculptul'c. The IlCtitC on the
tibiae It 1'0 I'uth('l' fhw.
Lellgth, 611101,; lll'(ltll1th, :It mill.
Akn.tu.r·lI\\u. nt'l\l' Wl'llingtull. My I'lpe('imen \\·1111 fuund h~' MI', A,
O'Connor.
large, "ith 0. IkL'l) Hemicil cnIul' C\lJlhlOll hCtW('l'II it I1tHI t.ho dentiform
secou(I onc, which ill Foitulltl'd nelLr tUll 111 i«I<11(', the pr01lli lIeut IlostOl ior
hnglc Cllll hftl'dly ho tel'lIlo11 n lohe; dillc lIlllch cll'vntud, VI lih n bronll
l'hltntHll 1'1'0111 front til rl'lll', n pnil' of Ht I'Ongl,\' l'h'vnh'd pl'nlllincllccB
fOllll tho llttcrd} hOllllc1ul,\' 01' th«' «·1111111101 ill I'l'ont, tllel't' ill It }lnil' of
I-llllllll'r OlH'H Ill,hillll, 111111 Itll0thcl', Il'HH I'll'vnh'cl hilt tUlII'l' I'I0ugllh', nOllr
the lIIi «MIll oj' tllC' hltHll; tho 1I1'nipt 111'(' itl g I'an nIaIl'. l~l~·trlL IIlO1'tl thaII
double the length of thlll'Ult., thuil' Hi«lel! IIl'IlI'ly VI'I t i('1I1, \\ ith hen'ate
11Iil.1'!('inll; on 1'0.('11 Cl,,'tl'lIll titero iR lin ('lollglltc tlll)(,l'cie ot the bnE4o, not
fal' f1'om tlJC IIninrl', 1\ pnit' behind tho l)1t"lIl 0111' hut 110111'01' thtl hide,
nn top of the declivity (potltol'iol'). lIOUl' thc 1'o1ltul'C. :I I'olludod pl'O-
1lIinence, nnd n pail' of IImalIel' onel:l 10woI' «lown; then' It1'e I'I01lle others
(Ill thu siutl, Ilnel n ulillute one hehind tho middle, nOlu' the 1I11tul'e; the
disc is sJmost flat along 1,h(> middle, witll nl',lJ'Jy sorin11' 1I,l'l.ulUJar bcuipturo.
.Autenulle \\lth nuo bt!t.J.C, tllllbO, hu\\o\'c., UII IILl! Unci. llllhlll joint Ilro
coal'ber nnll i>rullIooY; 2IId stout, OVifUl'lll; :~I'c1 olonguto, hilt llot quittl as
long aK the 4tlt uud .'th clltllbinctl. LOgH" ith ('111'I('«i tltllUllUiflll'1ll setae;
tibial:! ftel.l1UUh,
Unclel'sidc opltque, l'edcliRh; tlw Ult!tllloltOl'lllllll, IUlhul VtllltJonllootlgtllcnt,
and middlo of prostel'llllID with «liHtitlct gl'lIllllltlll! nnll V01'y ft!w yellow
setae, tbtl othor hegments llWI'C tinel~' sculptlll,..d, lin 11 kH of the I)I'Oht('1'llUlll
covered witll sl\ppy mntt<'l'.
An oi>bCl11 oly coloured t'lougnte IIpocics, with "touter le~tI thltll 1.',
imlm taf 11(1" I\nrl with ~\lto~ethcl' rlilierent HOulptul'c, the cl~'h'al pro-
tninenceH being distinctly sepnrntt!d from one atllltliel',
Length, 4~ nml.; breadth, If mm,
Mount Greenland, neal' RORS. From ~Ir, R, Halllilton'l! colleotion.
3221. Notoulus demissus Kp. nov, NIi/outulJ Bronn, MI,n. N.Z. Coloopt.,
p, 183 (.tblrdms).
Oblon~, ('011\'1'\, Kllho}lllllllC j UhHOUl'l' fllHCIO - l'lIi'IIIIH, thl:' depressed
scutt'lIar I'pgion IUlIl II ltll'Acl lll~lliltn Hllot IlCl'OkH l'Holt tlI~11'On Ilnrk fUFlCOUR;
lLogti f01'l'llginCIIIR, nntonnau 111111 tlll'Mi KOlllowhnt rulvl'Hccnt, tllCll'll.Ilic lobes
teHhlllCllUh; KplLrill!!,ly l'luthE'cl with Hhol't, mondel'. mOl'e 01' less cuded
flnvosoont Retac,
llead IlLrgo, 110nl'1y .IM bron<l ILH tlUJ thOl'IWil' 11i~C, with indistinct
gl'Uollulllol' Hculptul'c. 1110l'UX l11Jcmt lUI bl'ond nil long, excluding the
lateral lobel!; llllCVtlll, with u largo Illetliu.n illlPl'l'BIoliun which RCOlUS to
o'ttelui to th~ HidcR, cliHtilletly alld i1'l'oglllal'ly g'l'nnulltte; its sides
bl'oadly explnnate and hilnUe< 1 j the 11'IIutaliobe iFl largE', with itl! a.nterior
angle projeoting nbnoHt UII luI' all, yet cliKtllnt from, the centre of the
eye, the 2nd iH l'ylincli'itln.l Imll sitllUotl>cl bl't\'vcell the formel' a.nd the
base, pOllteriol' angll'H itUliHtinct, Elvtrn hiioc tho length of the thora.x,
with ebtuNe IIhoull1('I''', AO tIl nt, lit th~ lllIHO, they 111'0 1l() wi<l~'l' tha.n the
thorax; their Hides nl'e Hturlderl with A'1'1~lmleA, and appellor IIllhFlel'rate,i
420 'i'I'(/I/Mutl()IH.
thtlY are serillte-pnnctntc, l'egl1IUI'I,\ .lllllll' ~he, fool1ttU c, lIot hI) I)l'~'ond; the
scutcllnr region UI clcpl'ehhc(l; tIll'l'C lb Il (hhtmot, though not largo, hUMl1
elevation of tIlt' :ird iutci hticl'H, lind Oil thc bllllllllit or thc apical dC'clivit\
on each cl,\ tl'IIn tILCl'C iH n l"li!' oi' .. nudl llIulmliti('h. .
~\lltonnllo with the cluh Ill·moly PUbl'HC('nt, lwond HlIIT hiat,ticlliutc;
bLUml joitlt but littlo C\[lOHl'11 ubuve; lane! Ihid~, IIh long 111'1 hl'Cllul; :il'el
..ltlllUl"·, not l,lullgute, ~d lOllgcr than it ih Ill'Ollll; ,!til IIIHI !ith Hlight1~
lunger than 1.)1'(111<1; jointH 6-H \lUlU ilifurlll aull 1'1111011; Hth mthcr bl'uudCl'
thun it~ pl'cllccl'lo.SOl', not dull untl 1'I1b('bccnt, 111111 only nhont II third uf
tho width of thc club,
Only II fooinglo Sllt'cios, N. lirt'I,iR (1 'i:i:i) , l'chClIlblei! thih. which. ho\\-
ever, iR evon ~lllll11cl'. with mu!'e-cunVt'X eIytl' II , I'Olllldl,d hhul11cll'I'H, lind
difiol'ent thorilocic impl'eBRionR IIml 10001'1,
IJength, ucnrly 2 mm.; bl'cudtll, 11lull.
Mount Pil'!lngia. Doeembel', 190!!. Unielue,
In its nntural conllition it i~ ('ovcred with g'I'c,vj,.,h bILPl'J' lIIuttCI', so
that its rOlll ~culptul'o ILnu thuJ'II('ic lout's cmlnot hc ReCti. 1'ho removal
or that ilubi,tanco by dOgl'(,l'~ with thl.! point (If a nccIUe nnll hrushing
with ht'll?CIW ih 11 Vl·I·.'· cldi(,llt(, Illlel tClliollH OPl'l'utioll ill till' ('nllC of It
somewhat ru;PCI'n.ttl inFlOct UbOllt tho Hizc of n pill'l! hCI\cl. Atun," l'att',
tho cleaning v.ith()ut ciamago ullI1 the fJUbl!Cllllcnt elt-scription (If this
solitary hpecimon oooupiC'd II wholl.! (In,\',
Dr. Hharp's Bifr)111(1 Ilt'llafa (HI27) HhouM he 1,1nccll in this I-.rcnu!..
It ill not in tho least like tho ~l1I'()pl'lm Blt(J'nIfl, IInll lHn~' Ix- I'on(lily
separated from Clollofu/I by itl! hin!'ticlllltte club.
8225. Bitoma maura sp. nov. Bitoma Herbst, Man. N.Z. Coleopt.,
p.192.
Elongate, Suoucpl'esseu, entiroly dull black, with n fow elongatc, hculc-
like, yellowisll setae.
IIead rather lI111aUer than thorax, with granular bculptl1l'o. E~'L'8 of
moderate hizc. Alltcnllae illbcl·ted below the cdge ot' tho forehead i the
el.pObc<i portion of the bl\iUl.l juint not lUll!-lCI' than thc thick lIlld, which
iH nu.rl'o'Wecl towlll'd~ it.. bll.llc; :3l'c[ 111101\(101', longc!" tlum tl<lj(lini.ng ones;
4th and I)th ahout oqual, 10llgo!' than hl'cmcl; Htll aIle! !lth lIIonilif(lJ'lI1;
tho 10th ohollniclll, twico liS bl'Clltcl 1111 thc !It.h, uilout ttll long all hl'Clad i
11th oblong, 111'1 hl'lIIUIUb tht, 10th; tht'Ht,l t'I\'(I joilltH III't' (h'nHCly unll finel~·
ptloOKcent, uncI flll'llI the club. '1'hm'u,x lLot lflhnt~, "idcMt nelli' till:' huut,
Al'll.(lllUlly 11111'1'0\\'011 bliOkwlll'dll, \vith ohtuHO nll~loH; tll(,,' Hlll'fucc It little
llnCVI,m, with II llll'go dCPI'CBflioll on tIll' 1lIit.1I11c oi tIl(' <lih('; itH IIcnlpt1u'o
ill ill-Ilefinod, hut Upp~IIJ'" 10 111' gl'lUlulul'; it iH Mlightl," hl'(HUll'I' th,1Il
long. Elytl'u. ulUlost thrice the leugth of thol'll~, ovillolltly lll'ondcl', thci!'
sidell parallel; thc~' orc hI'OILlU," iJupl'(lNHCd bt'fol'tl the Illiilcll~ anll hchind
the Bcutl,llulll, their Rculptul't' BtlClllS to consiKt of olosely plnced scl'it!s of
granuleil.
So ill.l' a~ stlpel'ficial appellol'anct' is ('IIU('e1'l1ccl, thib I:!poricl! MtalldR alono,
'rht' cl11b is unusually elongate.
Our B, inRidariR ann. B, 1'I'('i'l1o (a4a and 344), so fat' as can be judged
without disl:Ieotion, will pt'obably remain in this genus. All the other
species are oel'tllinl~' different from the trpe of the genus, the EUl'opean
B. C'l'enata.
TJengtlt, 2i nun, j breadth, neaTly 1 mm.
Waill1al'ino; elevation, 2,600 ft, j January, 1910, One only COllIn
be fotmd.
nItOl.:~ -x flV (lfll( r(l alld 8ptrif'~ of ColcOpttll n. 421
butUl'e atlll arljoinillg illh'l'htioe, IlH Ilell Ith tht' {'Ill iniilll'lIl 4.th lind 6th,
(11) not <Illite l'c.ll'h the hahnl Illlll'gin, .Alltcllllltl! Ilith til(' 11th joint
indif'lti net.
lTnliol'hillt! l·ufl·!otCl·U!, hhilling, \\ ith I'lIthl'I' ('Olll'~t' }IUlwtUI'('h, I'uoh with
u lIIi1lut(' ~l'tll; lith Vl'lItl nl "('~Im'"t Idight1:· ('OIlI'II"l', ulIIl IIIIllOlo.t IIh
coal hel,' }IUl1"blh' 11101 tltl' IIthl'l'H,
.\ ('ul'('l'ul hOlutinl Ilr tho hllhO or till' (·1 I'll II \\ ill hl! 1111 lIill iu l1i,.,-
ot"imin.Ltiou. 'I'ht' lI~tt('I'iol' IIngleH 01' lltl' lil(lI"Il\ 1I1l' IIJIII'l' aeuto thall
tho~ (Ii lilt!), thu Illto1'111 1I11tl'giliH 111'(' thh'kl'1' th l'tlll~hl)llt, nIHI the dilio-
ouidal improhsiuuh al ~ !lot at ItIl sllul'ply hOl'dl'I'l'I1.
Length, 3! mDl,; hl'eallth, It Jlffil.
Gl'eyulOuth, Unique, JhlUlld by Mr. J. H. Ll'wil!1,
Oba. - From tho Iio!tlll!:' bom'Cl' IIjlel·illll·1l1-. \1 ('I'l' ll't'l,ivl!ll IIf H!Jlu p'lo
P. 10ll!lll111h, If I nm right UN l'ogUl'dR itlentifiolltiulI, the follClldng notes
will bo tIschll to othel' ilturlcnts: PI'Ot.tl·I'IlUIll 0Pltllu(' aud olllhl·ly puno-
tate; metm,tt!l'num gl(lRil~" mOl'l' (Joll.l'Nely hut ratlu:ll' oiHtttJltl~· punctured,
medially Rulcate behin!l; t!:'I'lllinnl vl'ntl'nl I>llgllll'Ut dl'pl'CI'IHecl 01' slightly
oonoave. ltil fl nntal 1tII1I',,dll inonl veil, thnt. Il'ILving' 11 t!'IIll"V(,II>e depres-
sion at the apl'X of the penultimate, Size, Dendy 4. rom, by 1~ mm..
G1'OUp ROTmUDJ~RIDAE,
Group Cnll"j'OPIlAOIDAK,
3229. Cryptophagus amoenus sp. nov. O'!JPtophagus Ht'rbf:ot, Man. N.Z.
Coleopt., p, 224.
Minute, clongatu - oulong, IIlCllleJ'utely tl'.L11hVUJ'I'.oly convex, nitid,
castJ.neO-l'UfOlU" logll uud tUl'wiuul joint of alltennllc flllveMJeut, remainder
of theso lotter I UfOUb j bl'll.1'ingl,\' clntiu:u "ith kuben,'Ct pale flavescent
setae,
Head eviLlcntl~' narl'oWor than thOI'ILX. not trigonnl, the flJrtlhead sub-
truucate. it it! IIl1looth on the middle, mouerately COal'Helr, proportion-
ally, punctllrtld nt the sidell, and ,vith a I\eries of coare.e pUJlctUles acr08S
the occiput. Eye!> COllvex, witll uit,tinct facets. Autennne inserted in
front of and just below the IlIuI'gin of the forehead, 11-llrticulute; basal
joint stout j !irll I'ather longer tlilm 211u; Hh and 13th as long us broad
and about equal, )'uthul' shorter than their pl'ed(lCeShOI' j joints 6-8 rathe!
&malleI' anu moniliformj nth I' ather nlll'l'owor than 10th, but larger than
8th; 10th tt'lmSVCI'lie j 11th conical. Thorll,( Hub'llladl'ate, r.lightly and
very gJ'ltduully narrowed townrds the obtuse front angles, its sides mar-
ginate but Ilowhore denticulate; base slightly bisinuate, with nearly
rectangular anglcs, it" length ILnIl bl'eadth about equal; the surface,
relatively, moderl1tely ooarll6lr punctute, Hcutellum transverse, smooth.
Elytra as broad aN thorax: at the baFoe. nlmost twice itl! length, slightly
narrowed pOHtel'iClI'I~' j with fine sutul"lll striae. their pUllotation a little
finer and mol'c llil\tnllt thl1l1 tha~ of til(! tlllll'tLl, nnn. hccOlning finer and
less distinct h~hind.
Tibiae somewhat dilatod tOWIIl'cIb till' extremity, unarmed. Tarsi
quadrial,ticulate, tho baeal three juintb of ahout O<lulll length, each of
these fumiHhel1 ",iill a Hlendel' clongat(· seta, the terminal rather longel'
than the othel's taken tug-ether, with dilltinct claws,
.A. sin~lo Hpecilllcn ou1r of this mhmw inl!lect bSl.ll been secured, It is
not a true Oryptopnaglt8, neithel' rloes it agl'ee with any of our OtJ,c'Ujhlae,
to which group nevul'thclCIIR it will no nouht bf' transfel'l'ed as the type
of u distinct !!,'cnm. if othel' speeimellH CIU1 b(' ohtained.
Length. 1~ mm.; breadth, ! mm,
Makatotc. FOllllfl aIllOl1f.,"Ht led-llJ()llltl enU('Ctctl fOl' IllC ill Febl'1II1.1'}"
1910. by Mr. W, .T. Ollinnt'Rs. •
nl'(lll p f..IATHlUllllll.\l<l.
IInother neill' the b:1Hl', dOHe to the hUllal Illal'gill tJIl'I'l' iH II tJ'ullMVerl!('
lineal' (1epl't)NHiou, ib. punl'111tioll ill il'l'cgllltLl' nUll mUll'I' eOllI'Ml'. HUlltel-
lum 1I1IInH, Elytl'n ohlnnJ.(. III111'h bl'cu\(ll'l' thnll t1ml'a..:. Ihl' Rhouldl'I'R
rounded hut not at ull elt·vutl'cl, Hlltlll'II1 Htl'ial' cri~titJ('t. tlll·it, I'ellltivdy
Cuo.rRC punctlll'eR nre Ill'ltl'I,\' 'Illite lIC.'I'inli', 1m! h(..'1'UIIIC thll'l' hl'hillCl.
LI'g'H pnheHcl.mt; IIntel'iol' tibiaL' Hlig'htlr bl'nt U('HI' Ihp e:.tn'luity;
tal'~i Hlt'!Hk'I', tCI'millu1 joint «uite IIH lCIllA' II~ tltl' hUHal h'(I l1uitt,,1
Om nel1l'eRt HpeciuR, (J. frrril'of(t (:27!'11), hUK II l'uthl'I' Khul'I('I', Hub-
oVllh.' hind-bod.,-, with tlilltinotl," l'UiHC,I Hhoulllel'M,
L(..n~rth, Ii mm,; bl'eudth, 1i mill.
Erua, JUlllltLI'Y, l!llO, Ouo. (ollm] UIIIOllg'st t1elltilClI\'CIl,
Uroup BYllUIIlDAU,
pUllctured 01" stl"i ne 011 the cl,,·tra; it mUlit therefore be distinct froUl
thili hl'llCiul! nnd P. lell,illi.
Length, ~-!) mnl.; Im.mdth, 5mm.
Bohl P('lIk, \r uklltilHl. .Anothcl· of MI'. H. Hamilton's cliHcovel·ies.
(h'ollp (',WHW.\Fl.
3232. Saphobius lepidus "p, IlOV. S"'pkobiu8 Kharp, Man. N.Z. Coleopt.,
p.255.
:411hl} uadrlttl.!, 1It(l,lcratl!l~' nitill, hcall'in~ llhlll't, rather linc, fiavellCent
IIllt.tC, "'1IiI)1I ou tho clytl'lll iuturtltil.ll.!H a 1'(' cliHpol:I(lll ill Illtlloht l'ogul:u'
duplicate scrie!!; n igreRclmt 01" l'uro-piccOUl~, lcgll tla l'k 1'U£lIllM, !l.ntenno.ll
lIud palpi I'll fo-tCRt:lCC'OUH.
HI.!l\ll llltl'rClwctl tOWUI'11H the Cmnt, biflcutll.te thlll'C, rather coarsely
and cloHel~' puuctllred. Thuru" twice as 1)1'01111 IlS long ill the lllicicUc,
wiclcl~' l'mul'ginatc nnd with nClltl! anglell in f"ont, the hidl'l> llcul'l~'
htl'aight fol' two-thil'dK of their ll!ngth, th()U obliqucl.\" 11 II I' l'owNl IlU-
tcriorh', the hasc slightly rounded, its anglCll, nevel'thdcilM, 11I!arl~' rect-
. angul~r; with Hllllllol\' ovirorlll impI'cRRiuJlH }'ather than pUlll.lhll·CII, the~(l
aro not vel'~ dOhl', IIlIll eal'h hnM II Hhlll·t Meta 1)1·llCl'l.!l1ing f"OIlI it, there
is olll~' II Klight IOll~itl1elilllll tll'lu'l'NHion helline!. Elrtl'a ul' exactly the
KaJUC wieltl! lUI till! tllOl'IlX ut till' 11111'10, l)l'oncll~' "oltnelell )l~KtcriOl'lr, and
Ilovcriug- the p~'g-i eli II III ; on l'lIch thol'l! ure 6 lillOR, which can hatrdly be
tl!rmcu Ktriac.
Tibial' finely lIetoRC', the uutel'iol' CIlI'Vate, gradunlly llillltC'd, ohli!Juel~'
tl'ltncute at the C'xtt'clllity uncI with IIcutely prominent e.rlcrnal allglcN,
there al't' 2 mOl'C' Ill!lltidl'toI 1111 the Huh'I' (·,lgl!. lntt:'I'lIIl'clinte anti hilll!
tlll'si well n('vl·lopt'cl, tlll' anti-rio I' Hlelltlcl' and nhhl'('viutcel, so that eluring
repose they do not C"xtNl<l outwards as far as the external angIe of the tibae.
~\ntl.mlluo iUllertoel bolow the llic1C'R of thC' hean; the>ir ('longlLte bD.Ral
joint, which iN aN lung tit! the following' five cnlllbinC'd. ill therefore pnrtly
I.lCll.cenleel from nhelvc; 211e1 cClnioltl, Htnlltel' than the ht; 31'd and 4th
1'11111111; :Jth nnc1 6th Hmne,vhut tl'unHvl'!'lml j oluh lllocI('rate, pubcsc('nt,
tria ,·ti('ulMo.
Un,lorNille Rhining, piOCOtlH, with minutc Rl'tne, the Rternmn coarse-Iv
punt'tnt(.', nbclolll('l1 finely. IIlctltHtcl'nUIn neal'I,\' Hnwlltll on the miclcUe. '
111 ntlwl' Hlll'llieH till' l'.'·l·H, thuugh nllt Ht lin )ll'ollli m:mt, 111'\' 1111 ite> c1 iM'
CCl'lIihlC" IIhovt·; tlll'v "xtPIlI) elown\\'nl'elll, nwl 111'(' Rituntccl at thC' hack
pl\l't of till' hCl\el .inKt inMiclt· till' thCII'ncio nnglclI, hut in thill IIpe-oiel! they
nrc nhuoHI inviHihlC' ahov", thuugh w(-11 ,11''f'(llopcel 1lI11lemcltth; these
Ol'gnnR, thl!l·c·Cm·I' , 1Ul<1 till' IWlttl." nI'I'amgcIl lIt-fllC on the> elytl'a, will enablo
thiFi Rpeei(,14 to he iell'ntifiNl.
L'mg-tll, 111'1111 (.'xHt'rtt'II, 4- 111111.; hl'euelth, 21 mm.
E"un, neill' W':\imlll·inll. Founci amongllt 11eoa.~'ing leaves on tho
grounel ('levlttiIlJl, 2.IiOO I'L), ,T 1111 11111'.'" JIlIO; nnd Rent ehning' ~llll'ch
b~' Cnptain II. H. Whitt·horn, 01' the Geological Altl've.\· Department,
mnongst veglltllhlc mnttel' collectcd at the- hene! of the Retllol'tlke River,
about five mileR fl'om EmIL.
Ch'oup MFJT,OT,ONTlllDA.E.
3233. Odontria nitidula RP, nov. Oilo'1llria White, Man. N.Z. Coleopt.,
p.265.
('on vex, Ruhnvn.te, shining j teetll.<.'OOUB j the ve,·tcx, milldlc of thol'ax,
and tibin.l teeth mOl'c 01' 1(>R1I infllRoate j forchead l'ufo-onRtaneous, spa.r-
426 l'l'lIlIlIarfloll".
Bold Pc,Lk. Wak ..ttipll. A pair, mounted on Cltl ullum rl. fot'" al ded
for im,pootioll by Mr. A. O'Connor, of Wellington. TIle 1111.1.10 '\ab CIIP-
tured b;r Mr. Hem'it" the femule h~' Mr. n. H,t1lliltOll.
8236, Costleya simmondsi "'p. nov. C'08t'leua Brou1l. Man. N.Z, Coleopt.,
p. 1115.
Bl'ulUlly oVItI. IIlc)lll'l'ntd,r l'el1lVl'\, 11 it icl; 11l!III'''' glnlJl'()uh, liaviJlg olJl~'
tI flo v.' ..ltlllclel' ,,('tlLc.l (Ill tlll' IH~I\(l atlll 1\141111-\ the Mitll'K, the p~'giclium and
tibiae.: with l'UlIl'Ht'!' om'l!; thl.' t'l~'Pl'l1h, l'l.,·tl'll. luul sidt'H (If t1torl~:3' rusco-
teHta.ceOllN tiul-\'l'd \\ ith gl·oc.'ll; till' huck or tIll' hCllcI nnll milleUo of thorax
light CUHCOIUI; tibino pil.·I\I)II". 1J10I'~ 01' It'Kfi vi!'ich·HOl·ut. the untl!l' ollllc Clf
the ttntl!rior l'(-d.dish; nllt~tlIlIH' pale cIUltanl'O\lS, club opo.qul' and ne'o.rly black.
Head il'l'Og'ullu'l,v, cOIII'Htll~'. but 1)ot clolldy llt1nctul'cel; ci,vlle\\s with
sotntlwhat l'etlextlll margins, 1I1lghtl,v JUC!elil\lly illOlll'Ved ill fl'uut. Thorax
nearly twice aK bl'lI'lcl aN it il:l long, HH KideH finely l'iUllllecl, mOl'O
na.rl'owed in hunt tlmu hdlillll, itH hlu«' Htrongly biKinllutl:!, tlle apex
deeply emal'gimtte. front ntl~kh projectiug' be~'oncl' tll(· mindle ()f the
eyes; its punctll.tioll 0(1111'11(', nut Vl!I'~' dOH!!, hut il'reg'1l11l.I'; then' ill II.
median illlpl"ehlilion JlC:U' the frout. ~}l~·trl1 of the lIame width as thot'ux
at the haMC.:'1 l'ltthcl' wid,,1' hohiIlIl tlw mitlcUC!. npiCC'K nbtl1l1clr roullderl
towards the Imhu'C!; with ruthcl' l)J'oad, olC1sl!l~' punctul'ocl Mtl'iae; intel'-
stices Rlllouth. th~ :Jl'cl. :;th, untl 7th bl,utul!:l' lind 11101'(' clevutell thun till!
othel'lI. thuil' HCl1lptUl'l'. hnwf.lvcl', heQelUlI'K illtliHtiuct twnl' the npiccII; the
latl'l'alll1ul'gillH nrc !!ClIllt'whltt l.''']l11111:1t(' lllUIlI'It to the l.'xh'clIlity,
428 7' 1'I1lIH(I('( /01//1,
3237. Talerax dorsalis till. nov. T(ll('fI1,'J:, Shl1tp, MILll, N.Z. Colf'opt.,
p.279.
Huhpal'llllcl, IIwclel'utcl,\' 11111'1'0\1, llitic1; hlllck; II'A'1I i'lIHcollH; thl' kUCl'lol,
t1l1'l:11, nncl npiclII lluu'gin of thol'lIX cIlHhml'ollH; l'ubcHcenco cincn'CluH,
Hltlnclcl', nnll clun~ltt(.), Hlight1~· t1avcHCl'llt !lllli thick(.)I' I~t the hlloKO of thu
thCII'ax.
HCIHI IlhllllAt !ll'l hl'onc] 1111 the (I'tlnt of thc thorn '<, rcehI~' Illl'ciinIly im-
PI'l'HHCc1, c]iHtillCtly but not vCI'Y dOHcly PUlIctut(', g~'('H llwI-."t, ntHI p,'o-
lIlincut. 'l'hol'lLX II third hl'oadcr thun 10llg, IlpiClll 1llltl'gin HUIlU.lwhnt
re:liuxecl nml hl'uatll~' l'tmlllleel, pUHtcl'iCll' IlTl~k'H l'ollUHt, 1'111.111'1 IOIl~, I~
little curve,l, not at nIl 11ivl'rgc>nt; oillc cuuvcx:, lliHtinotly ,Y'ut rathcr
finely but nOWhel'l' cl()scl~' PUJlctul'ed. Kuutcllllm lIlinutely soulptured,
Elytro. as )ll'on.d as thorax at tIle bltRe, tlll'icc it~ lcngth, tnpcring VCIT
gently towards tIle extl'tJmit~·; the slwllltlerl! unci onch sille tlf thc HlltUl'C,
at the b8J!6, ohtul!ely CltlVlltcd, 1!1ltul'nl Rtriac illclcnnite hut h!l'1uillating
neal' the apiccR in deep fnveif01'lU IICP1'CKHioUR, theil' Plln<:tutioll iH coarse'!'
than tha.t of the thorax anll IlI)penl'K Klightly l'ngoMo in HOllie IlKl'cctl!, there
nl'U 1l1HO RI)U1e obl!oletc I!tl'ioc noal' the mieMle.
l.egs VCl'~' finely pullCKutlut; tal'lIi Hlcmlor, Hitlll'Ie, hUHal joint IWIlI'1,\'
ilK Inn~ lUI nIl the following onOli. their ]lcnllitimnto joint with Klonelc!'
lubes, Alltellnnl' distinctly hut nut wi(ltoly Hl!]lllflltl'tl lit tliu 1IIIH(,; lRt
jnhlt IItont IlntI sOlllowhnt ctll'v()ll; 2nd VI'I','· Hhol,t 111111 i('('hl~' rlll'('Hoont i
:31'11 elongate, nil lUIlA' ilK tho lilt j jointH 4-10 HllhYCI'l'lIte, 1111 1111lgl'I' tltllll
ul'oltd, the 4t11, 1,IIWCVCI', ill hllol't(!1' tlllm 1I11juillillg onUH; 11th eI()lIgutc;
thl'Y bellI' diRtinct ini'IlHcut(' PUlJl'HCl'lICU,
1'. micnm (2:)66) C'Ollles nCll"CKt, hnt iR l'nt}i(ll' lurgel', itH thul'l\x is
II1m'c gloRRY IInrl il'Togulal'ly }>unctul'ucI, HO thut the lIIilltUl' 1l1Hl uther
IIpotf:1 0.1'0 nlnwllt Hlllouth, the olvtrn.l Hcnlptlll'C ill Illlll'(' Ildinitl', th€' hOFutI
elevations leSll bO, '
J.ength, 31 mm. ; bl'cn!lth, Ih m1l1,
l[ount Pi !'ongia.. One eaptl1l'erl by myself in Dccembcl', 19()9.
Group ELAT1!lRIDA1!l.
3238. Prote1ater diversus sp. nov. Protil(1/6f' Sharp, :Man. N.Z. Coleop1i.,
p.304.
Narrow, elongate, anil shining, pitchy black, clytral base sOlllewhat
l'ufesoent, the knees and cla.ws pale cast:nneotts j pubescence IIcnntv, slender
but diAtinct, greyiah-yellow. .
Bum N.-X I'll' VelieI'II lIml Sperit'R of ('olenph·l".t. 42~
3239. Chrosis dubitans flp. nov, Ollrl)sis f:!harp. Man. N,Z, Coleopt.,
p.296,
Uobust, not lJUI·ullL'l; plli>(,KCellC(l lliHtillct, pale urmll!l~'; nithl, rufu-
piceolls, antennae lligl'oRcent,
Head llwllerILtcl~' Ot)lu'Hcly Ilnd irrcg'ulal'ly PllllCt11l'0d, with It nearly
Hmonth apnec llCltl' each c~·c. .Antenl1uc finely pul'lCRCcnt, hu.l'dl~· attaining
tho 1)880 of thorax. :11'(1 joillt l'ntlwr longllr than 2nd. Thorax of ahClllt
cqunllon~rth II.ncl hrCllllth. g'I'llClI1UU~· ualTowccl antel'iorly, with carinate,
robUilt. und Hlightl,r cliv(ll·gent hURal !lng-IctIo; the liid08, befm'!.' the lIlirlclle,
arc hmadl,'· )·ct Hlig-htl," illllll'ellHccl, thl'ro is all elongate ll1eclio,l1 impl'cR-
Bion l1(MI' tho hUKe. I\nd lI!!III' tilt.' fl'out lUI illt1iNtinctl~' I.!Iovatetl lino, the
l1illo ill finely IIIHI l'utht'I' llilltinctl,\' }lunchll'l'cl. hut the 1I('nIpt lin' bt'ClllllOS
ClOSl']' nntI mlll'(' tlilltil\('t nC);~l' til(' Hid('11 UTIli 1l1'l'X. El~rtl'lI l'Iltlll'l' clcl!pl\"
Atl'iatC', tho (lnll'I' Htl'illl' l·vill('ntl~· pmwtntl.!; l11h'l'Htic(''I! llilltillotly hnt not
'\"el'~' cloHcly Pllllctul'l'cl. till' 21111 h('I'IIIII(,1I ohHo]cte Ill'hinll till.! millc1k·; IIpicCR
l'cmndotl Hillg1.,·.
Ncnl'l~' 111lil'(1 til (t. IJtrl'lmlfl (!i2!1). nnd pCI'hu}lH uIIly IInc or l!l'vcl'ul
vllrieticR. It iH cliKtingl1iHII!.'(l II,\' tlu' ruthc!' IliHilmt foIcnlptlll'E! nl the
thorax: IIncl the nhhl'('villt ion uf the 211~1 elytrltl intL·I'lIti(·(.'H,
Lcn~rth, 16 !lI1II.; h"(·llIlth. !) 1II1II,
MlIIlllt A1l'hu. lICU]' Wc1lin,.,rtoll. ,·'C)llnd Illlclt'I' HiollUK. at nil nliituoe
of 4:.70() ft,. II," Ah. H. W, RiullllondH.
tIu.' front, tmblMl'nlld hl.'hilHl; ittl hlltml :m~leR nncl,\ cnl'inntl.' IILovtl and
dil'cctell hack" 111'<111, tho IInte1'iol' PJ'Cllllilll'lIt hut ollh1KI.'; the KUl'face
ltwdel'!ltel~' cloKel,\' nnd I'athc',' thll'l~' ]Il1l1ctntl', HtiU 111111'1.' tillCly n01l1' the
hllRe; thl'l'C it< nn median illl)ll'l'KKion, l~lytl'n tlll'it'C 1111.' lC'll~tll of the
thorax, a littll:' \\'icl('t, 1WIlI' thc hillcl thil.\'hH HUll1 a1 tlIP "holllcle1'1l, tllcil'
nlliccK t'lIulI'gillute 0.1111 IlIiun1cl,\' ullf.\l1Iah· j t1U'~ Ul'e finely puu('tate-
Ktl'iatc, with Illiuutely fool'ulpltu'l'l1 intl'I'lJtiecH,
IT 1l11Cl'Ki d(' flllvCtlllcmt, VC'I'Y tin~ly clothc'cI IIlld p"llet III'L'cl, tho Htcl'nltl
HtJ'llOtlll'C :-limillll' to that of U, nnfipod/(I/I,
AlltC'llnlLC nlifol'm, IIttuining the bllKC or tho tlWl'U \, tlll'it' IIml jllillt
,plite half the ICll{,rth or :II'cl.
It may be ,'cnclily iclcntilk,J J)\' hcin~ IllUl'U bl'il.\htl\' clIluUl'ed than allV
NC'w Zeaiuntl membcr 01 thc g'C'llllH, U, (f'lti/IOr/llm '(:;:l~) iH jtH 1\eal't'~t
uIly, but the pll1I!Jtate c·l~,tl'al Ktt-i II l' llntl the ttifft'I'c'nH,\' rlll'lIlt'cl I1l1tl ICHH
denKl.'I~· Rf'ulptul'cd tho1';tx url" c!iHtillctive,
IJen~th, 14 rom,; bl'eudth, 4 111m,
Wnil'il'i. NC'lIwnl'cl Knikolll'UM, .\ Hillglc.' illclivicll1tli j'ollllfl hy MI',
W, L, Wallace, of Timnl'lI,
3244. Atopida basalis Hp. IlOV. Alopida WhitCl MILII. N.Z. Coleopt"
pp. :n 1 "ud 1141.
J<Jlollgllt,·, Hlightly niticl; hl!lIcl aull thlll'ax [uHco-l'11follK, clyl.I'!I. CILM-
tUlIOClll!! "itIL KIlf[uKl·(1 4111l'k 1lllll'1tK, Ieh'" 1I11cl pll.1pi tUHtU,CCllUMi 1l1ltennao
iniuRcntl' from tlltl 4t h jllillt 011" 111"11101, ~lIel nno 3rd ydlllWiRll, the blUml
joint I'UiOM"OHt i clothoei with cl('cllIlIlxmt ~'~'llowiHh hl~irR. tllol:!o on the
elytl'l~ oom'Mer ullcl gruyiHh.
HelHI llU'I,'tl, illclneling the Hlightly convex (.!~·eM, liS hl'oacl :111 tho front
of thorax, ol(JHl'l~' luul mthol' till(!l~' IHlIll'tuto-gl'o.nulollC. Thorax nearly
twioe lUI broad 1lI1 loug, lml:lo nnd npCl: RubtI' IInoate, with acutely rect-
angulnr ImgloH, tIlO alltel'iOl' HIIlllcwhat eleprclIIiCd, its "ddes sinuouflly·
narrowed behind, in Ol1e oXIJ,lUplc strongly l'ounded nt the mieldlo; the
sculpture rl(!nHe, 10HH oIoHC tit the lllLilal 11 utl'gin , not COIL1'YO, punctattl-
gl'linulolle. Rcnwllll111 tl'i'llngllIElI'. l'longate, Elytl"8. eviciently broadel'
than thorax tit tho ll1lHt· Illlel nhllnt fnlll' times itM length, Hllbparallel,
lllightly tl'ansvcl'Aely impl'oRfled IJcf(ll'e the mic1dlc, it'l'egularly but not
cOBrAel, punoturod, with 1\ t011(101l0y to. hooome Hubgl'lInulol'1() nen]' the
shoulllc I'S.
4:32 '1' ,.'1/1 VI( t /lJI"
Antt'nnlle I'l.!llcliing h.lr!, \\ nl'ch, to hl'rollcl till' h,IM' of tho cl,dm, thdl
hot joint bt(lllt, 2nd "hlll't lind IlIcmilifol'lIl, +tit thil·!.I'I' than :ll'd Illlt{ quite
lUI long,
In ~4., ,illllata (2:i!H) Il hmUlI Hillll()~it,\ UPIlJ' (,lIch hide of til(' thoJ'u\
CaUhll'l the pOhtoriol' IIl1gk'h tn IlJll'l';t1' III. if tht'~ \Wl't' Ililt'('l(·cl hllclnHIl'ch,;
thh-, i~ not tIl\! Cllhl' ill tht' }I)'t'hl'nt h)lI'C'il'h, \\ h i('h. IIIlIll'OVI'I', hml 1I101't'
lillel,\ hl'lIlptllletl d,ltl'll, (lilll'I'('nny colnlll'l'clllllh'III1I1(', 111111 bu'gt'J' Pyt'h,
fJc.'llgth, 1:q IIUIl.; l)1'olll1tll, :dlllllll.
Knitoke, nt'lI), Wellington. I lLlll iuticiltt'd to Ali', .\, 0'('0111101' fnl' .1
Pili! of 1oo1)(.'cillI(.'1I8.
3246, Cyphon pachymerus sp. no", 0Il'PMn Payk, Man. N.Z. Culeopt"
p,318.
, R\lb~paque, hrCllluly Ilvnl, Rnghtl~· oonvex; elyt1'8, densely olothed with
llloonsI>lcuoUR, MItmnl!l', oiuel'c()u8 pubescence; hod"" KlllO];:Y blnck, legh and
basltl two jointll of antennae rUlloouR. "
Head short and hroad, vel'Y lIlinutely grunulato. Antennae with the
::lrd joint ver~- small, yet l'ather longer than broad, 4th eviitenth- larger
than the foUov.·ing ones, Thol'ax strongly tl'nnsvene, its sides but little
rounded, their margins much filler than tlle basal, with subreoto.ngullll'
angles, the antel'ior "omcwhat dcfiexen, itll sculptul'e like that of the heau,
Soutellum large. ElI;vtrn. ample. witbout deprellSinnll, closely and :S.nel~
punctured.
BROtTN,-Neu' Gellera and 8peritB of Coleoptera, 438
C:I'1I1Jl ('lllltlll\J:,
3248, Phymatophaea griseipennis lip. nov. Phrrmatopli.aea Pa..coe. Man.
N. Z. ('ol('upt" p, aa4..
Elongato. hlli"clt'IIl'OL-lKUd, \\ itll llUmel'Otls CI'l'oi. IIIC'nder greyish setae,
subopa.que; nigrcH(,t'nt, hut the C'lytt·n. with the oxception of their ba.sal
portion, ar~ yellowilfh-grey j antenllae and tarsi infulloate.
Hea.d very 0101le-1y ann eOlLl'K('ly pl1nchu'ed. rrhorllox of about equa.l
length and breudth, nbtuHcly dilated la.~rlllly behind the mIddle. its puno-
tation hardly as clOl'it' and 000.1'110 111'1 that of the hend. finer in front. with
a. pair of smooth, slightly rniRC'd SpOhl l)C'fOl'(, th(' mir101e, AOlltl.!l1UD\ "lib-
triangular. Elytra. thrioe as long till thorax. not twice its width at the
be.ae, rather wider behind, without inequalities of surface j evenly punc-
tured, mOl'e coa.l'sely but not quite so closely as the thorax.
Legs pilose, antE-J'io!' I ihinf' JIItUl('1 ntC'l\' cu l'V(\tl. Tarsi with well-
rlE-velopetl mcmbl'a.nmlH IIp}l(,llclng'('H, l'la,,'; thiek~lllt'rl hut not dentate.
Eyes prOminCIJt, finC'ly fuC'cten. l'llllll'f_(illlltC' ill f!'Ollt,
T.ength, 5 mm, j hrendth, If mm.
Silversi,!'eo.m, I1C'AI' WC"llin,:tttlll. A l'jillglt- f.oIpetlimen, from Mr, A.
O'Connor.
US-T1'8tDI.
T rtWHarf 10118,
8249, Pannius violaceus sp. nov, Pa7'fniw 8harp. Man, N.Z, rolc'opt,
p, 331.
Elongate, &uuclepl'e8Ked, blii11 illg ; viulllOtlOUS, the tibiau and husal two
joint!> of antonnltO flHlco-tcstuceOllH, I'olllaiuing jointh :lnll th~ tllrsi uurkel',
tilc front of the fell'l:honu lIud tbe clYJlC'IlH IlI\l~. ,Yl'llo\\; the hody 11110 legs
with nlllUel'(1Il1! olllhtallllillg' ('UlIhpi('llllllh pullie\ luLil'h,
1l('lIel. illclllclin~ thl' l:u'g'1' PI'Ulllillellt C,I'CR, Ill! brond Ill> the milltlle of
thOl'IU, longitudinally bi-implC!!Kl'd iII fl'ont; it iH vCl'y irl'('gulll.l'ly, fillely,
Ilncl incliHtinctly puncturcd, rrhol'Ux oi ahnut C'I1Utl leJlJ,rth uno br(;'uutlJ,
a good deHl rlilutcd lith.·rall,\ 11('1\1' the mil1eUc'; it!! surface up}lllrently
illlpunctate hut finely transv('I'HcI,\' I'ugose. with Ull IIl1gular basnl imp!'es-
Ilion and a pail' ot' KIIIldl Hlwlllm 1'0'\"('11(;' lIt',lI ('nell hi ell', I~l~·t)'a II littll'
uneven, with slightly <,l~vntt'll il'l'I'g'ulllr I'U~I\(' ,
Antennae mthel' shOlt, thei!' hllslIl joillt Rtuut and evidently longer
than :alld; joiutR:J-fI IliffC'I' hut little, pach lunger than broa.d; olub tri-
:~I'ticull\te. its iniel'mecliate joint cOl'uate and hal'dly as long as the others,
Morc I'ubullt thalli the rcpl'l'K<'lItlltivo Rp<'cicR, 602, (n this s}lt'cies, IU
well 11M 602 60:3, IIn(1 203i, thel'e iH II IMI'C or lesR d('finitc erual'ginu,tion
(If the eyeH; tho author'R gl'Ilt'I'i<, tI i agnClHi~ thl'I'Pt'OI'P I'l'l(uiI'('I1 00l'1'1'01 illll,
IJength, 6t mm,; breadth, 2 mm,
Mount Quoin, De~c,'ibpd f,'om :I 1I101lutpn Hpccimen fOl'wlIl'ded by Alr,
Hubert W, Simmonds,
Ol'OUll .\NoHHn.u
:J21>O, Anobium inaequale lip, nov, A,nobium Fu hl'il'iu'l, M,m, ~ ,Z
Coleopt .• p, 339,
Cylindric, uneven, variegate; fuscoul'l. till' hltBt, and the clevut il)lIl!
of the elytra somewhat l'ufescGnt, legs and antennae rufo-pioeous, tarsi
obscure fUBco - testaoeouA; vCfltitllre un<'qllally distributed, varic·gated,
flavesoent and greyish,
Heat! vOl,tical in front. slightly nlll'I'OWel than thorn x, EyeR lttl'ge
Ilno subl'otllndnte, Thorn rnthpl' bron(1E.'r than it ill long, R01UC'what 011\1-
Ktrict<,d neal' the middle, apex slightly rounrluel, PORtC'rior anglell obtuNd,\
l't'ct::mgular; lliAtinctly gih1lO11R 011 the lIlirlrlle, itK Hcnlptnl'e cioRe IInel
~rltnulal" EI~-tl'n ratIler })I'Ollt1<,1' tlUlll thol'a 'C, theil' n.piccs inclivirluull,l
l'ounded and not entirely llovcl'ing thC' p,\·giclillln; with minutc, clollse,
rugose, and gramdll.l' Rculptm'(', lind HCl'i('H uf punctlll'es whil'lJ ,II't! l'C'gultu'
along the sides but mOt'C 01' less iutel'l'upten on tho do!'sum; OIl each
elytron, near the 8utUI'O, tllere iM n Kligltt hlUml (,It·vatinu. 1\ nUl'l'uwer but
more distinot elevatioll in liuc with till' hind thigll, lind n nol1t1llity 011
top of the apical declivity; ill Uc1YlIll<'e or tho IIlHt, hnt llel\1'~l' th(, Ride,
thN't' iH lin ohliqlle Ollt'; h('sidcs tlll'pe. ~Ilme MlIIallt'l, uHperitieR nre villible,
Antennae elongate, hasal joint stout, 2nd short. joints !l-8 inwardly
llerrate, 9-11 elongate, the 9th being almost as long aA the preceding foul'
oombined, 'I'arsi gradually expnnded, penultimate joint broadly exoavate
above, the 5th short and thick, dilateo towards the extremity, so as to
be of elongate-cordate contour, with thiok olaws,
Ma("l'QnobiuTIl t1"unrnturn (1618) ill the OItl~- Hpecies thnt ill at all
simila.r, but the antennal struoture is manifestly difterent,
Length, 5 rom,; breadth, nearJy 2 mm,
Titahi Bay, Wellington, A single moulltecl specimen from Ml' • ..1,
O'Connor, nnn one. minllR legs IInll antellnae. received fl'om 'Mr, Hubert
Simmonds,
RRIW"",- !I'BU' Ot11tra and SPBCIt'~ 0/ Coleopterll. io'i!'l
and IL little COILI'sel' 111:'111' the butUl't' than clst:lwl.u.lI'e, their sides have
!.cveral coa1'8e, Rmooth I'ov(.'ne; on uaoh el~'hol1 thel'e t\l'e 4 (liRtinot tuber-
cular clcvationll: tlw lilt if, 1>111'>111, 111'111' tlll' ~C"tc'lIl1l11, th' :tllll hl'fOl'I'thl'
lIlicldll" the :ll'd on top 0[' tilt' apical d('clivit,"; thl'lIe al'e IIoMly in line;
the 4th ill pluced just outNi,}t, IlIHI II Httll' ill .If(vallel' 01' lllt' ~lId; thl'l'l'
1I1't:! 4- KlIulller ont:lll nenl' the :11'11, the innel'lnollt pail- being neltl' the Buture,
Unl1ol'sitle iuscuus, O(l\'l'l'l'c! with N\tm'j tllWIl~' 'l'lnl', IlIh'I'lllt'llink>
('U"ac Itll fllr apart all tho I~ntol'iol' pair, the posterlCll' I:llightly more
IIppro:ciUh\ted, Basal ventral segment longer than the 2nd in the middle,
cuneifol'm between thl' C(lXIlA..', 4th mtlll'!' IIhclI'tl'" than :It'Il, :ith Hiulple,
Epipleurae broad nearly to the extremity
Antennae with short bl'!lSSY setae, their basal joint twice u.s thick but
!lot much longer thun the 2nd, which iM Il.hllollt hnlf the length of the
31'd, joints 4.--8 about equal; club finely pubescent.
..\. l'ather elongatecl species, with broud, obliquely trl11loate elytral
Il.l'icell, and unusually brightly ooluured palpi,
Length, 12 mIn, j bt'eudth, 51 mm.
Mount Quoin, .My specimen was found by 1\11', A, O'Connor,
G,'uup J)UPJilRIIIAI!l.
::\253, Menimus lineatus sp, nov, Memmm i:lll.l~rp, M.lm. N.Z. Coieopt"
p.360.
Oblong-oval, l:Ilightly tl'ansvel'Rely couvex, nitid j nCIlI'I~' glabl'ou&,
there being only a few slender, eruot, illconHpiouoUB gl'(.'yildl Metne on tile
hind-body; those ou the fOl'eht:lad and legs, however, are UWl'o ellsil)
seen, as they are more elongate awl Hnvet.ICclIt; hoc1,\' N01ll0Whllt iniu8Cllte
piceo-rufouB, the lateral margins and apioa.l portion of t'lytl'a plllt',;
legs, antennae, and palpi mOl'e or less forruginous,
Head smaller than thol'a'i:, anely yet quite distinctly but not clOliely
punctured, Eyes minute, 'rhol'lIx a third bl'ou.dol' tIlan long, very
slightly ourvecUy nUl'rowed :mtcl'iurly, lIC:'uJ'ly latl'llight I.Iebilld, \\'lu~l'e
the lateral margins I\l'e 1Il0l'C oxpaJllluci thuu th('y M'O in il'out; the
npex is biAinuate, \vith obtli!!U nngles; bul:ll.' s\1btl'uncltte, olosely app1i<ld
to the elytra, its angl~'H l'ectullgullLl' bllt. not M ull 1'1'lIlllilllmt.; thl!
punctation distant, an,l I'athol' fine I' thUJ! thaI of thtl head j jUllt in
iront of the nne basal mal'gin and parallel to it there is II fil\(! lillcal'
improJHsion which appears more definite when examined lIideWI\yll, 8cu-
tellum broa.dly h'ia.ugulm', finely puncta.te. El) 1.1'80 t\\ h:o the length of
tho thora.x, of a.bout the Name width, but with the l'l1ther o.cut(.' hUUlt:ll'al
angley extending just outside the hind angles of the thora.x; they at'e
gl'adually na.t'rowed posteriorly, with 1\ correspollding diminution of the
lateral margins, which when looked I\t from above seem quite obsolete j
their punctures a.re subseriate, ra.ther coarser near the sides and suture
than those of the thorax, and beoome indistinot behind,
.Antennae stout, rather short; 2nd and 3rd joints ra.ther longer tha.n
broad, each evidently longer thun the uncovC:'l'ed portion of the IMt; 4th
quadrate; joints 5-7 of about equal length, but suoceasively expanded,
na.rrowed towa.rds the base; 8th and 9th la.rge, transverse, na.rrowed
baokwards,; 10th large, subrotunda.te,
Legs modera.tely mendel''; tibiae with minute terminal &pUl'S; basal
joint of posterior tarsi rather aborter than the 2nd and 3rd oombined.
BROUN,,-.Ve1o Gtllern and 8peries of Coleoptera, 437
Hrnup Hl'lT.nl'lDAl~,
broad. Elytro. slightly brouclC'r thaJl tho I II \ .It the h.l"\" 11('I1I'1~ thrire it~
length. n. little ",idor just bE-fore the middle thon eIHcwhere. conhi(lE'1"8bly
narrowed pm.tol·;orl,l. 11101 II (,Il-<1"\'(')0l'"t! III 1I·(.lIIl" , ('.ll'lt 1'1" t I'on hus
eight series of tulltlO1 nte PI1l1l'tUll'I., two of these B1'C quilt! Intel'nl, the
thl'C(' nenrellt the lIuturo, bohind the middll', nrc IIUhflil'ilLh,: th(' inier·
Atices a.rc finely puucturerl.
Antennnc scal'coly afl long us tho henu and thOI'I\\, joints i-II a littlE'
broa.der o.nd more dilltinctly pubescent thnll the (lth('I'H, tho e'IIIIFled pOl'
tion of the basal joint is much stOUt(ll' than i he 2ncl but IULl'dl~ at a11
longer. the next is longcl' tha.n the 4.th. the terminal olollgo.tc-oval.
In ('. ckr!l80meloides the elyh'al PUllctul'efoo nro l(,HH Ilumerous and
quite foveiform. C. gellialis (20[)9) more 11eo.lly resembles. (hit, species,
in which, howevel, the ILDtcl'ior angles of the thorax are more broo.dly
rounded and the eoides almost ovenly cUl'veu, tho el)·t ('nl punctures are
nowhere co0.1'8e, a11d ill 2059 the 4th tmil'lIl1al joint '" ,111lIO..,j ,I' Ioug Uh
the Srd. 'rhe vestiture of the front tarsi is rather d('nRe and nearly
grey.
Length, 8 rom.; breadth, Si mm.
Advance Peak, Oto.go. One found by MI'. II'. ~. Oliver, and sent to
me by Professor Chilton. The pygidium is unnl\tura.ll~ dil'ltended and
uncovered, owing to so.turo.tioll with alcohol.
Group AN'rHH'IDAE.
325ft Cotes insignis sp. nov. Oates Sharp, Man. N.Z. Colpopt .. p. 410.
Elong ....tt!, bubllepl't'llbed, clothed With sII'uder fulvcscl'nt hOll'S, head
a.nd thorax shining, do.rk l'UfOUR j elytra, at the base, also rufous, of "
pale chestnut-I'ed aoross the middle, somewhat nigresccnt behind, but
light red at the a.pex; the legs, antellno.e, and pulpi I'ufescent, tarsi
yellowish.
Head broader than the thOl'tLX, the vertex smooth, with a. few in-
distinct punctures near the eyes. Thorax quite lllJlllLllillg in langth thE'
whlth of its basal portion, decplJ' oonstl'ich'll bdlillll the mitldlo, in
(rollt of tItc contmction it is subglnboso; it ('x:hihitH 110 cletinit.c IIculp·
ture. HcutelluIn trinngullLI', El,vtl'o. not, dUllhlt' tht' breadth oC the
thol'ax, but more than thrice its length; I'lLth<!)' finely :«·t q1lit!' pel'cell
tibly, but not pOl'icctl,I' serilttcly, JllIllohu'otl f('om tll(' blLfIa t.o tho hind
femora, the PC)Rt<~I'iol' Hculpturo lIhl4Uletc; ihore ih It hlight oi>tuHe eleva-
tion on each at the bOose, nnll tho pale centrlLl }>1''II'tinn is very R1ightl~·
depressed; the pygidium is unoovered, I~nd nearly testa.ccous. '
Eyes larl:,'O, pI·omilll.'nL, ,lI1d du,tj lIotI,' 1'.I('l.'lI'l! ,\ IItl'I11HHl tltout, diS
tinotly pubescent, reaching baokwul'ds to the milllne thighH, thE-ir 2nd
joint nearly as long IlR the Jst and more tha.n uulf the length of the
31'd, the 11th sco.rcely o.ny longer tha.n the penultiUlato.
('. ru,fa only, from Mokohinau IBlanli, l'esembleH this species j it is
lUl'gel', but in 2072 the pUlIc!.ation of the wing-caBell is much nlOl'e dis-
tinot, und, before the middl~. I:L broad urea. is covered with fine golden
pubescence.
l..ength, 5f mID. j breadth, If mm.
Kaitoke, near Wellington. My specimen is another of Mr. .A.
O'Connor's novelties. In bis specimen fine white hairs almost form a
fringe across the da.rk part of the elytra.
BnUIiN N ta' Gtllaa ,"ul Specie, of Coleoptt"fII.. 4-:l!l
nlolll \II.I.I'[)fIUU\I:
:-\257. Hylobia plagiata "'p. nov. HyZobia BIOUIl, Mnn. N.Z. ('oieopl.
p. 40:\.
Conve,\, ven elongate oval, slightly nitid, fUbco-cnstancous. legs a.nd
antennae fIlRC'C;-tl·'1tu('oom., tibiae hlllf fns('olU!: ench elyhon with 3
obscure lllrll-II'"lnceOUh spotF. lit the:' ... i(l(', behintl the middle; pUUescence
cinercou<l, V('I',I linc, I'nthol' denso, unci 1,1 iJlg olose to the dOl'Dl
Head ol .Ibotlt tlw SUlllO width bchllld as the apex or thbral.., dcfiexed,
with Hat oyeR, Thoro'\ II. little 1IIol,,1cl' tll.1ll Itlll'! .... Lightly rounded late·
rally, base bisinna.tc, llith 1\ depl·l· ...hlflll bot\\I'(,1l tlu' mic.l.dle lIud eaoh
side, its Rudn.oe "ilh minut(' trnuRvl·IK.ll loocnlptul'c, "llioh in some l~htR
appears gl·annlar. Rcutellum transversel;\" quadrate. Elytra four tunes
the length of thOl'U t, of the smile" Idtll .1 ... it 1'1 '1 t thc haKe, wiciel 1 "111'
the middle. neuminnt(' poRteriol'ly; their lIuture well marked and sli~htl~
rufescent, the 1I0111ptul'e similar to that of the thorax.
Antennae elongnto, 2nd joint rather shorter than 31'd, the following
Joints vel'~' gl uduully antI slightly ('xpandeli, 9th u.nd 10th subquadate.
Spurs of put-tel illl' tihi lie clOI'('l~' }leothltltC', hlll'dl~' nM lmlS nil the ba.snl
tursal joint. .\lltel·ior tn I'Ri lnodcmtely ('long-ate, baRal joint nearly the
lellgth of tlu' ,.,nbclundmte 2nd and 31 d takcn together, 4-th subquadrate,
angularly Illllnl'ginlito nt apel, 5th mth"I' lull~t'l tlUIl1 It... IlJ'C'tleceBRol',
ha.lf its width; claws Himple. •
SOllwwhnl Him illl l' to fl. ra7/(la (715), but reanily diHtinguiHhable b~
the mOI'c elong-lltc (mfli ne, sepal'aiely acuminatc a.nd trimnculate elytra.
Length, 5 IllllI. ; bl·eo.tlth, It mm.
Ho.ywal'tl's, n('o.r Wellington. 011e illtllVlciulIl, mouute,1 011 cllrdboIU'd,
from 1\11', II, W, RillllnondR.
nearly to the baSt', jth us long as the prooedillg two, with simple slender
claws. Spurs of posterior tibiae distinctly pectinate, as long as the
dilated tibiae, but Rhorter than the basal tarllal joint.
The pl'evliling indefinite Nctllptul'e lind appal'ont absence of the Beu.
telJum are good rliRtillglliRhing ehnractcl'S.
Length, 4 mill., hl'C'urlth, 12 tnm,
Mnk.ltotC', \II. W. ,J. r,lIinneHFI, whost' llltlll(l is besto\\erl Otl this
hpecies, fOl'VI'llI'rled 1\ pllckn~(' of II'.tf Illoultl \\hich ht' lJol1pch'ci neal' the
viu(luct in F\'bruHry. 11110. BC'Ridea Rome other rare Rpecies, one of this
was lonno umoJlgRt thE' decRying lellves.
I
ConIlIlPlU(,'
nllut.
I
I Alax
1I11U..
AmlllJ-
tulip
I ~:~tlO~~ l
I
1906. I H. m. 1 H. m'l
!I
Mm. I H.m.
,1,\n. 2 Il:l·j
408·7
209·11
.j 33·jj
I I
04
0·3
1 10
051 1
,. 18 I
22
1 3jj·!)
oj tH·O 42:H I Hi I 028
044-
Thi('kilning of line.
74-6.21 o 49 Thil!lccninp; of lint·.
.. 241
24- 22 1)1·2 2204·:J (1·4 Elld bctwe('n 22h. 09·4m. and 22h.
13·611\. while aHt'ndin~ im,u'U·
., 23 L 1i:l·1) I 0111
I Slight thillk(,IUng.
ment..
DGtl'.
\
I
l.omrnf.oro-I
mlut 111 ax.
I I I
--- - -
M.>x.
ADlpll·
hull'.
J)III.,tluD.
A)JriI14, 4 02·!}
.. 18
407.51
13 33·6 ~ :1:J.J 1
1;)(1 t Oriqin, Nl'w Rcbrid(·~.
Ha.n Franci<;('CI. Su blK'cluent trcllllll'h
at 17 )7·6. 18 111,7, 11) 20·7.
1401·0
6 15·4
.. 19 1 621·6
I I 14, 30·0 I 6·7 3 21
I - I
1906. H. nl. H. m. Mm. R. m.
Aug. 17 0 33·3 Va.ll~tl·di~".
( Sul"c..'qul'n( lI'('Jnor': II 33 to '7 14-,
o 57·3 1 36·1 mmutf.': 7 41 to II ()7, minute;
1 80·8
1 44-4- ]2-0 4 ]8
.. t!) 22 to 10 12. minute; 13 2:i to
13 25. lnr~f.': ,1nel minutf.' to
14 IS·2.
" 18 656·8
70I·.j 703'5 1·85 1 39
22 !
1945·6 1948·7 5·4 1 1Pt·by Beginning ,lncl end
('ontinlloU, tnomofh.
obh('urod
. :1 144.·11
162q·6
II 111·7
Ii 30·7
] 48·S
16 30·6
(134·8
Ii 4/1·2
0·8
1·25
3·3
I
0 21
y
I P"by continuou,
Bcqinning a.nd cnll.
tn'mON.
ob~('ul'f'it
303·5
338·7 1-4 128
HI 06·0 B.P., 15
, I
Ret'.
]9 11·2 19 ]J·i 2·4 o 10
" ]4 13 34·8 ]:i I).~·O O·R End obsourod by night tremors.
.. 14 16 ]2·1
HI 19·3 HI 30·2 20·2 I
103]·7
103;;·9 t End obAourcd hy nillht tl't'nlOl'R.
'" ]1) 243·2 003
.. 111 424·2 4 211·~ o 12
.. 17 8 ':;2·8
002·6 006·2 , End obscured by nill,ht (remOl'A.
" 21 ] 28·0
149·7 1.:;s·3 o 27
.. 21 20 08·2 Minute tremor. I:!hock noted a.t
20 10·0 a.t C1\ri'4t~hllrch, Cht'vint,
o.nd Kaikoum.
.• 29 1 ]4 ]2·7 0·8 ! Probably night ll'C.'mol"o.
Ol't. 2
2
0]0·2
1 1i1l·8
o 13·8 0·3 I) :18 I B.P., 15 Heo.
.- -'17 I 9 19·]
921-1
Mal'. 13
() 40'sl 103
I
B.P.. 15 'It'r.
ml.'l'Plv.
'l'hic-l,('nill~ of Iiue
"
.:i 08·4
1.; 1 734-0 :; .1~).0 I o 07
o 10 I Very s.li"qht.
Minlll<',
" 18
.. 27 I 0;)9·1 I '1:25
o 32 I .Minute hwpllin~.
.. 29 Ii 06·8 (108.61
I 1 II 08
,. 31 ? 22 13·8
I 22 17·9 2·:; I ltn mi(\,Uc ot luqht h~'lIl11r.,.
April 7 9 ;)4·4 1002·7 l<'ol1ow('t\ by ('()lItimlClllh tn·Ill"'....
,. 13 II 32·;,
7 02·0 730·9 .H 1
.. 18 21 11·4-
21 39·H 21 4S-1l I J·5 1 39
19 007·2 .. I
(I 32·0 o 31i·2 I I ml
n :;au I 2 13·;; :2 21·H I lot
22 ? IK 1Il·7 [n midtUo uf llip,bL IN·mol"l.
.. 241 2:J 4(1·11
24 02·(1 2~ 04·7 1·1 II :;ij 1
.:iK·:i I
41 I)
II (18·7 U ·IH·O I 2·7 y
IVery
End hy Illlfht
ObsOIlWJ lrontoJ'l'l.
9 829·9 S 33·0 I o 07 H.Ilght.
J2 I ? I
8 0~·6 R 06·1 1-1
I night tremOl'R.
o.ntl C'uc1 ()bhC'ured
Beginning b"
•
. 13
21 23·;') 21 26·7 ·i.g I
I
? ,: Ditto.
t. 14: 300·S 402·9 0·.(.5 o 32 I
000
"
17
23
27
1 22·6
!
I) 08.61
1 24-6
11 21·S
0·4
0-" ? IIn middlo of minute night tremors.
513·G 5 13·!! I 1·-1. } End obseured by night tremOT&.
I) 15·8
ao , I 2204.9 I 0-7 In middle of continuous treOOOfh.
" 31 I BegiuniDg and end o'baoured by
12 :')1-81 12 ;)K.() I 4-9
0-4
II night 1:1.'em0flj.
••Tune 5 346·2 4 2.1·9
Appel/(l/:.'t.
dO mont
I
JJ~et comnll'ncr-j- All)' IJUl.lhOIl J
I
-- -------
1907. H. m. H. III. MDl. H. m.
June 13 957·2 10 10·7 0·7 1 52
" 13 12 17·9 12 21·0 0·3 o 18
.. 18 I 906·3 II 07·9 1·9 011 l!'e1t in tU\\ ru. ""utll of 'I'ilnarll.
1728·6 005 Very smlLlI oarrllt·~h.\Jll'(L
.. 19, 1903·8
~~'21
" 19
1930·7 1·0 134
" 271 ? 19 ..
2243·0 22492 is:O±1 ? In mic.lcllc· ot contiulluu", tremoN.
July Ib I 048·1 0;)1·2 0.25/ 020
" 18 ;) 011·7 ;) 10·8 0·8 o 38
.. 20 I 14- 21·:, 0·8 111 midlll(' ot 1Illl,ht !J·('llIUrtl.
" 29 000·7 1 27·3 0·6 o 21)
19 51-4 20 ll·O 0-4 II r>3
" 29
A~. 4
I 6 39·1 644·8 0·3 ]00
., ;} 702·7 704·7 0·:1 I 10
13 21 54·4
., 1 2205·7 22011·8 1 17
,. 111 357-7 Small, ~\ldllr.'ll.
18 6 17·6 U 21·7 o 15 Swelling.
11) I 600·5 6 16·8 021 Very smAll.
., 2a 7 10·0 7 13·1 028
1 23 1338·0 13 n·s 0·1 023
.. I 28 1920·8 19 2a·9 0·3 050
.. I 31 843·8 848·9 0·3 o 16
Sept. 21 16 26·3
16 54-3 HI 55·7 1-5
..
? After·tl't'lU01'll for about twu houl'll.
,. 13 528·3
532·9 IS 36·6 2·2 ]30
.. 241
24-
600·7
1638·6 17 03·8 }04.
032
034
ThicJ.ening at till!.'.
-:-ate l~o;:;;-men(e-I
nllut.
\II<"
-- - -
1907. I H.
DI,(,. 30 II
m.
4.;:~
11.
..
Ill. I Mm. H. Ill.
1908. I I
Jan. 19 7 33·7 I 7 42·0 , 0·1 0 3a
.. 20 046·5 I) OIl I Small, t·.IlJ ul .~ha.}INt.
.J 24 o 46·8 0 '1'13 0 1 ()'9 I) 56
Feb. 6 150·4 II 37 Tbickonutg of limo.
.. 6 I 0 Oil I IimA.U, (,.1ll1l1 .~h.tJ)('(I.
"
..
7
14-
600·21
256·:;
103·1
300.6 I
106.21
O·l,j
0·7
I 0240 07
" 24 23 57·8 I
24 0],5 I 24 02·/i HI 0 38
" 27 021.6 I 003 Minute.
.. 27 919·6 I 044 ~htellinp;h.
" 27 123004- J·4 111 ruilldh.- of oontinuou~ tremul'\.
Maximul1I ~Ppe&nl "t. beginniug of
qllakf'.
.. 29 21 49·6 I 21 ,3:1·8 0·4 0 J2
Mar. 5 228·8 .. ..
2 62·] 2 39·8 HI 1 ~11
I'J RcgiDWlIlI,
II
.. 13 7 •• and end ubacuretl by
10 19·4 '10 20·4 27 7 night. b ('luom
" 19 3 08·1 3 11-7 0·5 o 4.8
.. 2] 425·8 o 88 I '('blc!o."uinll. of Jinl' preceded mtd
follOWt'!l by minute tremors.
23 .. II Bt~ng cd ond obscured by
OJ
.. 26
11 46.61
23 16·3
1
12 'OtI·2
.. I
I ~:9 ! ? I J night tJ ('IDOl'b.
23 27-6 24 09·0
24 29·7 ::23 I 2 ;>0
.. 27 4 11.61 4 53·5
5 ]5·2 ~:5.5 1
207
1
I
April 7 1 26·2 1 37·5 0·:; 1 o -l.'J I B.P•• ];'1·4 ~I-.
.. 10 003.6 " I
o 20·0 1 0 3]·8 I hj.3 140
I
t. 12 1109·] {I 1~1:I 0·1 o l7
.. 12 19 L8.4' 19 311·0 I 1)·75 o LO
.. 15 60tH fllNI 0·4 ()2H
.. 23 007·0 •. I
o 09·11 I 0 30·11 1111i :2 05
May 5 Ii 28·8 I II 03·8 I 1·0 I
II 05-4 I 140
.. 20 800·9 8 U·7 0·8 ? l1'ollowl-d by nijtht. t rcmol'H.
.. 21 7 LO·l 0\;> ISl~1..
Juue 3
.. 18
Y 21
1 39·11 I ]
:2;;·:;1
4:1-7 I J0
021)
I 014
l In uliddh' of (lontinuollS t mnol"l.
Aug. 17 10 .;0·8 I
10.39·,3 11 '28.4 I .> ;; 1
I
11 34·1
Sept. 2 ~
21 1.3·8 (HI 1.Beginnmg and end obsoured by
ziight trelDOl'b.
..
..
14
21
3 '32'51
6 1SO·7
336.71
7 n·8
0'4
0·9
0 19
144
.. 22 3 n·8 3 23·7 0·35 030
.. 26 I) 27·4.
Oot. 7
II 32-0
1 00'7
II 31·0
5 34-1
1 ~1'0
I 49
0·8
1 IS
046
J
13 1 42 tiwellinlt--
It
••
Appelld loL. 4t7
Dnto.
leo::::1-
mt'lls.
-
1_ _ _ _ _ _ IlUll:"~. __
1908.
Nov. 11
H. m.
21 22·3
fr. m.
21 30·0
l\lm.
HI
H. m.
(I 511 I ~ctlt'(l",\l·lling.
h~' Ili~ht 1!'('R1fl1",
.." 15 200·1:1 o 21 Sl1!~ht
I~_l''''
22 2252·7 2238·4- 0·2.1 0 21
" 27 038·0 O"H 0·:1 0 24
.. 30 212401 21 27·2 L·9 0 28
Dec. ] 305·4- o 38
2 IS ]2·5 1.') ]1)·6 0·25 023
200·3 206·9 0·]5 0 28
~I 026·1 026·4- 6·0 Il:!evere -.ho<·k In Nut th ('a.ntorbury
" o 27·S o Iii I a.nd ~light In South Od.nterbury.
16
b 07·7 In middlc (ott I!ontinlluu" tromon..
.. 28 1 440·0 43;)·3
:s 00·;;
.:s 07·8
I Ii 20·9
530·0 0·6 2 22 Hubbl'tfUeut tlcmorat 717·]. Origin,
IS 07·2
Ii 13·4 Italy.
tI 17·1
I IS 20·2
624,·3
I
1909.
Ja.n. 1 4 1].7 I o 07 Swclhng.
.. 3 21 46·5
21 ;)1·2 21 ;'2·2 6·4- lIn progroo... wbill' d.ttending to
21 ,),1,·3 023 I m'ltrumt'nt.
17 :l 17·9 328·2 0·4 025
" 21 238·4 247·7 0·5 029
" 23
,. 28 I
.. 29
::g
g0;)9·8 'I
1 15·9 0·3
L 37
004
045
TremOR.
Thickeumg meroly.
nate
----
I
Conuul'nl'e.!
-
RlPllt
MII'C~ -I Ait. IDII"lt~II:--_11 -
tuelt'.
--4-----
It"UHI"_
:1~'91
" 30 !
21 303·9 21 2·9 Bcgi.nuiUI\ "lw.·lIll·d by .l1r·ll'l'm,,1"I.
June 3 1852·7
1924·8
U 09·7
19 37-i
tI n·8
I :2·2 2 52
(130·7 0·7
1l1i3·2
7 13·9 ? I ~;ml ub.. I'\IIl>d b~' ,lil·h,(·lIllll'.
..
9
12
o tH·O
20 2:,·G
I 004- "lOry Eolllall: maximullI at ht'ginlling.
20 29·1 20 34·3 7-5 1 36 I
.. 14 7 31-1 7 :JS'4 0·4 02.2
'0 ]5 1 19·8
935·8 936·6 0·1
040
004-
I 'L'hit'kl'llilll,1, 111('1'('1,)'.
" 28 722·0
.. 27
7 30·3 735·5 4-7 1511
.. 28 ! 1529·0 0·7 1 I Beginning and I:'nd o1r.cured by .ioU·
July 1 13 03·7 13 10·9 0·3 028 I tremorl.
62:>·9 628·5 0·1 o 31
"
" 5
2 1 17 ;)1)·3 I 3·5 ? I BeginnillA I\nd end oI1'41'uL'f'(l b~' air.
10 13 40·3 1342·8 0·4 I tremOJ",.
II
14 11·3 033 I
" 26 22 06·6 2201H o 16 I Slight thi('I,('niug.
" 27 434·G 4- 36·1 0·2 o 29
" 30 11 16·3 11 16·,'S 0·8 I !<'il'bt llUIohlllluln at 1J('~inninq'.
11 31·2 234- YOri.gin• .MlIldcoCJ.
Aug. 4 6 HI·S 620·4- o·J o OU
..
••
Ii
10
346·0
II 511-1
5 ·,19·1
701·2
I (I."
0·2:1
o II
U22
,. 13 12 HI·7 J221·7 0·25 U (lU
.. 16 754·(1 !! li·7 0·4 (l 42
,. 18 o 3,')·4- I
04(1·2 o :;11·4- I 3·9 o 2!1 I Inin'ltrumellt.
II
pl"ngl'llhf> whil(' ntt(.'ndilll( to
.. 29 23 S9.S I 34- Thiokeni'l_14 of lim:
Sept. 3
t. l!5
8 11-2
12 29-6
I
!! HI·..!.
12 ~~'7
0·35
0·5
o 19
o 14-
B.P., [I;") ....'('.
Oct. 3 1 .28-8 2 15 I 'l'hiekenillll.
.. 4 13 53·2 14 ~.7.0 I 0·9 I Y I End obF.o\trcd by air.tremol'R.
.. 21 045-5 o 5;) I 'rhiIlkeningh.
II 23 ? 21.28·9 I I 'Beginllinp, obo.ctIl't'd by a.ir·tremOl'lt.
.. 27 132·3 133·3 I 0·35 o 36
..
Nov. 3
30 1033·8
619·0
11 02·0
.. I
0·9 y I End ohhcurt'd by air·tretnOl'll.
•• 10
623·9
5 ~·6 6 ~~4 i 3·2:1 1 16
011 I '.rhickcning of line.
•• 10 626·4
6 ~·6 (131:102 I 1 57 I
.A 1'1111/(/ /I' 119
1
1!109. H. III. 1 H. III. Mill. II. 10.
Nov. 12 I 10 Ool.·.t HI (I.;.; III Il IN N('WH' ",ho.·(c; (pit ill W('\IiIlIlLon,
N"II Zc'.IIllud.
101. I I) 01.73 ,,·I.j II 07
27 21 I!)·I 112H Nhu.h( thil·k,·nin~.
h·'.'IIIIl1'i.
.. 2H 107H
I 14·;; I l::i·n Itl 1201-
~ 2~·:l H :14·41 II;) I o 111 1>1'.,('(.'(1"11 .lIIel (ollowcd by IIllnn(1l
.m·t 1'1'11101 'i.
[)Pr. a 3 :J:l3 1 I) II o 1·(1
:l :i4.!! 1
.., II 1;·1;
!I
H 23~
~
21 I I 01
I Bl'ginlliug ,\Ild ('nel OlN'II;""c( II)' .111"
'I
I.; ;IIHI
21 2:).3
Hi ;)'1 I I
1
:~751 I tJ'(·mUl~.
I -\1(1'' 1: I I
D to
a,
()omm"II("
11I0llt I Ampll·
tlld.
UIII"tIUIl. 1 RMII,'I~~
HHO.
l\1.u'. 2\) I
III. m. I H.
N IMII
I
111.
I
l\Im.
(1·2.;
II. Ill. I
111-111111 II .11111 ~Il,ldl'll. 1,,(,1t ,It l'll!i~t.
H ;~l ~ I 0·1;) I ,lilllch.
301701·1) I
170:HI 17 W·O 17-0+ 1
30 2:i Jj9·.j. 241127 O·.j. II 2k
" 31 ;, :~7·2 Ii .j.(I!l 0·2 o 1-1
April I I 1335·3 1-1, 27 I! 0·9 I :IR·1i I
" .,I, 3 2:4·7 o 08·2 I 1:lIil.\ht I hil'l\('IlIllIl.
8 1 HI 5;),2 O·!I 1 1)1 ,mel (illl.llion "h~('IlH'd h~ ,\II.
I h'l!l\ltll'~.
12 () ;;0.:11 (I ,)tI.tj I I 0.j.·4 l'rl.','C,I(~1 .lIlIl
ajl·tr~IllUl ,
t(lIl"'\I~1 h... Iln 11 Ilh'
"
Itl
20
i 34-!1
2:2 ~.2~';;
I 224;,0 I II 4l'0
I 14·'"
Thick~llill~ lit iiIII'.
Il'II'tl'(II~1 h~ Ifllltil\lIlIlI~ .lIr·tu'Il1IJi'.
" 23 1
1:l r;2·~ I·a l 1'1 ,l.Iul ,hn.llion ,,1"""lIfl'll ";V lIir·
h"IlWI~.
271 :2 10 31 ":J3·( I 'l'hi('kl·llil\l.\~.
:&~
II
-I- 44-·7 () I I·!!
1 : r I 1 PI und ,hll,l! iUIl "h~'llrotl h.1
h('mul~,
d ir
Jun(' 1
,.
Ii
:2
502·6
I' Ii ()()·7
10 30·,;
II
II 22·1)
II 12-1)
10 82-0
;; 04,6
0·7
(;,1
0·2
0·1
I 21)·2
3 00·0
000·1
027·1
Y Origin. ~'iJi.
"
II I (j '-10.0 1
9 1 8 10,3
o 011·1
o 112·r,
I Millllw·.
..
" 13113 OS· 1 I 13 13-2 0·3 () ~9·5
.. 23 10 J(Hi 10 2()·' 0·4 020·9 I
,,24 2 lis·g o J 4.·3 I \'~ry tolll.(lIt.
,. 29 10 :i!·a Jl O:H!
11 08·' ]7"0+1 I!lnd ob'!l.'ul'I.'d by 111111.1..(' f"lIl1wiull,.
" 29 I Y 1421!·9 4-2 I 1:'1 o~ul'Ctl by JlI'I'\'l'Ilin!4 ql!.\ke
~nll(J(l l.ih. "'!Hlm,
July 5 1046,0
10 ~·1 1·0 I
10 54,4 1·1
I 1050·3 O·{I 1 ·ll.O I
7826·81 8 580j ]·0
9 14·1 I 0·8 .. I
. 11 20 38·0
9 2.'i,3 ,
20 30·4
20 4.j,o
1,1
102
1,3 I
1 ~.7-fi
o 113·;; I
I
12 21 08·0 I 21 12·1 7·0
2137.7 I 3·'; J 47.21
Ap/I(·I/(1!.1
]) ~t e. (omnll'uu-,
mrnt.
I
l!ll-O. I ff. m. I Ii. 111. 1.\11111. H. III.
,TlII~ 1.:1 I 12 08:i 12 II () 1 II
12 21 M 1·1)
12 :l:i·:i 1·0 I Ill·;;
1;; 21 4lj·7 21 iiI fl 0·2 Hw(·lhu!t,.
21 ;)1l·7 0·:1 II 1,,(·8 Pre<'l'dl'!l and foUllwrd h;'l minllt~
tJ'('ntDl·,.
19 I 19 :i3·r, 10 3n·3 0·2
1952·2 0·2 022·(1
24 1;; 27·7 I:s :lS·1I 2·0 o :n·o
29 to Slj·l 11 0]·:) 2·1)
11 14·:1 2·3 1 14·3
Aug . .:1 I;) 38·3 Iii :m·;) (I·ti o O;H)
.. tOl 20 40·6 21 08·2 0·4- 038·3
" 21 I :I SU·S :; 33·(1 70
Sept. 71 7 S3·9
:; 5.') I
7 :n·4o
3·0
7·0
23H
1 21l·0
I
" 9 900·0 II 19·8 ii·1
9 28·~ S.() o .').i3 I
10 I 12 2f.!1 1233·t. 1·4 o 311·a I
Ort. 7 7 1I.;·lj 7 11·0 2·4 Eurl oll'ol'uJ.'{'!) h\ ul·-fl't'mol'6.
II 2!i·;) 1·0 o IlH) I
7 I I 12 07·9
11 rl4-4 2·2 0530
't' 8 0()·2
J2
242'0
801i·8
247-7
1·0
4-0
oI 2H·(I
10·4-
I
" 18
" 30 747·.; 7 :;a·7 2·5 o :S1·7
Nov. 9 609'1:1 632·1 17·3 30:H I
1
.. 10 1228·4 I 1240·4 2·0 I) :ro·I) I r>ura.1.ion drmbtful owiul( tn I~ir
tremOl'h.
" 26 to 5(H I » 0~·8 1(1·8
,. 26 549·2 2·0
I '.l 01.0 2·1 End obioOurad by alr·tremor..
(l38·Y 1·.3 I 327·7
Dt>c. 1 352·a 3 ;).')·4 0·(1 016·5
.. 2 320·2 326·4- 1·5 023·9
3 7.38·3 804-5 16·4
838·0 1·0 1 21·7
3 413·8 4: 30·3 0·8 021-7
4 11 05·4 11 l()'(1 17·0
3·;, 1 4Ji.(I
. 10 I II :~7·4
11 30·0
II 47·0
10 OO·~
17·r,
5·0 11 36'3 1
11 3 :;7·0 404·2 0·7 020·5 1
" 12 23 55·9 021-7 0·8 o 4.2·3
12 25·(1
.. 13
14
16
20 54-4
14 1i5·3
1258·(1
21 02·7
15 20·3
1·6
l·n
5·1
] 16·5
] 07·3
I
15,1.1'8 3':; 217'6
I
1911.
Ja.n. 2 22 59·7 23 OIHI 14-0 ] 52·7 I
.. 3 23 46·3 026·6 1·4 Damtion llnOtlJ.'ta.iJl.
" 4 105·9 I·S 233·8 I
7 233·2 300·2 1·0 1 IS·5
8 16 24-7 1627·8 0·8 \' I End obsoured by a.ir-tremors..
.. 8 92U·8 932·0 2·S I
941-4 I·;;
..
••
Feb. 7
10
16
I 1637·7
9 OS·8
9 4:2-4
16 4Sol!
B 37·8
946·5
7-4
1·0
0·8
038.21
055·0
o 4.1·4 I
012·4
452 '/' /(/,'11111" IOIlR.
'Daw
•
I('ODllnclIQCoI
mpnt
HilI.
l<'eb. 7
I H. fl
m.
:i9·!)
I 10H. (12·0
Ul. MDI.
0·(1
H.
o IU·~
Ill.
.,
wi
12
1700·11
17 07·11
7 17011
7 Ill·1i
II
].I
0·1
1 14
o 14
I 14 -I6·{I
14 :16·1
15 1448·7 4-t)
1 14 :;2 ~
I I.i 0-1:; 1
:i'(I
lHI :1 1111 •
" 211 I III ,')7'31 HI 404 I·;S () 2!!·.i I
.Iuly 12 4 111-1 I " 311·0 :3-1 .• L.W. 1)I'LIin Ih. :!H·2m.
4 -!li·o l·5
4493 I no :i 211
J9 9 H2·~ I 9 05.8 ]·2
.-\ug. II 1 26·0
I 907·3
J 28·6
1·0
0·4
••
0]7.51
I 1~l1d oMcIU'\>U b:. trenulrA.
10 o 33.41 0 41)·9 , 0·4 (I 31
111 22 :;1·2 23 18·3 0-0 I L.W. commt'noe 22b.••jIN~ln.
23 21·7 6·0
tI·o
4~51
, 23 23·7
2327-4 ti-2
'J 19 2
" 21 1 16 S8-0
lu.sl 2 24·S!
16 41-11
()'S
4·0
o 13 1 Duration a.pproximate.
.Dura.tion obl!ourod by tremors.
t.!ept. II 1 17·9 1 20·7 0·6 1 12
'J 12
1
.. ]407·7 ]'';
I
IBeginning
'liremOl'lo.
a.mI enel ol».ourod. by
" .,
4.J..,
n,Lte. II C'nIDIl<I'Urc.'
mC'ut
Ala, ~I'
• hc P.
IDUldh·i RCIII&l~'.
I
1911. a. m. H. m. Mill. I H. In.
Hept. 15
,.
I 1225·2
15 13 44·8
12 21)04
]348·1:1
(1·11
2·() II 11
I J.}nd nh~('llred by Lnllowinq quak£>.
NOTE.-Pwvinu" to 11th Ma.y. B.P. = 14·5 ber.. 1 11110. = 0·49# ~tatic tilt.
On 11th May, 1911, the new t.Yllf.' of recording appunhl... I{1vi~ a time 'KlIIle aTI.
ploxima.tely lour time& a ... open as thl' old typo, wns illlltallNI. With t.hi9 reoorder the
lime >Oa.le ib :141 mm. to tlu.' honr.
Atter 11th May. H.P. =- 16·5 ReO"'. J mm. -= 0·4" Rtati(' tilt.
Prindpal Records of Milne /:Jmmll{fl'wpk No. 20. Itt Wl'llington, New Zea-
lall'6d. (Latitude, J.l° 17' H.; longitude, 1740 -17' K Obllervl'r, George
Hogbl'n.)
(The iDbtl'llDlcml is plaecrl in oJ, lo'[lOOiaJ 1"11111 In'low d. hCllI'IO lli:o.nciing about 30 ft.
hum the edge Ilf 11 rooky eliB ",bout MI ft. hilth. ~tll"lt(lll ilobclllt 2:',0 ~'al'flA franl the
ihort'·line 0{ WC'llington Harbour.)
-. - - I -
D••
_w.• lIumt c~1
I ('Qldllll'llI ,.I-
L-
\I,.'\.
\mpil· Ihll"tll,n.
___ I . _. Iud.·
I
-I
1006.
April 14
H.
T
m. H. m. Mm. I H. m. I
B.I'.. 111·(1 'IC'('. I't.
3.lo·n I.'~.
I
4- 02·:)
4: Ob·}
405·4 Hi ..
?
1623.31
16 26-6 16
16 3l).1)
~'4: I 0·6
19 41-2
l!O 05·8
I B.P., 18'5 sec.
P s•
r 1•
I 21 47-3 21 5i.o I
.. 26 I ?
630·0 635'71
0·8
3·2
P 1 ; obscured. by tremoth.
o 4O.:s
~31
Sept, 7 1908·0 D.P., 18'0 sec. P2'
1941-'7 1·0
.. 14 1341·3 19 f Obscured by tromorc"
16 12·8 s '
162701 10 ·sO.9 I 7 86 P
BoP., 15·9 Ml'C.
.AJIJlem/loi. .
11 t. c.nmm'll('l'o-I--M-I1:'t--;I"--l-!t-~'C-lI·-I-D-ll-I.\-h-ol-'·1------Buu-.U .....
.l" I ~'.J~t_ _~~I'::.~ ___-'-____________
-~I H. Ill. H. m. I Mm. H. JIl.
Oot. 2 1 .3tHI I B.I'.. 1N·;; 'l', • Plllhl\hl~ Judi-til
I (.)U''III1.
:2 OH.i
15
2 1;)·1
7
2 :1~.91 13·0 231
P"
21203·1
13 27·9 y I 2·,; Ob"('m+,l1
1'1'
oy tremor....
" 21NII 2 '19·9 13·0 1 5(j
29 1 39·11 1:'1'
I ;;2·4- I 33·6 I 1·4 Oh..mll~d b~ tn·nun ....
30 I 19 1:1·;
11) 111·3 Hl 11)·4 1·0
19 2:Hl
Nov. Ii 20 OfH
20 17·2
20 :16·0
20 ~.:s I 2-1
I B.I'.. IN·.i ..eo. PI'
., 14 17 22·4
17 47·2 l7 r~HI !
I
111 1O·1S I I 5 2;;
11) 727·{)
~~'1 I
735·2 I
7 4:8·0 7
10·3
8 Oil·S I
8 14·3 I 3 la
I
1909. I
Maor. 17 23 17·0:1:
2327·2 23 30·4
I tHl
I .. II Or 1'.1'. ('omDlenoe :23 12-8. 01' ('ven
1·24a.bOllt J 2.'1 0;;·0.
April 10 S 40·9
3 4!1·9 I 5 51-0 I 5·6 :2 42
I .. II 26 211·0. &e.
M.a.y 30 21 20·1 :.~ - I RO}le.tt., 0.1. 23 .i5·1. ::!4 .;:1·.1. 23 J!H.,
JuneU
21 32·2 21
1009·4
~.O) ••) I 4·0
11 02-1 HI 16,8 0·1i l' 2 02-6
0·.) I
~. 1"1·oIoU('8.
11 40·4
Nov. 3 Ii 21·7 (I 26·5 2·0 o 1.; I 'l'rcmor., befOl't' Itllfillfter.
Del'. 9 1545·2 I
133.3·8
I
11l1O.
.fullo 1 530·1
Ii 40·2
I I 27
I B.P., l3·;)~. PI'
Ii. W. l'acl1i.(.', nt'ILr New H.l'!lIido...
( Prl.lba.b~v S. W. }'acitie, about .j.,j()
" 16 I 140 mile<J north of New l..ea.IMul.
Ii 34-1
tI 4.)·2 649·3 14-5
1 I,'ollowed. by l.rcmol'b allli rcpea tb
until G55 on 17th .1 une.
.. 29 10 .30·5
LL 02·11 11 06·9 12·5 2 14
I Origin, 11° S., 1720 E.
" 29 1424·11
1427·8 1429·1) 9·0 1 38 Origin••Slo S., 1731° E.
July 29 1043.71 I Pl'
10 .32·3
I Pa· .
I
10.3704 10 51:1·0 3·0 P~.
Aug. 18 1240·')
1241·1 I 1·0
I p~~
I
1 } Lot'a.L
I)nt
G,
I('OIllIIIl'Il~I'
'Ilellt
~I,ll.
.lIn,..
\1111111.
huh'
I
Ullln!luu 11'·mu.I"~.
I
.~:,'
II III·:!
I' 2ij·2
2 a:HJ II II
24!l·:J 2 14·:;
2 411·1)
;\Inv. II II 07·3
II 11·4
(; l:{·a
(I 1;i·4
2U
II HHI
44(H) I
4 ;;\·2 1
, 5HI I fin·1 17·0 I
111'('. III 93M·1
II 41·7
Sl,IH'
12 (1:1·2
12 :~7·;' 12 411·101
14 .;1;·2
La 011·0
J;; 2,j·3
I!t 11.
.1.1lI. 2 22 il3. ~ I H.P., lli·a 10('.'.
22 511·2 , (I 41 DiHtllnol.', :t buut ::l,I(I() km.
23 01·4- 12·:4
224·7
251)·7
3 32·()
:l 4R.;'
3
I 427.31
23 ;;)0(1
PI 1
:H.1H l i I'll Y
I 2242101·2
~12·7
I
1
1':1 1 jV1'11I'1'.'ht.lm.
1'4 y
::lI -1M! I :H HI·7 1'(1
17 I:J·(I I 17 .~:I·;; I 1 112 \1'."'111. 1"'iJwiIHII I4hrwk, 2-a Hell.
I H(lK~i.lt·tml. V.
III :::;·11 ,I
IIi :17·1
I .. I II 24
j
I P=
I', I J)iMIUlIC't·, ILhllut 1,2(11) kill
III :J!oHI I 2·2 1
I
I 1':1
P:.
f It 1'., ::M1·7 hi'.'.
~: L jL}:~ l
,. :U 2 16·2
2 17-41
2 11)·2 I
220·5 2 22·5 I L·4 r 0 2211bllut
p r DiHtl,nnl.', about. l,UIl km.
llny 4 1401-4
14 04-4- : ..} O:H rn:~JlOO' II bout 2, LOO 1un.
;lL~.:}1 rl ~ L} I>iK1.llncl.',
1409·7
.. 5 23 39-0 14 12·8
24 22,4 1 02 about 8,000 klI
24- 28.9 u P: B.P., H)·,) Hl.'Cl.
NOD.-1 m.m. of amplitude = 0·70" of t.il!. (..tllti(l).
.Appelldi.c . 457
Datr. 1
I
00mmrnt'l"
m!'nt.
I
I
I
MAx. \ ·e·1
- I
DUlaHun. \
1911.
-
H. m.
I
I H. 111.
I Mm. H. m.
Ju.ne 7 U ::~ I
11 r13·7 I 11 56·2
I
: : ..}
J·.3·..0 2 30
{
B. P.. ] 8·4 He!', PI'
P,.
Ps' Long Wo.Vel!.
} Origin,
Moxico.
Tim811
1203·9 Ps' Probably tra.nl:lvetae Ilnoer·
W80V8ll. tain.
" ]0 O(H
HI 59·] 117 1·2 I t Local
17 04.1 17 05.2
.. 16 14 30·2 I
1432·4
.. I
..
..}I
0.8 j I
..
0 07
.~! o~{
736·7 7 :l7-5 B.P., 12·5 sec. Origin, about 200-
Oct. 5 230 milt1B e&Ht of Wel~
7311·0 f Origin. about 500 miles from WeI·
" 17 I II 41·0 946-4 lington.
I I I
INI)JDX.
1'\Uhl
HOGBl!lN, (J.-~Jnrthcllmk(,-III·iltin,.,
in tlll' HUIlt.h-\\'('Ht P,wilit· ill lOW 1:10
HOUBEN,O, (with MKDII. H. I<'.).-llt>(·ords oi I.\<liInt· McW1l01(l'l11lhH, If)()(I-1911 .J..J.j
HowllI!:!, O.-Now Klll·aieM o£ lJf'llillojlfll'(l, with Note, oil th(' I.,~rvlu· ,11111 PUII<II'
of Homl' Nc'w ZC'nhllld ButterJli,·, . • .. .. .. .. 203
IRO:NR1DE, ANNE 1~.-'I'11l' ,\n.lt.umit·nll'ltrnrttll'· of thl' Nc'\\ Z"nlnll,l /'il'lIl1flllt· " :I:UJ
.JunNsToN, ')'. H. (with H.\IIRIMIlN, 1•• ).- Otll~ ('ulll'plion 01 .llttllriJlhll(/1I Irum III(>
Kcrnut.llecH :11.13
Kuuc, H. B.-Koll1o 1,'c·ot,tu'Oh uf t,he ('irctliIlLory Kyhtem ur IlI/IIIIII'I/l/II finY/tll
Hondier .. 241
L.\BY, T. H. (with BUBII1.oGE, l.>. W.).-'l'be NI~tllt.,· of Ihulltllll n,~y,.,. Proooeu-
inJ\II, Part L :10
L.UNG, R. M.--SoJUo Nows on th(' Bnto.ny oC the' 1:!1'l'IlHl·t· AiountainH, wit.h u. I.ihi,
of tht· I:IpocieK ('olleuwu 110
LONGI-lTAEF, G. B.-On tht· NII1I1I'IlCintUl'l.· of t.lw 1."Jlidoptrm of Nc''\\' ZI'",lolld 108
1Il.u.ooLM, ,I.-')'hl' (Jompclllition of Homl' Nl'W Zl~~lllnd Foc)(iht.Ul'fh 265
1I[ABI!ltALL, P.-No]lhelinilt· ltuub in Nt·w ZCll~l,lnd
M.aYNeR, E.-
A R('vi~ion of the ('IM,ifioo.t.ion of till' Nl'W Zt'llhllllll '11l'11drillililt Ill!
Dc~('ril)tiollh uf Nt·w Zl'lIoland upidopfc'tcI 117
Ol.IVn, W. R. B.-
List of Lichens o.m.l l"ullgi collooted in tho Kt'l'madt'c' 1... 1","e1, in I!M)!! Nfl
The (leo(.\TlI.phic IWlationHhipH of thl' HiI'd, of 1.(11'1.1 HUWI', NUI'lulk, ,111<1 tbo
Kermali('(' lAl.u.nclH
P1rrBrH. D.-
DosoriptiOWI of .Now Native I:Ipeci('H I.lf Phllolll·l'llgo.DlH 179
On Danthotoo n.uda Oolld Trioditl ThotnROni •• IllS
PIm..l>OTT. A.-Dcsoriptitln~ of Throe .New Specie... uC lJf'pitloptcrtl illS
POl'l'lilL\VlDLL, D. L.-Notcs on the Plant <loVOrilll( of ('oclflHil l"lanel ",nIl tilt'
Rugged Itllands •• 76
l'R011T. L. B.-Note" on the Nom('ncilloture of the Nt·\\ Zcalt~lul (/t·lIIlltlriC/CII'. witIl
0. De-HeriptiOIl of a. New RppciCH. PrtK'c('dinI(H, Part JI .12
ItIGG, T.-?![ol1f.an Wa.x •• 270
1'11OIIY, H. F. (with HOGDlIIN, (~.).-Ht·c'I)1'(IH Ilf Mill1t' HciHlllClj.,'l'IlpllH, JI)(16-1011 141
HI.'EIGn!l', R.-A J'relirninnry Ac('onnt uf the IAWI'I' W"iPlIl'Il (lClI'I(('
TAYr.oB.. ('I ..\&A. 'JIIIU,LIOI11NT (with ('lAHTKIWII!lI,D, 'I'. H.). - 'I'JIIl Inh·rtU''Lillri Ilf 11'011
witb th,' HiqhC'f Fntly A"idH •• •• •• •• •• :1111
rrU,I..Y.\.RD. It. .r.- NCI{l'H Oll H()JUC' 1'h·8.j.\1lI1-IIiC·K fl'llm till' K('I'llII~clc'l' 1"IIIIICIH l:lll
W.\l'l'M, L1l. ll.-
Notl'H Oil New ~·u.land l~ht'H: NIl.:.I •• " 1114
AcltliticlllH j,(, the ~'iHh t<'lIounl\ of tilt· Kt·rnlllll ..c' l~b\l)c1H. l't·lIc·{'(·clinp,H. J'nrl, I 2M
WU.D, L. J. -'I'hl' U('()I(lgy oC till' BlIlIT, .N c·w ZC'l~lo.lIIl. • :117