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Quaternary Stratigraphy and


Paleoceanography of the
Canada Basin, Western Arctic Ocean

U.S. GEOLOGICAL SURVEY BULLETIN 2080

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Quaternary Stratigraphy and
Paleoceanography of the
Canada Basin, Western Arctic Ocean

By Richard Z. Poore, Scott E. Ishman, R. Lawrence Phillips, and David H. McNeil

U.S. GEOLOGICAL SURVEY BULLETIN 2080

Climate and paleoceanography of the western


Arctic Ocean for the last 1 million years interpreted
from four marine sediment cores

UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON: 1994


U.S. DEPARTMENT OF THE INTERIOR

BRUCE BABBITT, Secretary

U.S. GEOLOGICAL SURVEY

ROBERT M. HIRSCH, Acting Director

For sale by U.S. Geological Survey, Map Distribution


Box 25286, MS 306, Federal Center
Denver, CO 80225

Any use of trade, product, or firm names in this publication is for descriptive purposes only and
does not imply endorsement by the U.S. Government.

Published in the Eastern Region, Reston, Va.


Manuscript approved for publication December 7, 1993.

Library of Congress Cataloging in Publication Data

Quaternary stratigraphy and paleoceanography of the Canada Basin, western Arctic


Ocean / by Richard Z. Poore ... [et al.].
p. cm. – (U.S. Geological Survey bulletin ; 2080)
Includes bibliographical references.
Supt. of Docs. no.: I 19.3:2080
1. Geology, Stratigraphic–Quaternary. 2. Paleoceanography—Canada Basin.
I. Poore, Richard Z. II. Series.
QE75.B9 no. 2080
[QE696)
557.3 s—dc20
[551.46'083327] 93–50.143
CIP
CONTENTS

Abstract........................................................................................... 1
Introduction...................................................................................... 1
Acknowledgments ........................................................................ 4
Lithostratigraphy................................................................................ 4
Unit J ....................................................................................... 4
Unit M...................................................................................... 4
Units L, K, and I......................................................................... 7
Magnetostratigraphy............................................................................ 7
Foraminifers ..................................................................................... 7
Paleoceanography............................................................................... 10
Glacial-Interglacial Cycles .............................................................. 10
Deep Waters............................................................................... 11
Conclusions...................................................................................... 15
References Cited................................................................................ 16

FIGURES

1. Maps showing location of Northwind Ridge in western Arctic Ocean and coring sites discussed in text........ 2
2. Diagram showing coarse-fraction data and assignment to Arctic lithostratigraphic units for Northwind
Ridge cores 9, 5, 3, and 7 and central Arctic cores 228 and 270....................................................... 5
3. Photograph showing brown beds and color banding in parts of Northwind Ridge cores 9, 5, 3, and 7.......... 6
4. Diagram of magnetostratigraphy, lithostratigraphy, and foraminifer abundance data for Northwind
Ridge core 5 and oxygen-isotope stages of the deep sea.................................................................. 8
5. Diagram showing foraminifer abundance data for Northwind Ridge cores 9, 5, 3, and 7.......................... 9
6, 7. Graphs showing abundance variation of selected benthic foraminifers in Northwind Ridge cores:
6. Core 5.......................................................................................................................... 13
7. Core 3.......................................................................................................................... 14

TABLES

[Tables 2 and 3 follow References Cited]

Core designation, core length, water depth, and coordinates for Northwind Ridge cores 9, 5, 3, and 7
from the western Arctic Ocean ................................................................................................ 3
2, 3. Benthic foraminifer abundance, in percent, in samples from Northwind Ridge cores: -

2. Core 5.......................................................................................................................... 18
3. Core 3.......................................................................................................................... 27

III
IV CONTENTS

CONVERSION FACTORS

Multiply By To obtain

Length
micrometer (plm) 0.000039 inch
millimeter (mm) 0.0394 inch
centimeter (cm) 0.3937 inch
meter (m) 3.281 foot

Mass

gram (g) 0.0353 ounce avoirdupois

Age designation.—Carbon-14 dates are expressed as yr B.P., meaning years before


present (A.D. 1950).
Quaternary Stratigraphy and Paleoceanography of the
Canada Basin, Western Arctic Ocean

By Richard Z. Poore," Scott E. Ishman," R. Lawrence Phillips,” and David H. McNeil’

ABSTRACT topographic highs in the central and western Arctic recov


ered sedimentary sections with extremely low average
Lithologic, paleontologic, and paleomagnetic analyses accumulation rates (about 1 to 2 mm/1,000 yr; for example,
of sediment cores from Northwind Ridge in the western see Clark and others, 1980; Scott and others, 1989). Age
Canada Basin provide a stratigraphic framework for inter and environmental interpretations of these cores are com
preting the upper Quaternary paleoceanographic record of plicated by the low resolution of the records. The cores
the western Arctic Ocean. Analyses of the cores, which available from the deep Canada Basin sampled Holocene
were taken over a depth range of 945 to 3,513 m, reveal turbidites (Hunkins and Kutschale, 1967; Campbell and
distinct cycles in the abundance of planktic foraminifers that Clark, 1977; Goldstein, 1983), for which sediment accu
we interpret as glacial-interglacial cycles. Variations in mulation rates range from 4 to 462 mm/1,000 yr, with an
preservation of foraminifers and changes in benthic foram average of 83 mm/1,000 yr (Campbell and Clark, 1977).
inifer assemblages through time and along the depth However, the turbidites contain much reworked material,
transect show that the carbonate compensation depth and interpreting a paleoceanographic record from them is
became deeper during successive interglacial stages of the very difficult.
late Quaternary. The abrupt abundant appearance of Ori The U.S. Geological Survey obtained a suite of piston
dorsalis in benthic assemblages between 3,513 and 945 m cores in 1988 from the U.S. Coast Guard Cutter Polar Star
depth during an interglacial stage that we correlate with in the area of Northwind Ridge, which is a prominent
oxygen-isotope stage 9 marks the development of near feature extending from the Chukchi Sea continental slope
modern deepwater structure in the western Arctic. into the Canada Basin of the western Arctic Ocean (fig. 1A,
B). The cruise was part of an ongoing geologic sampling
and geophysical program designed to understand the geo
logic evolution and oceanographic history of the western
INTRODUCTION Arctic Ocean. Initial examination of the 1988 Polar Star
cores indicated that they contained a relatively complete
The Arctic Ocean is an extremely important compo record of the last million years and that accumulation rates
nent of the Earth's climate system, yet the paleoceano in some cores were higher (>4 mm/1,000 yr) than those for
graphic and paleoclimatic history of the Arctic is still poorly existing Arctic cores collected from topographic highs.
known and controversial. Documenting the Pliocene to Thus, these cores preserve a more detailed, yet still con
Quaternary history of the Arctic Ocean has been hampered densed, paleoclimatic and paleoceanographic record of the
by lack of adequate material for study and by the perennial western Arctic Ocean.
ice cover that limits access to many areas of the Arctic. This study presents the stratigraphy and environmental
Most coring in the central and western Arctic has been done interpretations of four cores recovered from the crest and
from floating ice stations that cannot be directed toward a east flank (Northwind Escarpment) of Northwind Ridge in
particular area or objective. In general, only short piston 1988 (fig. 1C, table 1). The cores form a transect that
and gravity cores are available, and most cores from extends from 945 m down to 3,513 m depth. Cores 9 and 5
are from near the crest of Northwind Ridge on opposite
sides of a central depression that extends along the ridge
axis. Cores 3 and 7 are from the Northwind Escarpment.
'U.S. Geological Survey, Reston, VA 22092.
*U.S. Geological Survey, Menlo Park, CA 94025. Cores 9 and 5 recovered relatively long and apparently
*Institute of Sedimentary and Petroleum Geology, 3303 33rd St., undisturbed sections—626 and 476 cm, respectively. The
NW, Calgary, Alberta T2L 2A7, Canada. sections recovered in cores 3 and 7 are complicated by
2 QUATERNARY STRATIGRAPHY AND PALEOCEANOGRAPHY, CANADA BASIN, WESTERN ARCTIC OCEAN

Sea of
Okhotsk


u -
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- 300 CESAR
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Location of
study area
INTRODUCTION

C
159°00 158°00 157°00' 156°00'

75°00'
S
Z
S
&
S’
S

74°50'

74°40' -

74°30 -

74°20'
0 20 KILOMETERS
——

_f Figure 1. Location of study area (A, B) and four coring sites on Northwind Ridge in western Arctic
Ocean (C). Bathymetry contours in meters. A, Setting of the study area (at star) near the North Pole. Base
map modified from Mapart Update 92 (MicroMaps Software, Lambertville, N.J.). B, Location of the
study area on Northwind Ridge from which the U.S. Geological Survey collected the four cores described
in this report. CESAR indicates area where cores were taken during the Canadian Expedition to Study the
Alpha Ridge (CESAR) (Aksu, 1985; Scott and others, 1989). C, Location of Northwind Ridge coring
sites (solid dots) discussed in text. B and C modified from Phillips and others (1992, fig. 1).

Table 1. Core designation, core length, water depth, and coordinates for Northwind Ridge cores
9, 5, 3, and 7 from the western Arctic Ocean.

Core designation Core length Water depth Coordinates

Full Brief (cm) (m) Latitude (N.) Longitude (W.)

PI-88—AR—P9 9 626 945 75°01.38' 158°14.23'


PI-88–AR-P5 5 476 1089 74°37.35' 157°53,04'
PI-88—AR–P3 3 581 1909 74°35.60' 157°39.59'
PI-88—AR-P7 7 536 3513 74°37.76' 157°23, 17'
4 QUATERNARY STRATIGRAPHY AND PALEOCEANOGRAPHY, CANADA BASIN, WESTERN ARCTIC OCEAN

internal erosional unconformities. However, most of core 3 UNIT J


and the upper part of core 7 do not contain discernible
sediment gaps (Phillips and others, 1992). Pleistocene Unit J, with its upper and lower sand peaks, is one of
sediments recovered from Northwind Ridge consist of the keys to identifying the lithostratigraphic sequence. In
unconsolidated, interbedded mud, silty mud, sandy mud, cores from the central Arctic, the lower sand peak of unit J
and gravelly sandy mud. The cores exhibit distinct color is characterized by a thin, but well-defined, pinkish-white,
banding due to intercalation of pinkish-white to white, clast-rich layer (PW1 of Clark and others, 1980), and the
clast-rich layers, dark-brown muddy and sandy mud beds,
upper sand peak typically contains scattered pinkish-white
and gray to tan muds and silty muds. Calcareous microfos
clasts (pink-white clast interval of Clark and others, 1980).
sils are concentrated in the dark-brown beds, most of which
are bioturbated and contain variable amounts of ice-rafted At Northwind Ridge, near the edge of the Arctic Basin,
sediment. PW1 is well developed at the base of unit J, and the
pinkish-white clast interval is present as a well-defined bed
at the top of unit J in cores 5 and 7. Pinkish-white clasts are
ACKNOWLEDGMENTS not present in the coarse-fraction peak at the top of unit J in
core 3. An unconformity occurs between units J and L in
We thank Kevin Foley (U.S. Geological Survey, core 3, and the top of J and all of unit K are missing (see
USGS) for technical assistance throughout this study and Phillips and others, 1992). The lower coarse-fraction peak
David L. Clark (University of Wisconsin) for discussions and PW1 are well represented in core 3 (fig. 2).
about the lithostratigraphy of our cores. The manuscript
benefited from comments by Debra Willard, Paula Quin
terno, and Arthur Grantz (all of USGS). We thank Meyer UNIT M
Rubin (USGS, Reston, Va.) for preparing graphite targets
used for carbon-14 dating of upper layers of cores 9, 5, and Unit M is another key unit. As defined by Clark and
3. Dating of samples from core 5 was done at the National others (1980), unit M commonly has an upper and a lower
Science Foundation Accelerator Facility for Radioisotope
sand peak; the lower peak is marked by a pinkish-white,
Analyses at the University of Arizona, Tucson, Ariz.
clast-rich layer. They noted that this clast-rich layer (PW2)
Dating of samples from cores 9 and 3 was done at the
is commonly distorted and intermixed with the surrounding
Center for Accelerator Mass Spectrometry, Lawrence
sediment through an interval of up to 15 cm. In the
Livermore National Laboratory, Livermore, Calif.
relatively expanded sections recovered from Northwind
Ridge, PW2 has been identified as two separate 1- to 2
cm-thick, clast-rich beds approximately 5 to 10 cm apart.
LITHOSTRATIGRAPHY Following consultation with D.L. Clark, the lower limit of
unit M in Northwind Ridge cores has been placed at the top
of the first dark-brown, foraminifer-rich muddy sand bed
Clark and others (1980) defined informal standard
just below PW2. Thus, in Northwind Ridge cores, PW2
lithostratigraphic units for the Arctic based on analyses of occurs near the bottom but not at the base of unit M. D.L.
cores taken from the T3 Ice Island. These standard litho
Clark (oral commun., 1993) placed the base of unit M on
stratigraphic units have been recognized and used for
top of the brown bed below PW2 because the underlying
correlation throughout the central and western Arctic Basin
(Morris and Clark, 1986; Darby and others, 1989; Scott and unit L always has a dark-brown bed at its top. The split
others, 1989). The lithostratigraphic units of Clark and PW2 in our cores is readily traced among cores 9, 5, 3, and
others (1980) have been identified in the Northwind Ridge 7 (fig. 3). The split PW2 shows up as a double peak in the
sedimentary sequence (Phillips and others, 1992) and form coarse-fraction plots for cores 9, 3, and 7 (fig. 2). No
a primary means for correlating between cores. Sediment double peak is evident in the coarse-fraction plot of core 5
texture, pinkish-white to white, clast-rich beds, and dark because of burrowing and the closer spacing of the PW2
brown muddy and sandy mud beds are important features beds (fig. 2).
for identifying these lithostratigraphic units. Figure 2 shows The upper sand peak of unit M in central Arctic cores
the percentage of coarse material (>63 p.m.) in Northwind commonly contains one or two white layers that have been
Ridge cores and sand-sized content of selected cores used designated W2 and W3 (Clark and others, 1980; Minicucci
by Clark and others (1980) to define the lithostratigraphy and Clark, 1983). The white, clast-rich beds W2 and W3
for the central and western Arctic Ocean. The coarse are present in all four of our cores. The upper and lower
material in Northwind Ridge cores is mostly sand. Detailed coarse-fraction peaks, the presence of W2 and W3, and the
logs and photographs of cores 9, 5, 3, and 7 were provided split PW2 allow ready identification of unit M in cores 9, 5,
by Phillips and others (1992). 3, and 7 (see figs. 2 and 3).
Cosearsiceg-fnrmtaceatngioten units
2.
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6 QUATERNARY STRATIGRAPHY AND PALEOCEANOGRAPHY, CANADA BASIN, WESTERN ARCTIC OCEAN

PW2

Unit M
Unit L

Unit M
Unit L

Figure 3. Parts of Northwind Ridge cores 9, 5, 3, and 7. In each core, the two light bands are the split PW2 bed, which
is clast rich. Centimeter scales are visible along the cores.
FORAMINIFERS 7

UNITS L, K, AND I and that the Jaramillo normal polarity event is in the lower
part of unit J to the upper part of unit I (see fig. 4). These
A distinct, 20- to 25-cm-thick, pinkish-white to white, results match compilations of paleomagnetic results from
clast-rich bed, not recognized in previous Arctic cores, Arctic cores, which show that the Matuyama-Brunhes
occurs within the interval we have assigned to unit L. This polarity reversal horizon typically falls within the middle
pinkish-white to white, clast-rich layer, labeled NPWB on part of unit K and that the Jaramillo is associated with the
figure 2, occurs near the middle of unit L in cores 9, 5, and lower part of unit J (for example, see Darby and others,
3. An alternative stratigraphy for our cores would be to 1989).
interpret NPWB as PW2 and to consider the split, clast-rich
beds discussed above as a new clast-rich unit. Interpreting
NPWB as PW2 would change the lower boundary of unit M FORAMINIFERS
but would not change correlation among our cores.
Several thin, pink or pinkish-white, clast-rich beds are
The abundance of foraminifers (in number per gram
present in the lower part of unit L in core 7. However, unit
dry weight) and a census of planktic foraminifers were
L in core 7 is complicated by one or more unconformities,
obtained from the -150-plm size fraction of samples from
and the multiple pink to pinkish-white beds seen in core 7
cores 9, 5, and 3 and the upper part of core 7 (fig. 5). We
were not observed in any of our other cores from Northwind
selected the P 150-plm size fraction to make our planktic
Ridge. The multiple pink to pinkish-white beds in unit L of
foraminifer census data consistent with published modern
core 7 may represent very local depositional features.
core-top data (for example, Kipp, 1976; Kellogg, 1977) and
Units L, K, and I do not have distinctive textural
to minimize problems of juvenile and unidentifiable speci
signatures like units M and J but can be identified from
mens in our census data. For samples containing abundant
other characteristics and stratigraphic position. As delin
planktic foraminifers > 150 p.m., a split of the total sample
eated by Clark and others (1980), unit L has a high but
containing approximately 300 planktic specimens was used
variable sand content, and foraminifers can be abundant in
for the faunal census and for calculating abundances of
the middle to upper part of L but are rare in the lower part
planktic and benthic foraminifers. For samples containing
of L. Unit K is characterized by a lower sand content than
=300 planktic foraminifers, all planktic and benthic foram
units J or L and commonly contains abundant planktic
foraminifers. inifers were picked from the -150-plm fraction and used to
calculate foraminifer abundances and for the planktic
Unit L, as identified at Northwind Ridge, shows a high
foraminifer census. Inspection of the 63- to 150-plm frac
but very variable coarse-fraction content. Dark-brown,
tion of our samples indicates that the abundance data for the
foraminifer-rich beds occur in the upper part of L in cores 9,
> 150-plm size fraction accurately reflect changes in abun
5, and 3. Foraminifers are essentially absent below the top
dances of foraminifers in the P63-plm fraction.
of unit L in core 7 because of dissolution (see discussion
Census data for planktic foraminifers were tabulated
below). The expanded section recovered in core 9 ends in
unit L.
by Foley and Poore (1991, 1993). These data show that
Neogloboguadrina pachyderma constitutes over 90 percent
Unit K is distinguished from units L and J in core 5 by
of the planktic assemblages in all samples; left-coiling
the presence of upper and lower dark-brown, foraminifer
Neogloboguadrina pachyderma usually exceeds 95 percent
rich beds and a slightly lower average coarse-fraction
of the N. pachyderma population. The remainder of the
content. Unit K is distinguished from units L and J in core
fauna consists of intergrades between N. pachyderma and
7 by a distinctly lower coarse-fraction content. Unit K is not
N. dutertrei (P-D intergrade of Kipp, 1976; dupac of Foley
present in core 3. The top of unit J in core 3 contains
and Poore, 1991). Inspection of the 63- to 150-plm fraction
slightly inclined laminated beds that are truncated at the
shows that Globigerina egelida and Turborotalita quinque
contact between J and L. Slumping or erosion probably
loba are present in a few samples from the upper part of unit
removed unit K from the sequence recovered in core 3.
L and from unit M; however, Neogloboguadrina pachy
Unit I was identified in cores 5, 7, and 3 primarily on
derma is still the dominant member of the assemblages in
the basis of stratigraphic position below unit J. Unit I is
the 63- to 150-plm size fraction. We observed no first or last
complicated by flow-in problems in core 3 and is not plotted
occurrences of planktic foraminifers in the Northwind
on figure 2.
Ridge cores that would be considered standard biostrati
graphic events.
Previous studies of benthic foraminifers in deep-sea
MAGNETOSTRATIGRAPHY sediments have used a variety of methods and size fractions.
Our examination of benthic foraminifers in Northwind
Paleomagnetic data from core 5 (Poore and others, Ridge cores incorporated different faunal census methods as
1993) indicate that the polarity reversal horizon represent part of an effort to find an optimum procedure for future
ing the Matuyama-Brunhes boundary occurs within unit K, work. Splits of approximately 300 specimens -63 plm were
8 QUATERNARY STRATIGRAPHY AND PALEOCEANOGRAPHY, CANADA BASIN, WESTERN ARCTIC OCEAN

Core 5
Standard Arctic 0xygen-isotope stages
Polarity lithostratigraphic Planktic foraminifers 6'80 (std. dev. units, Imbrie
ZOnes units Depth >150 plm per gram and others, 1984)
(cm)
6000 2 1 0 -1 -2
15,186 yr B.P. ººzze
7,800 yr B.P. 2"

100

N 200

| 300

benthic foraminifers
400

; > benthic foraminifers

500
0 300
Benthic foraminifers
>150 pm per gram

Figure 4. Magnetostratigraphy, lithostratigraphy, and foraminifer abundance


data for core 5 and relation of foraminifer abundance to generalized deep-sea
oxygen-isotope record. Composite deep-sea oxygen-isotope record is aligned so
that the Matuyama-Brunhes polarity reversal horizon in core 5 coincides
approximately with the stage 20-stage 19 transition. Figure modified from
Poore and others (1993).

counted for selected samples in core 5 that contained progress on box core top samples from Northwind Ridge
abundant benthic foraminifers. Benthic census data for the shows that the P.106-plm and > 125-plm benthic foraminifer
remaining samples with abundant foraminifers in core 5 and census data provide the same basic oceanographic informa
all samples with abundant benthic foraminifers in core 3 tion as the P63-plm census data. Thus, we plan to use
were obtained by tabulating the first 200–300 specimens in quantitative census of the P 125-plm size fraction in our
strewn slides. Tabulation of specimens began with the future studies of foraminifer assemblages from the Arctic
> 150-plm size fraction and extended to the 63- to 150-plm Ocean.
size fraction only when it was required to obtain >200 The planktic foraminifer abundance data show that
specimens. All benthic foraminifers were picked from the foraminifers are concentrated in distinctive layers (fig. 5)
>63-plm fraction of samples, which contained rare benthic that generally coincide with dark-brown beds. Three inter
specimens. After cores 5 and 3 were completed, represen vals of abundant planktic foraminifers occur in unit M (M1,
tative samples from cores 9 and 7 were scanned for M2, and M3) and in unit L (L1, L2, and L3). The interval
comparison with the assemblages noted in cores 5 and 3. labeled L3 in core 3 contains relatively few foraminifers
Census data for benthic foraminifers in cores 5 and 3 (<500/g) along with common iron manganese grains that
are presented in tables 2 and 3, respectively. Work in are molds and internal casts of foraminifers. Thus, L3 in
3,513 -150
=
depth
Water
m foraminiſers
Planktic
Depth
um
| Foraminifer
5.
Figure
K,
L,
M,
1993).
(1991,
Poore
Foley
from
are
Abundance
7.
and
3,
5,
9,
cores
Ridge
Northwind
for
data
abundance
aiothroasre
I
lJ,
fArctic
planktic
designate
L3
L2,
L1,
M3,
M2,
M1,
2).
fig.
text
(see
(1980)
others
and
Clark
of
units tmraitnigirafpehirc from
liathuoysare
within
peaks
Mabundance
aunits.
5
core
for
event
normal
Jaramillo
and
horizon
reversal
polaritytmra-tiBgruanpheisc
foraminifers
Benthic
pm
-150 raminifer plot
Boenthic
]
f[
plot foraminifer
Planktic
[]

gram
per
5,000 EXPLANATION
7
Core
gram
per Brown
E
beds

o
400

0
(cm)
0 100 200 300

1,909 -150
=
depth
Water
m foraminifers
Planktic
Depth
pm
M2
foraminifers
Benthic
um
-150

0
(cm)
gram
per
5,000
3
Core
gram
per

F--

o
400

10 20 30 40

1,089 -150
=
depth
Water
m foraminiſers
Planktic
um

foraminifers
Benthic
um
~150

(cm)
5,000
dgram
per
5
Core
gram
per

4o
00

Depth
foraminifers
Planktic
um
-150
0

horizon
reversal
polarity
Matuyama-Brunhes
->
polarity
normal
Jaramillo
event

945
=
depth
Water
m
foraminifers
Benthic
um
-150
400

9
Core
gram
per
rT— (1993).
others
and
Poore

o o
Depth
(cm)
100 200 300 400 500 600–
10 QUATERNARY STRATIGRAPHY AND PALEOCEANOGRAPHY, CANADA BASIN, WESTERN ARCTIC OCEAN

core 3 is the residue of a foraminifer abundance peak. Core mun., 1993) show that ice cover eliminates or greatly
5 shows two foraminifer abundance peaks in unit K. Core 7, curtails biogenic and lithogenic sedimentation and that
which was taken from a depth of 3,513 m, contains sedimentation resumes as soon as open-water conditions
significant numbers of foraminifers in unit M and the top of return. Productivity is highest near the ice margin. The
unit L (L1). Foraminifers from deeper intervals in core 7 are Northwind Ridge area of the western Arctic has 1 to 2 m of
rare and, where present, are corroded and pitted. The top of ice cover during most of the year but is near the northern
core 7 was lost during coring, and the upper foraminifer margin of waters that are normally ice free for several
peak represents foraminifers in burrows that extended down months in the late summer (Untersteiner, 1990). Leads and
from the unrecovered surface unit. Burrows with brown
polynyas of open water were observed over Northwind
infilling are evident on the split face of the core. The peaks Ridge during the 1988 cruise, and brown, planktic
M2, M3, and L1 are well defined.
foraminifer-rich muddy sands were recovered in the tops of
The stratigraphic distributions of the planktic foramin
most piston and gravity cores from Northwind Ridge.
ifer abundance peaks in units M and L are very similar in Analyses of Neogloboquadrina pachyderma from the sur
cores 9, 5, and 3. The poor preservation of L3 in core 3 and face foraminifer-rich bed in core 5 yielded carbon-14 dates
the absence of L2 and L3 in core 7 represent increased
of 5, 186+91 yr B.P. at 0–2 cm depth and 7,800+ 118 yr
dissolution reflecting the deeper location of cores 3 and 7
B.P. at 4.6–5.6 cm depth (fig. 4). The age of 5,186 yr B.P.
(see following discussion). The similarity of the records in
for the top interval probably reflects the effects of burrow
the shallow cores and the upper part of core 3 support the
ing and the loss of the uppermost sediment during piston
lithostratigraphic correlations and indicate that the foram
coring. A carbon-14 date of 8,480+80 yr B.P. was
inifer abundance variations reflect regional oceanographic obtained from the 3-cm level of core 9, and a date of
events and are not features caused by localized sedimentary 4,830+ 100 yr B.P. was obtained from the 1-cm level of
processes on Northwind Ridge. core 3. Thus, the brown, planktic-foraminifer-rich surface
sediments of Northwind Ridge were deposited during the
current interglacial stage. Seasonally open water results in
PALEOCEANOGRAPHY high productivity, abundant planktic foraminifers in the
water column, and preservation of planktic foraminifers in
GLACIAL-INTERGLACIAL CYCLES the underlying sediments. By analogy with surface sedi
ments, we infer that the repeated planktic-foraminifer-rich
The environmental significance of foraminifer abun beds occurring at depth in Northwind Ridge cores record
dance variations in Arctic cores is controversial (see Her past interglacial stages.
man and others, 1989, for summary). One interpretation The positive correlation of planktic and benthic foram
concludes that foraminifer-rich intervals represent intergla inifer abundances throughout most of the shallow-water
cial conditions reflecting seasonally absent or reduced ice cores 9 and 5 and in core 3 (fig. 5) indicates that the major
cover leading to increased productivity; intervals barren or variations in planktic foraminifer abundances in these cores
nearly barren of foraminifers are considered to represent are not caused primarily by variable dissolution of sediment
glacial conditions with thicker ice cover and lower produc with a similar foraminifer content. Neogloboguadrina
tivity (Clark, 1971; Aksu, 1985). An alternative interpreta pachyderma is very resistant to dissolution (Berger, 1979;
tion concludes that foraminifer-poor intervals represent Malmgren, 1983), but many benthic foraminifers are even
climates warmer than today when low-salinity surface more resistant to dissolution (Berger, 1973, 1979). Abun
waters due to increased river runoff or melting ice resulted dance variations due primarily to differential dissolution of
in low productivity of planktic foraminifers (Herman, sediments having initially similar foraminifer components
1983). Foraminifer-poor intervals have also been inter would result in variable ratios of planktic to benthic
preted as dissolution events occurring during interglacial foraminifers and would include associations of common,
and early glacial intervals (Morris and Clark, 1986). Many robust benthic foraminifers with low numbers of planktic
factors, including dissolution, surface-water productivity, foraminifers. Additionally, no major changes in fragmen
downslope transport, and dilution by ice-rafted material, tation were observed, which would be expected if the
influence the abundance of foraminifers in Arctic sedi observed abundance fluctuations in the shallow cores were
ments. Interaction of these factors is complex, and no single caused primarily by changes in dissolution intensity (Malm
factor can explain all of the variability seen in Arctic gren, 1983). Thus, we interpret the major variations in
sediments. We conclude that the foraminifer-rich intervals planktic foraminifer abundances in units M through K of
in the Northwind Ridge cores included in this study repre cores 9, 5, and 3 to be primarily a function of productivity
sent interglacial deposits for the following reasons. related to ice cover rather than a function of changes in
Sediment-trap studies in the Fram Strait (Hebbeln and dissolution.
Wefer, 1991) and the Norwegian-Greenland Sea (William The record from core 5 extends through the Brunhes
Curry, Woods Hole Oceanographic Institution, oral com into the Matuyama. The number and stratigraphic distribu
PALEOCEANOGRAPHY 11

tion of interglacial, planktic-foraminifer-rich horizons in unit I could be dissolved remnants of foraminifer-rich beds
core 5 (fig. 4) are similar to the number and stratigraphic representing interglacial events.
distribution of oxygen-isotope minima (representing inter The Northwind Ridge data show that distinct cycles in
glacial stages) recorded in the isotope records of the the abundance of foraminifers occur throughout the Brunhes
Brunhes interval of marine cores at lower latitudes. Most Normal Polarity Zone. Our interpretation of the foraminifer
lower latitude records indicate that the Matuyama-Brunhes abundance variations as resulting from glacial-interglacial
polarity reversal horizon is associated with the stage 20–19 cycles implies that, with the exception of major interglacial
transition and that interglacial stage 21 occurs near the end stages, the Arctic Ocean since stage 21 has been covered by
of the Matuyama (Imbrie and others, 1984; Shackleton and thick sea ice, which resulted in low productivity of foram
others, 1990). We infer that the foraminifer abundance peak inifers. The Northwind Ridge Brunhes record of foraminifer
just below 3.5 m in core 5 represents stage 21 and the abundance fluctuations supports previous conclusions that
foraminifer peak that begins just above the base of the unit M represents oxygen-isotope stages 1–8 and that stage
Brunhes, around 3.4 m, represents stage 19 (Poore and 9 occurs at the top of unit L; these conclusions were based
others, 1993). We correlate the top four foraminifer abun on variations in sand-fraction data (Clark and others, 1980)
dance peaks with stages 1, 5, 7, and 9. Matching the and limited oxygen-isotope data from Arctic cores (Aksu,
remaining foraminifer abundance peaks with specific iso 1985; Scott and others, 1989). Our results also support the
tope stages is equivocal. Inspection of figure 4 indicates that conclusions of Boyd and others (1984) that climate varia
several plausible correlations between the foraminifer peaks tions in the Arctic are in phase with glacial-interglacial
and interglacial stages are possible. cycles observed in other proxy records of the Brunhes. The
Previous records from the central and western Arctic Northwind Ridge data are too limited to speculate on
pre-Brunhes conditions and their relation to events within
indicated that foraminifers are generally abundant through and outside of the Arctic Ocean.
out most of unit M and in the upper third of unit L (Clark,
1971; Aksu, 1985; Scott and others, 1989). In contrast, the
relatively expanded record in our cores demonstrates that DEEP WATERS
foraminifers are concentrated in discrete layers that we
correlate with global interglacial stages. Abundance varia Changes in bottom-water conditions and structure
tions of planktic foraminifers (> 150 p.m.) in the relatively through the late Pleistocene are indicated by differences in
expanded section of T3–67–11 (Darby and others, 1989), preservation of foraminifers and by changes in benthic
from the slope of the Mendeleyev Ridge, also indicate that assemblages within and between cores from different
foraminifers occur in discrete beds separated by intervals depths. Although most of the variation in foraminifer
that are barren or nearly barren of foraminifers. The abundances seen in cores 9, 5, and 3 and the uppermost part
differences between foraminifer occurrences reported by of core 7 reflects changing productivity, significant disso
Clark (1971), Aksu (1985), and Scott and others (1989) and lution is evident in much of core 7 and in the lower parts of
those found in our study may represent regional variations cores 5 and 3. Dissolution effects are seen by comparing the
in the response of the Arctic to climate cycles or may be due foraminifer abundances in cores from different water depths
to nonlinear sediment accumulation rates within and (fig. 5). Foraminifer peaks M1, M2, M3, and L1 are in all
between cores. It is likely that bioturbation and low accu four cores. Foraminifer peak L2 is present in cores 9, 5, and
mulation rates have blurred the foraminifer record in many 3. Peak L2 is not recorded in core 7. However, L2 may be
central and western Arctic cores. For example, the com missing from core 7 because of erosion. L3 is well defined
bined thickness of unit M and unit L in CESAR (Canadian in cores 9 and 5 but is represented by a residual assemblage
Expedition to Study the Alpha Ridge) core 14 from Alpha in core 3. Some calcareous specimens are present, but most
Ridge is about 42 cm (Scott and others, 1989), whereas the of the foraminifers are preserved in core 3 as iron manga
thickness of units M and L in Northwind Ridge core 9 is 626 nese molds and internal casts. No indication of L3 was
Cm. observed in core 7. Thus, during deposition of L3, the
Two foraminifer-rich beds occur in unit K of core 5 carbonate compensation depth (CCD) was near but just
(fig. 4). The section below unit K in core 5 does not contain below the depth of core 3. Unit K was identified in cores 5
foraminifer-rich horizons, although the magnetostratigra and 7. Two large peaks of foraminifer abundance occur in
phy shows that this interval includes the Jaramillo normal unit K of core 5, whereas, in core 7, unit K is essentially
polarity event and thus spans several hundred thousand barren of calcareous foraminifers. Unit I of core 5 contains
years, during which several glacial-interglacial cycles two intervals with low abundances of benthic foraminifers
occurred. Apparently, interglacials are not represented by (fig. 4) and rare planktic foraminifers that we interpret as
distinctive foraminifer-rich beds below unit K as they are in dissolved assemblages. Unit I of core 3 yielded only a few
units M through K. However, the sparse planktic and agglutinated specimens. The short interval of unit I identi
calcareous benthic foraminifers at 425 cm and 440 cm in fied above an erosional contact in core 7 is barren of
12 QUATERNARY STRATIGRAPHY AND PALEOCEANOGRAPHY, CANADA BASIN, WESTERN ARCTIC OCEAN

foraminifers. The pattern of preservation of foraminifers in cene sediments of the Arctic but is common in low-oxygen
Northwind Ridge cores documents a general increase in environments and areas of sediments with high organic
depth of the CCD of the Canada Basin with each successive matter content in the North Atlantic (Burgess and Schnitker,
(younger) interglacial stage through the late Quaternary. 1990). The benthic foraminifer assemblages in the upper
The changes in carbonate preservation seen in Northwind part of unit L and in unit M suggest that deepwater
Ridge cores are in agreement with previous studies that conditions in the area of Northwind Ridge were very similar
show that the CCD in the central Arctic has progressively to modern conditions during the last few interglacial stages.
deepened during the Pleistocene (Morris and Clark, 1986). However, lower oxygen conditions prevailed during M3
Information in tables 2 and 3 and figures 6 and 7 on and L1 (probably stages 7 and 9), perhaps reflecting
benthic foraminifer assemblages in cores 5 and 3 was increased productivity and thus greater supply of organic
obtained by quantitative and qualitative methods using material to the sea floor.
several size fractions and cannot be used for detailed A distinct change occurs between L1 and L2. Except
interpretations. However, the data (figs. 6 and 7) reveal for one sample in unit K of core 5, Oridorsalis, which is
several broad features that reflect changing deepwater significant in assemblages in the upper parts of all cores, is
characteristics. Assemblages from M1, M2, M3, and L1 of absent below L1 (figs. 6 and 7). Cassidulina teretis contin
cores 5 and 3 are characterized by Oridorsalis and Stetsonia ues to be common in the assemblages from L2 and L3 of
horvathi. Stetsonia is a very small form that is not found in core 5; Bolivina arctica is an important component of some
the P 150-plm size fraction, and it is well represented only assemblages. Core 3 assemblages from the lower part of
in samples in tables 2 and 3 that were tabulated at >63 p.m. unit L are slightly different from the core 5 assemblages. In
Inspection of the 3150-plm fraction, however, shows that core 3, the L2 assemblages contain abundant Bolivina
Stetsonia is abundant in most samples that contain common arctica and Stetsonia with Fontbotia wuellerstorfi as an
to abundant foraminifers. Our general observation is that, important accessory. L3 is represented by two samples in
where present, Stetsonia horvathi is the dominant taxon in core 3 (table 3, 396.5 and 401.7 cm). The upper sample
the “150-plm size fraction. (fig. 7) has abundant Stetsonia and Epistominella exigua,
Differences between benthic foraminifer assemblages whereas the lower sample yields a sparse assemblage
in the upper part of unit L and in unit M of cores 5 and 3 are dominated by Bolivina arctica. Thus, Bolivina arctica
explained by the depth distribution of modern assemblages appears to have taken the place of Oridorsalis in L2 and L3
in Holocene Arctic sediments. For example, published assemblages. Epistominella exigua, which is abundant
distributional data (Lagoe, 1977; Scott and Vilks, 1991) and between 2,000 and 3,000 m depth in modern benthic
our data from Northwind Ridge box cores (Scott Ishman assemblages in the Norwegian-Greenland Sea (Belanger
and Kevin Foley, unpub. data, 1993) show that Cassidulina and Streeter, 1980; Mackensen and others, 1985), is essen
teretis has its maximum abundance between 1,400 m and tially absent from L3 assemblages of core 5 (one specimen
about 600 m depth throughout the entire Arctic and in the occurs at the top of L3 in the sample from 226.4 cm). The
Norwegian-Greenland Sea (Belanger and Streeter, 1980; absence of Epistominella exigua from core 5, coupled with
Mackensen and others, 1985). Cassidulina teretis is a the abundance of Epistominella exigua in core 3, may
common and consistent component of calcareous assem reflect the presence of a water-mass boundary between
blages in the upper part of unit L and in unit M of core 5 cores 3 and 5 during L3 .
(1,089 m water depth) but occurs sporadically and in much Cassidulina teretis and Bolivina arctica are consistent
lower abundances in equivalent foraminifer-rich levels of components of benthic assemblages in unit K of core 5 (fig.
core 3 (1,909 m water depth). Similarly, Fontbotia wuel 6). Fontbotia wuellerstorfi occurs sporadically along with
lerstorfi and Eponides tumidulus, which generally are more Pullenia bulloides. Cassidulina reniforme, which occurs in
abundant below 1,500 m in the modern Arctic, are impor a few samples from unit L, is a common element in the
tant components of core 3 assemblages in the upper part of assemblages from the lower part of unit K. A few speci
unit L and in unit M but are poorly represented in equivalent mens of Oridorsalis occur in one sample near the base of
assemblages from core 5. unit K.
Inspection of selected core 9 and core 7 assemblages The assemblages recovered from units J and I in core
from units L and M shows that core 9 assemblages are 5 generally contain Cassidulina reniforme, Cassidulina
similar to core 5 assemblages and that core 7 assemblages teretis, and Bolivina arctica (fig. 6). Pullenia bulloides
are similar to core 3 assemblages. Core 7 assemblages, dominates one sample (table 2, 439.6 cm), and the agglu
however, show distinct signs of dissolution below M2; tinated foraminifer Haplophragmoides obscurus is abun
fragments are much more common than in the other cores, dant in several samples from unit I (fig. 6).
and test surfaces are pitted and etched. Bulimina aculeata Studies of core tops from the Norwegian-Greenland
occurs in M3 in all four cores and in Ll of cores 9, 5, and Sea (Mackensen and others, 1985) indicate that Cassidulina
3 (assemblages found in Ll of core 7 are sparse and poorly reniforme is abundant in assemblages from water depths
preserved). Bulimina aculeata is not reported from Holo between 500 and 1,000 m, which is above and partially
C

"HCEtaphBulimina
and
unitsCisolaBolivina
Otprsiahotrdmiaouugrlnmlsiaoeniplahdniepeaks
Eponides
Stetsonia
Fontbotia fascorteretis
planktic
wauhorvathi
marctica
exigua
eilnSpp.
eirfsetor fi
tumidulus
aculeata
obscurus
reniforme 6.
fFigure
benthic
selected
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Apercent,
5.
core
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from
sin
more
or
30
yielding
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for
plotted
are
opbreuacnmidnmaienfcesr.,s
Arctic
Standard

0-
M1
-
-
-
-- i
T-
50

º
lM2 M3
-
-
-
--
-
I
|
I
H
- .
L1 |
I
l
ll–
—-L2 5
L3
--
-
--
-
-
--
-
Lioxsthrptoalsitmrinataitgerfiaseoph.rny
planktic
location
and
text
See
5.
figure
from
fpeaks
benthic
obtaining
for
methods
eof
abundance

M
i -
-
--
-
L
S.-
-
-
-
-
--

|-
4–
|
I-
K-
- l
|
J1.
-
-
-

co.,
O
c
C.
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C

+T T H+ + + TTTTTTTTIT+ T+TT+ T+T TI+


500

f
F
P-
d
450
|
_*
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-
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-

-
200-
#
l
º
– - - Pºm
-


350

-
-
-
-T=
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| –
400
Tl-
T
I
-
-
-

- - - H TTºm
-
-
- .

-
+
100
_rTºm-
--
- -

Abundance,
percent
in
Tºm
*
-
-
-
-
-

- I-
F-
P-
O
-
-
-
-
-
- - -

5
Core

- -

- --

- - -

- - -

- -

1:
T-
--
-
- - -


150
-
-
-
-
--
-

300

-
-
-
-
-
--

Q)
-
-
-
-
-
-+ 250
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-
-
-
-
-
--
-- 3.C
-
-
-
-
-
-
-
-
-
-
-
T.

lCEiEponides
and
units
OrptStetsonia
ihbosdrtoorreasmltpilhgniiresadlpihuamc exigua
Bolivina
Bulimina
Fontbotia upeaks
fhorvathi
planktic
eolSpp.
rearmstionrfifer
warctica
tumidulus
aculeata
excavatum foraminifers of
benthic
selected
of
percent,
in
Abundance,
7.
Figure
yielding
for
plotted
are
Abundances
3.
core
Ridge
Northwind
from
samples sixopthreolasctmirnamtaietgnrifasoeph.nry
more
or
fL30
planktic
location
eand
text
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5.
figure
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benthic
obtaining
for
methods

Arctic
Standard

0M1
k
* !M2
º - -
-
-
-
M3
S.
--
-
300-L
©
-
-
-
-
- L1
T
I
Tl- L-E-+E-O2
|
L _ºm-T-ºm H
400
—E.
L
a 3
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-
-

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-
-
-
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-
-
|

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co
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HHHHH
TTTTTTTTT
TTTTTTTT

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-
-
-

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Abundance,
percent
in

- -
É
H_*
-
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-
-
- |
- - -

3
Core -

- - - - -

|-
- - - -

- -
100
-
-
-
-T-
Th–
-
|
P
-
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-
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-
-
- - - -

efosrtaimiantiefse.r
abundance

- -

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- - - -

200
-
-
-
-
--

500
-
-
-
-

E- - .9
- -
-
--
- §
-- -----
-
-
CONCLUSIONS 15

overlapping the zone of maximum abundance of Cassid size fraction. Similarly, placing the base of unit M lower in
ulina teretis. However, the factors controlling the distribu our cores (below the pinkish-white bed designated NPWB
tion of these taxa are unclear. The increase in abundance of in figure 2) would not alter the fact that, in contrast to the
agglutinated foraminifers in unit I of core 5 and the sparse CESAR core 14 record, significant occurrences of Font
agglutinated foraminifer assemblages found in unit I of core botia wuellerstorfi occur stratigraphically below the first
3 represent increased dissolution in both cores. significant occurrence of Oridorsalis. In addition, Clark
Clark and others (1990) documented benthic assem and others (1990) reported significant occurrences of Font
blages from unit G in several Arctic cores. They concluded botia wuellerstorfi in assemblages from unit G.
that increased exchange between the Arctic and the Our data on benthic assemblages from Northwind
Norwegian-Greenland Sea through the Fram Strait occurred Ridge indicate that the history of deepwater evolution in the
western Arctic and its communication with the eastern
at the beginning of deposition of unit G and that the
increased exchange caused lowering of the Arctic CCD and Arctic and the world's oceans is complex. Substantial
introduction of several North Atlantic taxa such as Font changes occur in both the abundance and composition of
botia wuellerstorfi (as Planulina wuellerstorfi) and Oridor late Quaternary benthic assemblages from Northwind Ridge
salis (sparse and sporadic as Eponides tener) into the both within cores and between equivalent horizons in cores
Arctic. Clark and others (1990) suggested that there is a from different depths. The changes seen at Northwind
general trend toward increasing diversity of Arctic benthic Ridge by us and at Alpha Ridge by Scott and others (1989)
foraminifer assemblages through the Pleistocene and that cannot be completely explained by simple changes in
connections of the Arctic Ocean with the North Atlantic
variations in assemblages are common but that the controls
through the Norwegian-Greenland Sea. Studies of core tops
and significance of the changes are unknown.
from the Arctic and the Norwegian-Greenland Sea show
Scott and others (1989) conducted a detailed study of
Quaternary benthic foraminifers in three CESAR cores that Oridorsalis is more abundant in deeper waters than
Fontbotia wuellerstorfi, but they have very similar upper
recovered from a relatively narrow depth range of 1,370 to
1,555 m on Alpha Ridge of the central Arctic Ocean (fig. depth limits. In addition, Epistominella exigua, which is
strongly associated with Fontbotia wuellerstorfi in modern
1B). Benthic assemblages in these CESAR cores, of which
samples from the Norwegian-Greenland Sea (Mackensen
one, core 14, extends through the entire Pleistocene, show
and others, 1985), is absent from Holocene and unit M
changes that were interpreted to represent increased
exchange between the Atlantic and the Norwegian sediments from the central and western Arctic (Lagoe,
Greenland Sea. Scott and others (1989) found that the first 1977; Scott and others, 1989; this study).
Additional sampling and study of Pliocene and Pleis
significant (>1 percent) occurrences of North Atlantic
tocene sections from Northwind Ridge and other Arctic
benthic species in CESAR cores are within unit M and that areas that include careful documentation of benthic assem
the first North Atlantic taxon to appear is Oridorsalis
umbonatus in the lower part of unit M, followed sequen blages along depth transects are required to develop a better
tially by Eponides tumidulus and Planulina wuellerstorfi understanding of the four-dimensional history of the Arctic
(=Fontbotia wuellerstorfi in this study). Scott and others Ocean and its influence on global oceanic circulation.
(1989) inferred that the changes they observed in CESAR
benthic assemblages must have predated any similar
changes in the western Arctic. CONCLUSIONS
Aside from the sparse occurrence in unit K of core 5,
the Northwind Ridge benthic data also show a dramatic first 1. Lithostratigraphic units established for the upper Qua
appearance of Oridorsalis, but the first appearance is in the ternary of the central and western Arctic Ocean (Clark
upper part of unit L. In addition, Fontbotia wuellerstorfi and others, 1980) are recognized in sediment cores from
forms a significant percentage of assemblages in units L Northwind Ridge. However, the Northwind Ridge sec
(cores 5 and 3) and K (core 5) (see tables 2 and 3). The tion is expanded and contains one or more coarse,
Alpha Ridge benthic data were tabulated from the P63-plm clast-rich units that are not present in the central Arctic
size fraction, whereas our data are a mixture of quantitative Sequence.
and qualitative data, some of which were based on the 2. Foraminifers are concentrated in discrete intervals
>150-plm size fraction. However, the basic differences within the Brunhes Normal Polarity Zone. Available age
between the sequence of events at Northwind Ridge and that control and stratigraphic distribution of the foraminifer
at Alpha Ridge cannot be explained by differences in rich beds indicate that they represent interglacial condi
methodology. The first appearance of Oridorsalis at North tions.
wind Ridge is very evident in all size fractions, and several 3. Differential preservation of foraminifers within and
of the significant occurrences of Fontbotia wuellerstorfi in between cores from different water depths indicates that
unit K of Northwind Ridge core 5 are based on the total the CCD became progressively deeper during successive
assemblage -63 p.m and quantitative splits of the P63-plm (younger) interglacial stages of the last million years.
16 QUATERNARY STRATIGRAPHY AND PALEOCEANOGRAPHY, CANADA BASIN, WESTERN ARCTIC OCEAN

4. Changes in benthic foraminifer assemblages within and Goldstein, R.H., 1983, Stratigraphy and sedimentology of ice
between cores of the Northwind Ridge depth transect rafted and turbidite sediment, Canada Basin, Arctic Ocean, in
indicate that the character and structure of deep waters in Molnia, B.F., ed., Glacial-marine sedimentation: New York,
the western Arctic evolved substantially during the late Plenum Press, p. 367–400.
Quaternary. The most striking change occurs at an Hebbeln, Dierk, and Wefer, Gerold, 1991, Effects of ice coverage
and ice-rafted material on sedimentation in the Fram Strait:
interglacial stage we correlate with oxygen-isotope stage
9. At this change, Oridorsalis becomes a dominant Nature (London), v. 350, no. 6317, p. 409–411.
component of benthic assemblages that were deposited Herman, Yvonne, 1983, Baffin Bay; Present-day analog of the
central Arctic during late Pliocene to mid-Pleistocene time:
between 3,513 and 945 m depth.
Geology, v. 11, no. 6, p. 356–359.
Herman, Yvonne, Osmond, J.K., and Somayajulu, B.L.K., 1989,
Late Neogene Arctic paleoceanography; Micropaleontology,
REFERENCES CITED stable isotopes, and chronology, in Herman, Yvonne, ed.,
The Arctic seas: New York, Van Nostrand Reinhold, p.
Aksu, A.E., 1985, Planktonic foraminiferal and oxygen isotopic 581—655.
stratigraphy of CESAR cores 102 and 103: Preliminary Hunkins, Kenneth, and Kutschale, Henry, 1967, Quaternary
results, in Jackson, H.R., Mudie, P.J., and Blasco, S.M., sedimentation in the Arctic Ocean, in Progress in oceanog
eds., Initial geological report on CESAR —The Canadian raphy, v.4, The Quaternary history of the ocean basins: New
Expedition to Study the Alpha Ridge, Arctic Ocean: Canada York, Pergamon Press, p. 89–93.
Geological Survey Paper 84–22, p. 115–124. Imbrie, John, Hays, J.D., Martinson, D.G., McIntyre, A., Mix,
Belanger, P.E., and Streeter, S.S., 1980, Distribution and ecology A.C., Morley, J.J., Pisias, N.G., Prell, W.L., and Shackle
of benthic Foraminifera in the Norwegian-Greenland Sea: ton, N.J., 1984, The orbital theory of Pleistocene climate:
Marine Micropaleontology, v. 5, no. 4, p. 401–428. Support from a revised chronology of the marine à"O record,
Berger, W.H., 1973, Deep-sea carbonates; Pleistocene dissolution in Berger, A., ed., Milankovitch and climate—Understand
cycles: Journal of Foraminiferal Research, v. 3, no. 4, p. ing the response to astronomical forcing: Boston, D. Reidel,
187–195. p. 269–305.
1979, Preservation of Foraminifera: Society of Economic Kellogg, T.B., 1977, Paleoclimatology and paleo-oceanography
Paleontologists and Mineralogists Short Course [Notes] No. of the Norwegian and Greenland Seas; The last 450,000
6, p. 105–155. years: Marine Micropaleontology, v. 2, no. 3, p. 235–249.
Boyd, R.F., Clark, D.L., Jones, Glenn, Ruddiman, W.F., McIn Kipp, N.G., 1976, New transfer function for estimating past sea
tyre, A., and Pisias, N.G., 1984, Central Arctic Ocean surface conditions from sea-bed distribution of planktonic
response to Pleistocene Earth-orbital variations: Quaternary foraminiferal assemblages in the North Atlantic, in Cline,
Research, v. 22, no. 1, p. 121–128. R.M., and Hays, J.D., eds., Investigation of late Quaternary
Burgess, M.V., and Schnitker, Detmar, 1990, Morphometry of paleoceanography and paleoclimatology: Geological Society
Bulimina aculeata Orbigny and Bulimina marginata Orbigny: of America Memoir 145, p. 3–41.
Journal of Foraminiferal Research, v. 20, no. 1, p. 37–49. Lagoe, M.B., 1977, Recent benthic Foraminifera from the central
Campbell, J.S., and Clark, D.L., 1977, Pleistocene turbidites of Arctic Ocean: Journal of Foraminiferal Research, v. 7, p.
the Canada Abyssal Plain of the Arctic Ocean: Journal of 106–129.
Sedimentary Petrology, v. 47, no. 2, p. 657–670. Mackensen, A., Sejrup, H.P., and Jansen, E., 1985, The distri
Clark, D.L., 1971, Arctic Ocean ice cover and its late Cenozoic bution of living benthic Foraminifera on the continental slope
history: Geological Society of America Bulletin, v. 82, no. and rise off southwest Norway: Marine Micropaleontology,
12, p. 33.13–3324. v. 9, no. 4, p. 275–306.
Clark, D.L., Chern, L.A., Hogler, J.A., Mennicke, C.M., and Malmgren, B.A., 1983, Ranking of dissolution susceptibility of
Atkins, E.D., 1990, Late Neogene climate evolution of the planktonic Foraminifera at high latitudes of South Atlantic
central Arctic Ocean: Marine Geology, v. 93, p. 69–94. Ocean: Marine Micropaleontology, v. 8, no. 3, p. 183–191.
Clark, D.L., Whitman, R.R., Morgan, K.A., and Mackey, S.D., Minicucci, D.A., and Clark, D.L., 1983, A late Cenozoic
1980, Stratigraphy and glacial-marine sediments of the stratigraphy for glacial-marine sediments of the eastern Alpha
Amerasian Basin, central Arctic Ocean: Geological Society Cordillera, central Arctic Ocean, in Molnia, B.F., ed.,
of America Special Paper 181, 57 p. Glacial-marine sedimentation: New York, Plenum Press, p.
Darby, D.A., Naidu, A.S., Mowatt, T.C., and Jones, G., 1989, 331–365.
Sediment composition and sedimentary processes in the Morris, T.H., and Clark, D.L., 1986, Pleistocene calcite lysocline
Arctic Ocean, in Herman, Yvonne, ed., The Arctic seas: and paleocurrents of the central Arctic Ocean and their
New York, Von Nostrand Reinhold, p. 657–720. paleoclimatic significance: Paleoceanography, v. 1, no. 2, p.
Foley, K.M., and Poore, R.Z., 1991, Planktic foraminifer census 181–195.
data from Northwind Ridge core 5, Arctic Ocean: U.S. Phillips, R.L., Grantz, Arthur, Mullen, M.W., Rieck, H.J.,
Geological Survey Open-File Report 91–346, 6 p. McLaughlin, M.W., and Selkirk, T.L., 1992, Summary of
1993, Planktic foraminifer census data from Northwind lithostratigraphy and stratigraphic correlations in piston cores
Ridge cores PI-88–AR P3, PI-88—AR P7 and PI-88–AR P9, from Northwind Ridge, Arctic Ocean, from USCGC Polar
Arctic Ocean: U.S. Geological Survey Open-File Report Star, 1988: U.S. Geological Survey Open-File Report
93–218, 11 p. 92–426, 110 p.
REFERENCES CITED 17

Poore, R.Z., Phillips, R.L., and Rieck, H.J., 1993, Paleoclimate Shackleton, N.J., Berger, A., and Peltier, W.R., 1990, An
record for Northwind Ridge, western Arctic Ocean: Paleo- alternative astronomical calibration of the lower Pleistocene
ceanography, v. 8, p. 149-159. . timescale based on ODP Site 677, in The late Cenozoic ice
Scott, D.B., Mudie, P.J., Baki, V., MacKinnon, K.D., and Cole, age, Transactions of the Royal Society of Edinburgh: Earth
F.E., 1989, Biostratigraphy and late Cenozoic paleoceanog- Sciences, v. 81, no. 4, p. 251–261.
raphy of the Arctic Ocean; Foraminiferal, lithostratigraphic,
Untersteiner, Norbert, 1990, Structure and dynamics of the Arctic
and isotopic evidence: Geological Society of America Bulle
Ocean ice cover, in Grantz, Arthur, Johnson, L., and
tin, v. 101, no. 2, p. 260–277.
Scott, D.B., and Vilks, Gustav, 1991, Benthic Foraminifera in the Sweeney, J.F., eds., The Arctic Ocean region, v. L of The
surface sediments of the deep-sea Arctic Ocean: Journal of geology of North America: Boulder, Colo., Geological Soci
Foraminiferal Research, v. 21, no. 1, p. 20–38. ety of America, p. 37–51.

Tables 2–3 begin on p. 18


o,

Percentages abundance
included.
are
samples
two
or
one
only
abundances
low
occurred
that
Taxa
row.
bottom
the
is
sample
each
in
specimens
number
[Total
rounding
of
because
100
to
add
not
may edoltehrinoamstirfaintacgitrafpieohirnc
because
ior
for
text
See
taxa.
lrare
fof
Planktic
1980).
others,
(Clark
I–M
units
shown
are
L3
and
L2,
L1,
M3,
M2,
M1,
peaks 0.
0
O
1eBluelgiam.nitnselima wuel erstorfi
Fontbotia
.
O
0
2 0obscurus.
0
OHaploph.ragmoides
. 0
O
. O
bulloides
Pullenia 7arctica
0
2Quinqu5eloculina
. 0Stain.forthia
1
O
0.
concava. Triloculina
.
frigida.
6
3
0
O
l
1

1excavatum
0
OCrib00roelphidium
. sGluobogca.lsoibduolsina
O
00.
|
foraminifer
Planktic
peak
M2
M1

Epistominel a
.
exigua.
0
O

Cas idulina
.
laevigata
0
O
2 .
tumidulus
Eponides
1
O
0

Cas i.dulina
.
reniforme.
0O
00

1
0
O
. 0
arctica.
Bolivina O
0
0.
.
aculeata
Bulimina
.

and
6
figures
6]
figure
in
plotted
are
specimens
more
or
30
yielding
samples
from
taxa
benthic
common
for
data
Abundance
7. |
Lithostratigraphic
unit
M
Unit

Aobruanmdinainfcer,s
2.
fTable
benthic
selected
of
percent,
5.
core
Ridge
Northwind
from
samples
in

3spe30c6i.mens.
of
number
Total
.
30.
386
0
10
527
690
0
425
1
O

247
4
04teretis.
O
58
Cas i0dulina
. 80
28
O
.
horvathi
Stetsonia
1

0
O
Valvulineria
.
arctica.

(cm)
depth
1Sample
9
6
3
39
35
32
29.2
26
02
21
17.1
13
44
40

5Orid5
.
spp..
2100
0
39
449
7 o16rsalis
: S

.
tumidulus
Eponides
O
0
0
I. Stain.forthia
.
0concava
0
I3
O

fM2
Planktic
peak
oraminifer
M3

eB7
.
O
0luelgiamnitnselima 1V4
.
arctica.
0
Ol
11
alvul7ineria

Epistominel a
.
exigua.
O
0
6 Haploph.ragmoides
.
0obscurus.
0
1O

T1
.
frigida.
0
O
1
2
4
5rilo5culina

w6
Fontbotia
.
0 uel e.rstorfi
02
1
5
O 21
.
horvathi
Stetsonia
6
O
0 3

f–oCroanmtiniufedr.s
benthic
selected
of
percent,
Abundance,
2.
5Table
core
Ridge
Northwind
from
samples
in

Lithostratigraphic
unit
M
Unit
.
aculeata
Bulimina
O
0
56
6
5
1
50

.
arctica.
Bolivina
O
00 . C5
.
laevigata
3
O
0as i3dulina 2
C0as i.dulina
.
reniforme
0
O

0
0
O
0.
1C11
.
4teretis.
30
14
32
33
21
23
as id. C0ribroe.lphidium
.
excavatum
04ulina 0
O

QO
.
arctica
8
0uinqueloculina

(cm)
depth
Sample
65
81
59
54.5
50.8
47.6
79
78
75.4
73
69
89
87
82
94.4

0sGluobogca.lsoibduolsina
.
0
s454
of
number
Total
9.
pec29i3mens.
318
19
413
0
333
467
174
44
34

00.
0.
bulloides
Pullenia
.
O59
.
spp..
5
0rido.4rsalis
33
18
0
66
64
50
24
17
22
46
S

rCO
.
0aesnif.dourlmien.a
0.
01 Episto.minel a
.
exigua.
0
3
O
0 H0
.
obscurus.
0aploph.ragmoides
O .
bulloides
Pullenia
0
O S0
.
concava.
tainforthia
2
O
1 Trilo.5culina
.
frigida.
2
1
04
3
O
0 s10
of
number
Total
.peci36mens.
670
48
78
2
87
62
2
15
1265
O
064

eB1
.
0
Oluelgiamnitnselima C50
1
.
teretis.
88
69
49
25
31
5
9
0
O
30
53
74
as id4
5ulina VO
.
arctica.
00alvuli.neria
1

|
Lithostratigraphic
unit
M
L
Unit O
.
horvathi
Stetsonia
3
5
0

O
.
arctica.
Bolivina
2
1
0 Cas idulina
.
laevigata
O
0

wuel e0rstorfi
Fontbotia
.
O
1
0

QO
.
arctica
2
0uinqueloculina

f–oCroanmtiniufedr.s
benthic
selected
of
percent,
Abundance,
2.
5Table
core
Ridge
Northwind
from
samples
in
38
.
aculeata
Bulimina
1
3
19
42
4
O
57
0
50
33
13 1
5 sGluobogcalsoibduolsina
.
O
0 O22
.
spp..
3
2
3 rido813rsalis
127
4
O
0
10
13

tO
Eponides
.
01umid3ulus.

foraminiſer
Planktic
Ms
peak

(cm)
depth
Sample
140
136
130.2
127.2
124
121.4
120
118
113.4
110
107
104
103
101
98

C0ribroe.lphidium
.
excavatum
0
: te

187.5
::: : 7

185 : ::: 67
3
3

183 : ::: :
7 7

L2
179 : :::
1 5
540

177 : ::: 3
3

174.4
3::::

f–oCroanmtiniufedr.s
benthic
selected
of
percent,
Abundance,
2.
5Table
core
Ridge
Northwind
from
samples
in :;: :
Lithostratigraphic
unit
L
Unit
::::
0.::::

0::::

::: 0:

::: :
0 0

::: 0:

0::::

0::::

foraminifer
Planktic
peak
(cm)
depth
Sample
167
163.2
161
160
155.3
151
146
141.2
171

sGluobogcalsoibduolsina obscurus.
. Haplophragmoides
.
Cribroelphidium
.
excavatum specimens.
of
number
Total
.
eBluelgiamnitnselima
.
wuel erstorfi
Fontbotia
. Quinqueloculina
.
arctica
Cas idulina .
.
laevigata rCaesnifdourlmien.a
Epistominel a Eponides
.
exigua. Stainforthia
.
concava
Cas idulina
.
teretis.
.
tumidulus Valvulineria
.
arctica.
.
aculeata
Bulimina .
bulloides
Pullenia . frigida.
horvathi
Stetsonia Triloculina
.
.
arctica.
Bolivina Oridorsalis
.
spp..
§

0.
.
bulloides
Pullenia
O
0 S1
.
concava.
O
|
0tainforthia

Episto3minel a
.
exigua.
3
O
0 Triloculina
.
frigida.
0
O

Cas i.dulina
.
reniforme.
02
0
O1 0
O
Valvul.ineria
.
0arctica.
2

|
foraminifer
Planktic
peak
L2
L3
sG0luobogcal.soibduolsina
.
O
0

0
O
.5
arctica.
1Bolivina eBluelgiamnitnselima
O
0.
2
4 Haplophragmoides
.
obscurus.
O
2
0
3 0
.
horvathi
Stetsonia
1O 50

f–oCroanmtiniufedr.s
benthic
selected
of
percent,
Abundance,
2.
5Table
core
Ridge
Northwind
from
samples
in .
tumidulus
0Eponides
O
0 . 0 wO
Fontbotia
.
3uel e3rstorfi

Lithostratigraphic
unit
L
Unit

0Bulimina
O
0.
aculeata
. C7as 00
.
1teretis.
O
0.
80
0
100
82
63
33 id7ulina s11
of
number
Total
.
0
O1p521
17
3
0
5
2
1 e954c.i4mens.

Ori00dorsalis
.
1spp..
0
O

Cribroe7.lphidium
.
excavatum
O
6
0
33
0

Quinqueloculina
.
arctica
0
O

(cm)
depth
Sample
195.5
192
190
211
208.5
239
236
232.5
230
205
201.5
226.4
221
217
244

Cas i.dulina
.
laevigata
0
0
290 0::
: §

7
28
::: 0:

0::286

283 0::

279 0::

274 2::

270 O::
f–oCroanmtiniufedr.s
benthic
selected
of
percent,
Abundance,
2.
5Table
core
Ridge
Northwind
from
samples
in

LiUnit
unit
thostratigraphic
L 267 O::::

2
26

7 0::
25

255 0::::

253 0::

251 ::
0::::

0::::

foraminifer
Planktic
peak
(cm)
depth
Sample
247.3
247

sGluobogcalsoibduolsina obscurus.
. Haplophragmoides
.
Cribroelphidium
.
excavatum specimens.
of
number
Total
.
eBluelgianmtinseilma.
.
wuel erstorfi
Fontbotia
. Quinqueloculina
.
arctica
Cas idulina .
.
laevigata rCaesnifdourlmien.a
Epistominel a Eponides
.
exigua. Stainforthia
.
concava.
Cas idulina
.
teretis. .
tumidulus Valvulineria
.
arctica.
.
aculeata
Bulimina .
bulloides
Pullenia .
horvathi
Stetsonia Triloculina
.
frigida.
.
arctica.
Bolivina Oridorsalis
.
spp..
§

O
0 . O
0Bulimina
.
aculeata eBluelgiamnitnselima O
1
0. 0laevigata
0Cas .idulina
. 0spp...
O
0Orid.orsalis
. Triloculina
.
frigida.
O
0
0

Quinqueloculina
.
arctica
O
0

2
O
0
10
42
.
horvathi
Stetsonia
2

1.
0
OsGluobogc7alsoibduolsina O
5
9
0
1arctica.
8Valvul2ineria
.

O
0
20excavatum
11
0Cribr7o.elphidium
.
Litnit
unit
hostratigraphic
UUnit
KL

0
3
O
8
0
.
bulloides
Pullenia

A–obCruoanmdtianinufcedr,.s
2.
fTable
benthic
selected
of
5percent,
core
Ridge
Northwind
from
samples
in

3.
tumidulus
Eponides
0O
0 .

56
64
21
100
186
0
50
O
58
Cas i2dulina
.
teretis. 1obscurus.
0
1
8
OHaplop4hragmoides
.

32
32
16
0
80O
.
arctica.
Bolivina
4
3.5 2
70.
Osp053
of
number
Total
2649
1154
394
12
57
20
5 e97c.imens.

foraminifer
Planktic
peak
(cm)
depth
Sample
356
352
48.
341.6
339
335
332
327
317
3.11
308
304
302
297
292

0
10.
wuel 2.erstorfi
Fontbotia
20.
50
2rCaesni6.fdourlmien.a
32
01
5
0
27
17.
0exigua.
0
3
0Epist.ominel a
. Stai.nforthia
.
concava.
08
51
0
2
: S.

eBluelgiamnitnselima
O
0.
2 0.
O
0w1uel .e4rstorfi
Fontbotia Trilo.culina
.
0frigida.
O
0

Stainforthia
.
concava
2
0
O
8
1
|

0.
0
Otumi.5dulus.
2Eponides
KLithnit
unit
J
UUnit
I ostratigraphic
nit

0
24
O
0
1Bolivina
5
1.
arctica.
5.
3
7 0 O
.
aculeata
Bulimina Cribroe3lphidium
.
excavatum
O
0
3 1Stetsonia
4
05
0
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73
50
82
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11027
0
1
515
11
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364
389
383
414
420
428.8
375.5
372
404
403.2
399
395
392
415.4
424

14
2
0
67
7
11
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Percentages abundance
included.
are
samples
two
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only
abundances
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that
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benthic
common
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Abundance
7. |
Lithostratigraphic
unit
M
Unit
:::::46
2
5

5
0::

Aobruanmdinainfcer,s
fTable
benthic
selected
of
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3.
core
Ridge
Northwind
from
samples
in
3

3
3
:::::
0:::::9.

:24 :::: 7
4

19
::::: 5

14 :: : : i
O i::
7 : :i 8

4
:::
Mpeak
|foram.
Planktic
inifer
: :: i: 1 8

(cm)
depth
Sample
1

sTurbgolcbihgearmin fiornmias
.

Globocas idulina
.
subglobosa Haplophragmoides
.
obscurus.
Cribroelphidium
.
excavatum aQukinqeurlioacunlain.a
. spe14cimens.
of
number
3Total
.
313
381
743
421
eBluelgiamnitnselima
.
wuel erstorfi
Fontbotia
. Quinqueloculina
.
arctica
Cas idulina
.
reniforme .
gordialis
Glomospira
Epistominel a
.
exigua. Stainforthia
.
concava.
Cas idulina
.
teretis. Cribrostomoides
.
sp. .
tumidulus
Eponides .
wiesneri
Textularia Valvulineria
.
arctica.
.
aculeata
Bulimina
-
.
horvathi
Stetsonia Triloculina
.
frigida.
.
arctica.
Bolivina .
frigida.
Buccella Cyclam ina
.
sp... Oridorsalis
.
spp..
.
sp.
Reophax
&

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.
sp.
O
0 sG0
.l7
O1u1
obogcals0oibduolsina
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.
O
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16
1

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00ibroel.phidium

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arctica.
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.
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0pistominel a w0
Fontbotia
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O
4
3 el e0rstorfi
1 G0
.
gordialis
lomospira
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2
1

.
sp.
Reophax
0
O1l7 concava.
S0
. tainf.orthia
01
3O

f–oCroanmtiniufedr.s
benthic
selected
of
percent,
Abundance,
3.
3Table
core
Ridge
Northwind
from
samples
in |
fM2
Planktic
peak
oraminifer
.
frigida.
Buccella
0
O .
horvathi
Stetsonia
0
134
47
3
3
6
16
O
4
21
9
11 469 V3
.
arctica.
0O
1
18
a4
2lvuli9
neria

LUnit
unit
ithostratigraphic
M
0sTu0
.
0
O rbgolcbih.gearmin fiornmias

HO
.
obscurus.
0a1
0 ploph.ragmoides

eBO
.
0l8
7
4
1
3uelgiamnit9nselima

t0
Eponides
. u1mid950ulus.
616
O9
7
21
2
6
12
5 T27
.
frigida.
O
03rilo2c1ulina
11
5

97
93.5
(cm)
depth
Sample
107.5
102
143
139.5
136
132.5
130.5
125.5
118
114
174.5
165.5
156.5
152
147.5

s0
of
number
Total
. pe16
cimens.
11591
319
283
351
355
53
29
14
314
332
290
307
300
Q3
.
arctica
4u0
2
0
7
6 inque1loculina

O22
.
spp..
rido0rsalis
172
0
31
21
66
23
25
12
69
84
42
34
28
i §

Cribroelphidium 0
.
excavatum
O
0
2
1 1Cribr00
O.
sp. ostomoides Glomo.spira
.
gordialis
O
0
0
1 sTurbgolcbihgearmin fiornmias
.
O
0

Cyclam ina
.
sp.
O
0 aQukinqeu.rlioacunlain.a
.
0
O
0 O
0
.
wiesneri
Textularia

Mpeak
|fora.
Planktic
minifer

.
tumidulus
Eponides
0
O Haploph.ragmoides
.
obscurus.
0
0
O

3
O
0
2
.
arctica.
Bolivina
1 Cas idulina
.
reniforme.
0
O w0
Fontbotia
.0uel e.2rstorfi
O
1
1 O
0
8
Valvulineria
4
.
arctica.
2
1

O
0
1
Cas idulina
.
teretis. Stainforthia
.
concava.
0
O
1

.
sp.
Reophax
O
0
f–oCroanmtiniufedr.s
benthic
selected
of
percent,
Abundance,
3.
3Table
core
Ridge
Northwind
from
samples
in

Lithostratigraphic
unit
M
Unit Trilo.culina
.
frigida.
0
5
0
O
1
2
7

sGluobogcalsoibduolsina
.
O
0
1
3

s11
of
number
1Total
.
0
0
2
Ope31c.i1mens.
237
337
354
2107
313
345

3
0
O
43
2
53
4
.
horvathi
1Stetsonia
1 7
0

eBluelgiamn5itnselima
O
0.
1
1

O
1
0
.
aculeata
Bulimina

196.
190.5
184.5
179.5
(cm)
depth
Sample
280.5
278.5
275
267
261
253
246.5
238.5
231.5
223
216.5
209.5
202.

Quinqu.eloculina
.
0arctica
0
Epistominel a
.
exigua.
0

.
frigida.
1Buccella
021 1 Ori00
.
1spp..
0
37
766
94
44
40
79do1rsalis
§

569.5
O :: º
:: :
7

541.5
515.5
493.8
0 5 6 0.
4 3 |
f–oCroanmtiniufedr.s
benthic
selected
of
percent,
Abundance,
3.
3Table
core
Ridge
Northwind
from
samples
in

LI
unit
JUithonit
Unit stratigraphic : :::
0
3
3 0
6
8

492
i:::
O

O:::::
2 0
8
7
1

: ::0 : 3
9
4
1

28:i:
1

0 O:

0 0:::::

4i:ii:

O:::::

0:::

foraminifer
Planktic
peak
482
469
465.5
461.5
454.5
449.5
445.5
(cm)
depth
Sample

sTurbgolcbihgearmin fiornmias
.

sGluobogcalsoibduolsina
. Haplophragmoides
.
obscurus.
Cribroelphidium
.
excavatum aQukinqeurlioacunlain.a
. specimens.
of
number
Total
.
eBluelgiamnitnselima
.
wuel erstorfi
Fontbotia
. Quinqueloculina
.
arctica
rCaesnifdourlmien.a
. Glomospira
.
gordialis
Epistominel a
.
exigua. tumidulus.
Eponides
. Stainforthia
.
concava. .
wiesneri
Textularia Valvulineria
.
arctica.
.
aculeata
Bulimina Cas idulina
.
teretis. Cribrostomoides
.
sp. .
horvathi
Stetsonia Triloculina
.
frigida.
. Buccella
arctica.
Bolivina .
frigida. Cyclam ina
.
sp... Oridorsalis
.
spp..
.
sp.
Reophax
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