Professional Documents
Culture Documents
World of Bats
One of the greatest mysteries surrounding bats was how they found their way around and located their food in the
dark. It was only relatively recently discovered that many species of bat are able to orientate in the dark by emitting
very short, but loud, pulses of high pitched sound called ‘ultrasound’. This behaviour – echolocation – works in a
similar way to sonar on submarines. Bats listen to the echoes that return from their calls to construct images of their
surroundings. In order to ‘see’ where they are going, bats can emit up to 15 – 20 calls per second at a volume that can
be louder than the sound heard when standing next to a jet aircraft taking off.
The development of electronic devices (‘bat detectors’) that convert ultrasound into sounds that the human ear can
detect has been a major breakthrough in making it possible to listen to bats. Due to the different environments bats
feed in, and the variety of food they eat, each species has developed echolocation calls that are specialised to their
needs. Listening to these calls is fascinating because it provides an opportunity to identify many different bat species
and to get insight into the behaviour of these otherwise very elusive animals.
All of the 17 species of bat found in the UK use ultrasonic echolocation calls to orientate, for communication, and of-
ten for finding food. Using a bat detector at night is an ideal way to discover what bats are flying in your garden, local
woods, or even around streetlights in towns and villages.
By Jon Flanders
Jon, who studied at the University of Bristol, is a member of ‘The
Bat Ecology and Bioacoustics Lab’, an organisation conducting re-
search on the ecology, behaviour and conservation biology of bats.
The 3 types of detector
How they work, their advantages and disadvantages, and their usage
Heterodyne Bat Detectors
How do they work?
Heterodyne detectors combine the incoming bat call with a constant signal generated within the detector so that the
sum and different frequencies are generated. For instance a bat call at 45 kHz (end frequency for a Common Pip-
istrelle) and internal frequency of 43 kHz produces output frequencies of 2 kHz and 88 kHz. The 88 kHz frequency is
inaudible and is filtered out and the 2 kHz frequency is fed to a loudspeaker or headphones.
Advantages Disadvantages
+ The least expensive and best for beginners. - No sound analysis possible.
+ Exaggerates frequency changes of bat calls. - Narrow frequency band means species outside range can
+ Allows sound recording of calls for playback. be missed.
+ Allows real time analysis of calls. - Tuning is continually required.
+ Immediately determine approximate frequency of the - Louder bats can bias population estimates as they are
ultrasound. most frequently heard by heterodyne detectors.
+ With expertise they allow immediate identification.
+ Easy to recognise Doppler shift (bats in flight).
+ Able to detect bats at long distances.
The user estimates the species likely to be present and tunes to the appropriate frequency, once call is heard the frequency is
Usage adjusted until the clearest sound is heard (much like tuning in a radio!). Heterodyne detectors are recommended for non-profes-
sional surveys, or in professional surveys when used with other detector types.
Advantages Disadvantages
+ Requires no tuning when in FD mode. - Frequency changes in calls not exaggerated (unlike het-
+ Broadband recording means that all frequencies are erodyne), which reduces ability to identify species.
detected at once. - All calls are heard at same time, which can make it difficult
+ Records continually so nothing can be missed. to distinguish species.
+ Allows for more calls to be recorded within a given - Picks up all ultrasound signals, can ‘clutter’ calls.
time period than TE - reduces missed calls and false - Sound analysis made using these recordings not as good
identification. as those made with TE detectors.
+ Recordings can be used for sound analysis. - Louder bats can bias population estimates as they are the
+ Preserves rhythm of calls, which can be helpful in aid- most frequently heard.
ing identification.
Dual FD/heterodyne detectors are useful for professional consultancy surveys and transects, especially as many detectors include
Usage a function to record voice notes, and all calls missed using the heterodyne function can be seen and measured on a sound analysis
later. These detectors are able to provide identification to a standard that is adequate for determining the quantity of a species.
Advantages Disadvantages
+ Broadband recording means that all frequencies are - Doesn’t provide real-time/continual recording, which
detected at once. means less accuracy and some bats may be missed.
+ Unlike other detectors, it uses original signal, preserv- - Whilst a recording is being played back nothing is being
ing the whole waveform, making analysis easier and recorded, so bat calls are sampled in small segments
more accurate. - Automatic recording (possible to use manual) initiatied by
+ Recordings can be used for the most accurate sound false triggers (non-bat ultrasound) can be an annoyance.
analysis of any type of detector. - As with other types of detectors: louder bats can bias
population estimates as they are the most frequently heard.
The user has several options with this detector. Firstly, they can use the heterodyne function as normal and record the TE output
Usage for sound analysis later. Secondly, they can listen directly to TE output. Lastly stereo headphones can be used to simultaneously
monitor both the TE and heterodyne output.
Choosing the right detector
Also available Pettersson Designed to be left in the field for up to two weeks, the D-500X detects the full
D-500X spectrum of ultrasound and records in real time. Thus, it provides much more
for long-term
detailed data than either frequency division or time expansion detectors, and is
monitoring: #178758
particularly good at detecting those ‘hard to detect’ species such as the Brown
Long-eared bat.
* Can be output to tape recorder, mp3 player with input feature etc. *** Built to order.
** Note that it is possible to use rechargeable versions of each type of battery.
Features in-depth
Other factors to consider while choosing your bat detector
Type of Microphone and Frequency Range
The type of microphone used in a detector determines its sensitivity. Electret microphones are
most sensitive at low
frequencies (<30 kHz). Piezo microphones are particularly sensitive around 45 kHz, whilst
capacitance microphones do best at frequencies greater than 80 kHz. All good detectors are
able to scan a frequency range of 20 to 120 kHz, meaning that all models will detect all Eu-
ropean species. Thus, regardless of the microphone, you will be able to detect any species.
The only difference is that if you have a piezo microphone (optimally operating at low frequen-
cies) the distance you can detect a bat operating at high frequencies (e.g. lesser horseshoe)
will be less than if you have a capacitance microphone. Some detectors improve sensitivity by
combining more than one type of microphone, such as the Pettersson D-100 with its combined
electret-piezo microphones.
Recording Calls
You can record bat calls from a bat detector on to most commercially available recording
devices. Ideally the recorder should have a ‘line in’ socket to ensure the sound quality from
the detector is preserved. If recording from a frequency division or time expansion detector,
you can then use sound analysis software to identify species. The recording is turned into a
sonogram (sound graph) on your computer for subsequent analysis. To transfer recordings
from your recorder to your computer, the computer must have a ‘line in’ socket, which most
modern computers now have (some laptops combine the ‘line in’ and ‘mic’ socket, which is
also fine to use).
Listening to Calls
Most bat detectors have both an inbuilt speaker and socket for headphones. The speaker allows multiple users to listen to
the same detector, such as when on bat walks. Using headphones rather than the speaker provides better sound quality
and allows you to hear the weakest bat signals. Background noise interference is reduced when the speaker is not produc-
ing sound. If using stereo headphones, some bat detectors allow you to hear the heterodyne output in one ear and the
frequency division output in the other.
Bandwidth
When using heterodyne detectors, the bandwidth can be an important consideration for the user. A small bandwidth
(e.g. 4 kHz) makes it easier to judge where a call starts or finishes, which can be useful when targeting a specific
species. Conversely, larger bandwidths increase the likelihood of detecting a species when undertaking a general survey.
For example, a detector with a bandwidth of 10 kHz, when tuned to 50 kHz, will cover 45 - 55 kHz simultaneously. Finally,
remember the size of the bandwidth does not influence the ability to tune in accurately to a particular frequency.
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Latest news from NHBS
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