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ABUNDANCE,
Introduction
Interpretation of palaeoenvironmental changes is an important facet of many disciplines
including archaeology, botany and geography. Each of these disciplines discerns past
changes in different ways. The prevailing conditions during particular palaeoenvironments are archaeologically inferred by several means, including phytoliths, micro and
macro-fauna1 remains, seed material, pollen and charcoal as sources of information
(Brothwell & Higgs, 1963; Shackley, 198 1). Credence is lent to particular interpretations
and conclusions if similar trends are evident from independent, although complementary,
forms of investigation (Deacon & Lancaster, 1988).
The taxonomic identification of charcoal in particular yields data depicting changes in
the floral species composition of the area under scrutiny. Combined with analysis of
anatomical features such as vessel diameter (Deacon & Lancaster, 1988; Scholtz, 1990)
and botanical knowledge of the ecological requirements of extant species (also represented in the charcoal sample) it is possible to predict some details of the climatic
conditions prevailing during the different periods (e.g. Deacon et al., 1983; Prior, 1983;
Deacon et al., 1984; Prior & Price-Williams, 1985; Prior & Tuohy, 1987; Dowson, 1988;
Tusenius, 1989).
Trends of increasing or decreasing abundance of chosen indicator species should be
accorded greater emphasis than erratic changes in the abundance of rarer species. A time
sequence of dates and species composition data should be set up to emphasize differences
in the abundance of species (e.g. Prior, 1983).
Functional interpretation of charcoal data sets rests upon acceptance of the Principle
of least effort (Scholtz, 1986; Tusenius, 1986). It is the validity of the underlying
assumptions that are the focus on this article.
OWits Rural
bDepartment
Africa.
Facility,
P.O. Box 7, Klaserie 1381, South Africa.
of Botany,
University
of Transkei,
P.B. Xl,
Unitra
5100,
South
631
03054403/92/06063
I+07
$08.00/O
0 1992 Academic
Press Limited
632
C. M. SHACKLETON
AND F. PRINS
The PLE assumes that all species were collected as fuelwood in direct proportion to their
abundance in the environment. If, however, particular species were actively avoided or
selected, then the charcoal remains would fail to provide an accurate reflection of the
composition and abundance of the species at that time. Although Prior & Price-Williams
(1985) Scholtz (1986) and Tusenius (1986) amongst others, have questioned the validity
of this assumption it cannot be falsified directly through archaeological excavation.
Several ethnographic and biological studies, however, suggest that the assumption may
indeed be false (Best, 1979; Ford, 1979; Jelenic & van Vegten, 1981; Gandar, 1983).
Unpublished work by Shackleton (in press) shows that the avoidance of several species
CHARCOAL
Fresh
4
Strong
wood
633
ANALYSIS
Dry wood
I
I
I
I
I
1
High
Wood
abundance
per
person
The model proposed (Figure 1) ideally applies to fuelwood collected for domestic use but
it will also apply to those species selected for specialist functions like smithing and the
firing of ceramics. It describes the degree of selectivity for different fuelwood species
relative to the total abundance of wood in the environment. As such, it can be used to
predict when the PLE may or may not be in force. It rests on two conditions. Firstly, if
wood is plentiful then inhabitants will select species that are the most desirable for the
purposes on hand. Initially, selection will be for dry wood of a particular species-it being
preferred to fresh wood that is heavier to carry and takes time to dry before it can be used.
Secondly, if wood is in short supply, fuelwood collectors will probably exploit whatever
wood is available to them. In this instance the PLE will apply. Additional factors such as
the annual burning of the woody vegetation and clearance of large tracts of woody
vegetation for agricultural fields may influence the process. However, these factors are
not considered here and they require further research. For ease of interpretation four
generalized areas are described in the model. Most settlements probably experienced a
sequential transition from one to the next during the course of occupation.
Area 1. This represents the typical situation encountered at a new settlement in a relatively
unexploited area. There is plenty of fuelwood, both dead and live, with a considerable
diversity of species available. Thus, there is maximum wood abundance per person and
hence individuals may select dry wood freely, taking that which is most suited to their
needs. Even though there is maximum selectivity there is very little effort involved in
fuelwood collection since total abundance is high, i.e. inhabitants do not have to go far,
nor search for long, to obtain dry wood of the particular species that they are seeking. In
summary, therefore, area one is characterized by a high dry wood availability, maximum
634
C. M. SHACKLETON
AND F. PRINS
selectivity and low effort. Modern analogues are scarce but this situation may apply in
some nature reserves where controlled exploitation of natural resources by indigenous
peoples is allowed (Brunton, 1980).
Area 2. This area is defined by a medium abundance of dry wood, declining selectivity and
maximum effort. This situation applies near settlements with a declining dry fuelwood
resource. This may be a new site in a previously exploited area. More likely however, it is
the same settlement as in (1) above, after several decades of occupation. The size of the
settlement has increased and the cumulative effects of fuelwood collection over time have
decreased the total availability of dry wood. The degree of selectivity is decreasing since
the abundance of the most sought-after species is also decreasing. Some individuals are
using increasing amounts of previously unfavourable species. Maximum effort is required
at this stage, since it takes time and energy to seek out the few remaining dry branches of
the most preferred species. Situations with declining dry fuelwood resources may even
occur in relatively low population pressure areas where traditional settlement patterns are
maintained such as was observed in tribal areas near Willowvale in the Transkei (Prins,
unpubl.).
Area 3. As time progresses, dry wood of the preferred species is depleted to levels where
the effort required to seek it out is too great (although it is not necessarily exhausted
completely). This stage is characterized by low wood abundance, little selectivity and
minimal effort. In the most extreme situation any wood present is removed, even if its
fuelwood properties are poorly rated. In this situation the PLE would apply. Resettlement
areas in Transkei and Gazankulu, with a relatively high population pressure on the
available natural resources, exhibit this pattern.
Area 4. Demand for dry wood far exceeds supply, even if the least favourable species are
included. Supplementary supplies are obtained through the collection of live wood. Since
this is a new source of supply, many different species are available and the degree of
selectivity will therefore be high. The situation is essentially a repetition of Area 1 in the
model but applied to fresh wood. The model follows the same course for live wood as
described for Area 2 and 3 pertaining to dry wood. A modern analogue would be longestablished areas of resettlement in tribal areas such as were observed in northern
Transkei and the eastern Transvaal Lowveld. However, it should be noted that some
overlap between Areas 3 and 4 is likely to occur as certain highly favoured species could be
removed as green wood, even though dry wood of less desirable species occurs abundantly
near the homestead. This would especially be the case where wood is required for specialist
functions or ceremonies. Nevertheless, the transition from Area 3 to 4 represents both an
anthropological and ecological threshold.
Predictions and implications
CHARCOAL
ANALYSIS
635
latter. Ultimately, whatever is available will be collected. Should inhabitants adopt the
first strategy, species not present in the area may be imported into the charcoal deposits
leading to erroneous interpretation.
Even if (a) is adopted, under conditions of high
fuelwood demands and long occupancy option (b) will ultimately be taken. In this way
PLE may apply during the later occupational phases.
3. Highly preferred species may be selectively reduced with increasing harvesting pressure.
This will come about through either an increase in the number of people in the settlement
and, therefore, a greater fuelwood demand, or continuous removal over long periods.
Thus, the relative abundance of less desirable species will increase. At some threshold
abundance, the amount of effort required to seek out the most favourable species is too
great even though a few sparsely distributed individuals may remain. At this point an
increasing proportion of the less desirable species would be evident in the charcoal record.
Theoretically, the change in focus for particular species from the most desirable to those
of intermediate preference (not actively sought initially, nor actively avoided) would
provide a respite to the few remaining trees of the most preferred species. This would
provide time for them to increase in relative abundance, whilst the other species experienced a decline in relative abundance similar to that which the former previously underwent. Since generation times of woody species are long, these changes would take several
decades. Were that the case, then young fuelwood harvesters would be unable to recognize
the resurgence of the most preferred species, having being taught what were the most
desirable from an impoverished assemblage of species. An example of this is the inability
of young fuelwood collects (younger than 16 years old) to recognize the less abundant fruit
trees (e.g. Annona senegalensis) in the Timbavati communal grazing lands-and yet older
community members easily identified it as auseful fruit-bearing species (Shackleton,
in press). However, at some threshold of decreasing abundance of the species recognized
as desirable, an increasing proportion of the initially most favoured species would again
be collected.
This theoretical projection could result in cyclical changes of increasing and decreasing
abundance of species in both the environment and the charcoal remains. This extrapolation of the model may be extreme. However, if plausible, it is significant in that it
projects changes in species abundance in the absence of climatic changes. Furthermore,
stable limit cycles are not unknown from biological literature (Whittaker,
1975;
McNaughton & Wolf, 1979). If other animal species may induce such cycles why not
humans?
4. If cyclical changes in species abundances exist in the environment (and hence
the charcoal record) brought about through human interference important thrusts in
ethnographic research would be to identify the threshold points of the model. Thus, at
what threshold densities of wood (total and relative species) is there a change in the
situation from one to which the PLE is not applicable to one where it is?
5. Even if the extreme, theoretical extrapolation
(discussed in implication 3) is
unattractive, the model still demonstrates (although to a lesser degree) that changes in
relative frequencies of species in the charcoal record through time may not necessarily be a
result of altered abundance induced through climatic changes. Thus, as argued by Scholtz
(1986) they may have been the result of the selective use of the most favoured species.
For example, data from Deacon et al. (1983,1984) demonstrate changes in abundance
of several species throughout the stratigraphic record. This is interpreted as indicating
climatic change. This may well have been the case. However, the possibility also exists that
these changes were induced or exacerbated by human selection and localized removal of
some species over others, promoting the exploitation of alternative species.
636
Similarly, data presented by Prior (1983) indicate that the frequency of Dichrostachys
cinerea increases throughout the stratigraphic record, with a concomitant decrease in
Sclerocarya
birrea. There is little discussion of these trends, but they could function
Acknowledgements
Thanks are due to the colleague of CS at the Wits Rural Facility for comments and
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Cambridge: Cambridge University Press.
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ANALYSIS
631
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