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Rules of attending the class # max. 15 lately # noisy forbidden # keep HP silent # fit dressing Presence at least 75% Grading: >= 80 A Distribution
66 79,9 56 65,9 46 55,9 <= 45,9 B C D E Mid exam Assignment Course Prac Final exam 25% 20% 30% 25%
Scope of Ecology
Ecology is primarily concerned with those biological (and Biogeochemical) processes that control the functioning of populations, communities, and ecosystems over large spatial (communities to global) and long temporal (days-millennia) scales.
Ecosystem Properties:
Structure: Species diversity: plants, animals, and microbes; Community structure; Foodweb structure; Soil type: structure, texture; Carbon and Nutrient Pools Function: Energy capture (primary productivity, yield); Energy flow; Nutrient cycling; Population regulation; Stability and flexibility; Disturbance regime; Succession.
BIODIVERSITY
What is Biological Diversity or Biodiversity? Biodiversity or biological diversity is defined by the United Nations Convention on Biological Diversity as: "The variability among living organisms from all sources, including, inter alia [among other things], terrestrial, marine and other aquatic ecosystems and the ecological complexes of which they are part: this includes diversity within species, between species and of ecosystems."
Within that definition, there are 3 distinct levels of biodiversity: Species diversity: diversity among species present in different ecosystems. This is the diversity of populations of organisms and species and the way they interact. Genetic diversity: diversity of genes within a species and processes such as mutations, gene exchanges, and genome dynamics that occur at the DNA level and generate evolution. Ecosystem diversity: genetic, species, and ecosystem diversity of a given region. This is the diversity of species interactions and their immediate environment.
Names and word definitions of food chain Producers. Organisms, such as plants, that produce their own food are called autotrophs. The autotrophs, as mentioned before, convert inorganic compounds into organic compounds. They are called producers because all of the species of the ecosystem depend on them. Consumers. All the organisms that can not make their own food (and need producers) are called heterotrophs. In an ecosystem heterotrophs are called consumers because they depend on others. They obtain food by eating other organisms.
Consumers i.e. : Herbivores are those that eat only plants or plant products. Carnivores, on the other hand, are those that eat only other animals. Omnivores are the last type and eat both plants (acting a primary consumers) and meat (acting as secondary or tertiary consumers). Trophic level. The last word that is worth mentioning in this section is trophic level, which corresponds to the different levels or steps in the food chain. In other words, the producers, the consumers, and the decomposers are the main trophic levels.
FOOD WEB
The concept of food chain looks very simple, but in reality it is more complex. How many different animals eat grass? How many different foods does the hawk eat? One doesn't find simple independent food chains in an ecosystem, but many interdependent and complex food chains that look more like a web and are therefore called food webs. A food web that shows the energy transformations in an ecosystem looks like
One way to calculate the energy transfer is by measuring or sizing the energy at one trophic level and then at the next. Calorie is a unit of measure used for energy The energy transfer from one trophic level to the next is about 10%. For example, if there are 10,000 calories at one level, only 1,000 are transferred to the next. This 10% energy and material transfer rule can be illustrate with an ecological pyramid
QIUZ I
1. Apa pengertian Ekologi? 2. Sebutkan struktur dan fungsi ekosistem? 3. Apa pengertian Keanekaragaman hayati? 4. Sebutkan dan jelaskan level kehati? 5. Kenapa kehati itu penting?
BIOGEOKIMIA
Succession
Komunitas yang terdiri dari berbagai populasi bersifat dinamis dalam interaksinya yang berarti dalam ekosistem mengalami perubahan sepanjang masa. Perkembangan ekosistem menuju kedewasaan dan keseimbangan dikenal sebagai suksesi ekologis atau suksesi. Suksesi terjadi sebagai akibat dari modifikasi lingkungan fisik dalam komunitas atau ekosistem. Proses suksesi berakhir dengan sebuah komunitas atau ekosistem klimaks atau telah tercapai keadaan seimbang (homeostatis).
Succession
A directional, cumulative change in the species that occupy a given area, through time. Primary vs secondary Autogenic vs allogenic Progressive vs retrogressive Cyclic vs directional
Primary succession the establishment of plants on land not previously vegetated (volcanic explosion)
Secondary succession The invasion of land that has been previously vegetated (fire, logging or cultivation)
Autogenic succession both the environment and the community change and this metamorphosis is due to the activities of the organism themselves (environmental stress adapted) Allogenic succession Due to major environmental change beyond the control of the indigenous organisms (Env. Change Changes the pattern of vegetation)
Progressive succession lead process that the communities with greater and greater complexity and biomass Retrogressive succession Lead process that the community toward simpler (fewer species)
CROP ECOLOGY
EKOLOGI: ilmu yang mempelajari hubungan timbal balik antara faktor biotik dan abiotik. Scope: Distribution and Abundance EKOLOGI TANAMAN: pengembangan dari ekologi dalam lingkup tanaman (budidaya pertanian).
Goals of Agro-ecosystem Management Provide an adequate income to the farmer Maintain the resource base on which future production depends Produce enough food to meet the demands (of the farm family, local community, region or nation, or globe)
Maintaining the resource base (soils, biodiversity) is the core of most definitions of sustainability. Definitions of Sustainablity by The American Society of Agronomy: "A sustainable agriculture is one that, over the long term, (i) enhances environmental quality and the resource base on which agriculture depends, (ii) provides for human fiber and food needs, (iii) is economically viable, and (iv) enhances the quality of life for farmers and society as a whole."
Environmental conditions to refer to the things, both abiotic and biotic, that influence the rates and efficiencies at which plants capture (or lose) supplies of these resources. Resources to refer to the things plants consume in their growth and reproduction, and
Resources It is axiomatic that crop plants must consume resources to grow and produce a harvestable yield. In most agro-ecosystems, crop productivity is limited by the availability of one or more required resources, most often nutrients, water, and light. The amount of yield achieved by a crop is a function of both the level of limiting resources available to the crop, and the efficiency with which it uses these resources.
Environmental factors
I. CLIMATE Important features of climate include: light temperature humidity precipitation Wind Climate includes both: Resources [light, precipitation (actually, soil water is the resource)] Conditions (e.g., temperature, day length, humidity, wind)
Environmental factors
II. SOILS 1. 2. 3. 4. 5. 6. 7. SOIL CONSTITUENTS MINERAL (INORGANIC) FRACTION SOIL ORGANIC MATTER SOIL STRUCTURE SOIL TYPES SOIL ORGANISMS SOIL pH
Environmental factors
III. RESOURCES Light Carbon dioxide Water Nutrient
CLIMATE
I. LIGHT (Solar Radiation) The seasonal distribution of light is controlled by latitude. [How does the light environment of tropical latitudes differ from that of temperate and boreal latitudes?] Plants (including many crops) show photoperiodic responses to day length, particularly in their phenology.
CLIMATE
I. LIGHT (Solar Radiation) Phenology has been defined as "the sequence of development events during the plant's life cycle as it is determined by environmental conditions" (Hall, 2001); these include flowering, bolting, tuber formation, etc. "Long day" (LD) plants; "short-day" (SD) plants; and "day neutral" (DN).
CLIMATE
II. TEMPERATURE The seasonal and diurnal variation in temperature increase with latitude Temperature also decreases with increasing altitude The rate of temperature change with altitude is called the lapse rate and is about 1 0C 100 m-1 for dry air and about 0.6 0C 100 m-1 for wet air.
CLIMATE
II. TEMPERATURE Most plant processes have an optimum temperature. Respiration increases with increasing temperature. Plant development is mostly controlled by temperature. Plants sense environmental temperature in terms of degree days --the cumulative number of the degrees above a base or threshold temperature. Crop plants exposed to higher than normal temperatures develop at a more rapid rate (for example, flower earlier), which could decrease yield.
CLIMATE
III. PRECIPITATION GO TO ANOTHER SLIDE : RAINFALL N CROPPING SYSTEMS IN INDONESIA
SOILS
I. SOIL CONSTITUENTS Atmosphere Water Mineral (inorganic) materials Soil organic matter (SOM) Soil organisms The atmosphere below ground in the soil difference substantially from that aboveground. The soil atmosphere is higher in CO2 and lower in O2
SOILS
Soil provide an important environment for plants/crops due to: 1. Plants need anchorage, so that there should be adequate soil layer. 2. Plants need water, so that soil should hold adequate water and supply. 3. Plants need oxygen for respiration, so that soil should be able to provide it without any interruption. 4. Plant roots release CO2 during respiration, and soil should be able to regulate the movement of this gas without allowing it to build up to toxic levels
SOILS
5. Plants need nutrients from soils, which are absorbed by roots, so that soils should have some characteristics to supply and retain nutrients. 6. Plants add a lot of dead material (OM) and the soil should have able break them to some form so that they will not interfere with plants and their root systems. 7. Some plants through root exudates add to soil toxic chemicals (allelo-chemicals) and soil should be able to decompose them to avoid root damage.
SOILS
8. During heavy rainy periods, large volumes of water are added with a very high intensities and the soil should be able to handle these volumes without severe soil losses 9. There are toxic gases released when animal and root systems grow in soils and soil should be able to either release these gases to atmosphere or convert to non-toxic form by other reaction
SOILS
10.When both plant and animals live in soil, it should be able to maintain suitable temperatures required by those living beings
SOILS
Therefore Soils is suitable for everything at anytime It is required to treat the soil with the right knowledge of it in order to receive benefits the mankind wants soil always have many associations and interactions among these factors (physical, chemical, physico-chemical and biological factors)
Physical factors
Soil texture Particle size distribution (clay, silt and sand) In general Coarse sand 0.25 2.0 mm Find sand 0.05 0.2 mm Silt 0.002 0.05 mm Clay < 0.002 mm
Physical factors
Bulk density and porosity Both factors related to: 1. Capacity for gas exchange 2. Root growth and penetration 3. Drainage and retain water 4. Infiltration and percolation
Physical factors
Soil structure Composition of pores and soil aggregates Pores consist of : Micro pores (capillary water retained) Macro pores (gas exchange and drainage) Crumb structure best for agriculture 50 % each of micro and macro pores.
Physical factors
Soil water content Saturated condition Field capacity Permanent wilting point Soil temperature Increase root growth and activities Increase microbial population Increase organic matter decomposition Increase seed germination
Chemical factors
Nutrient contents in soil Gas content Chemical reactions
Physico-chemical factors
(good for agriculture) pH (6 7) CEC (Cation exchange capacity) (> 40 mg/100 g soil) EC (electrical conductivity) = water quality parameter (0.4 0.7 m mhos/cm)
Biological factors
Micro and macro both fauna and flora Important activities: Mineralization of organic matter Nitrogen fixation in legumes Micorrhyza promoting P absorption Enzymes activities and nutrient transformation in soils Improve porosity by earthworm (tunneling) Improve root absorption activities
RESOURCES
Light Quantity Full Sunlight: 200-500 Wm-2 or 1000-2000 mol m-2 s-1 (W = J s-1) Cloudy sky: 20-90 Wm-2 or 100-400 mol m-2 s-1 Seasonality: The highest monthly (i.e., growing season) maximum light levels are at higher latitudes. Crop yields in the tropics (compared to temperate zones) are ultimately limited by: incident radiation cloudiness-compare wet season and dry season yields
RESOURCES
Growth and Yield are ultimately related to light interception. At the leaf level: There is a minimum amount of light required for a positive net photosynthesis to occur, called the light compensation point. At the canopy level: Some leaves in a canopy will be shaded by other leaves, some below, and perhaps some below the light compensation point. Rates of canopy photosynthesis are usually proportional to LAI
RESOURCES
At the crop level: Crop growth (and yield) is generally a function of leafarea duration (LAD), the area under a curve of LAI vs. time. LAD is proportional to the total amount of light energy absorbed during the crop's growing season, and thus to yield.
RESOURCES
CO2
The direct (physiological) effects of this increase in atmospheric CO2 are: increased rates of photosynthesis, especially in C3 plants, resulting in higher crop yields. increased water-use efficiency. higher C:N ratios in plant biomass. Higher CO2 concentrations induce partial closing of the stomates, which increases the resistance to the flow of water vapor, reducing transpiration and thus increasing water-use efficiency.
RESOURCES
Higher leaf temperatures (caused by stomatal closure) associated with increased [CO2] can lead to increased leaf turnover rate (higher leaf temperatures and more rapid leaf aging), Decreased specific leaf area, reducing the CO2-fertilization effect.
RESOURCES Soil Water Field capacity is the amount of water held in a saturated soil after all excess water has drained off; the water potential at field capacity is -0.1 to -0.2 MPa. Permanent wilting point is the point at which a (particular) plant can no longer absorb water from the soil, for most plants in most soils the water potential at the permanent wilting point is about -1.5 MPa.
RESOURCES
Available water is the amount of water between field capacity and permanent wilting point. Soil water content is influenced by both soil texture and soil organic matter (SOM). Fine-textured soils have a higher total pore volume, and hence can hold more water. Clay particles hold water more tightly. SOM functions similar to clay particles in affecting soil water-holding capacity and soil water potential.
RESOURCES
Nutrition Macronutrients, those required in rather high amounts by plants, are nitrogen (N), phosphorus (P), potassium (K), calcium (Ca), magnesium (Mg), and sulfur (S). Most fertilizers contain N, P, and/or K. Micronutrients are elements that are also essential for growth but are required in lower amounts; these include iron (Fe), copper (Cu), zinc (Zn), boron (Bo), molybdenum (Mo), manganese (Mn), cobalt (Co), and chlorine (Cl).
Nutrient cycling
Refers to the processes that transfer nutrients to and from plants and the various soil (and atmospheric) pools. These pools can be characterized as: active, inorganic forms and microbial biomassvery rapid turnover; slow, new crop residues and coarse particulate organic matter; and Passive, fine particulate organic matter and humic substances-very slow turnover.
Factors that influence crop yield are of several types and include: Resources not under grower control: light, CO2, water (precipitation), nutrients released by mineralization. Environmental conditions, not under grower control: temperature, wind, seasonality, topography, length of growing season, relative humidity; soil type, soil depth, SOM, soil pH; pest, weed and pathogen populations (in part). Resources under grower control: nutrients (from fertilizer), water (from irrigation).
Factors that influence crop yield are of several types and include: Environmental conditions under partial grower control: pest, weed, and pathogen populations; SOM; soil structure; soil pH. Crop varieties. Management: land preparation, choice of cropping system; choice of cultivars; date of planting; plant population; timing of nutrient input; timing of pest, weed and pathogen control; date of harvest; management of residues. Infrastructural or institutional factors: access to credit, suitable varieties, extension services, inputs, markets.
Interactions Among Species in Agroecosystems This part of the course considers some of the other organisms, in addition to crops and soil organisms, that occur in agroecosystems, particular herbivores (mostly insects) and their predators, and competitors (weeds). Pathogens are discussed only briefly.
Herbivores
Why don't insects (and other herbivores) consume all available plant biomass? That is, Why is the world green?-most likely answers are plant defenses that limit which herbivores can feed on which plants, and predators that keep herbivore populations in check. Groups of herbivores: Vertebrates-birds, mammals Invertebrates-insects, arachnids (mites), mollusks (snails, slugs). Of these groups insects cause the greatest crop losses in most agroecosystems.
Herbivores
Plant Strategies to cope with herbivory: Escape-short life cycle Tolerance--Compensation for tissue loss Defense--protection of tissues Ecological problems associated with insecticide use: 1. Insecticide resistance 2. Pest Resurgence 3. Secondary Pest Outbreaks
Competitors (Weeds)
Characteristics of Weeds High seed production, competitiveness, low attractiveness, seed longevity, seed dormancy, rapid emergence. Most weeds evolved from early successional species; many are crop relatives
Competition/Niche Theory
Two species can occupy the same habitat and not compete if: The species use different resources. This is often true for animals, but seldom true for plants. Resources are sufficient for both. For example, plants in the desert seldom compete for light. The species obtain their resources from different parts of the habitat. I.e., the species have a somewhat different niche with respect to resource acquisition.
Competition/Niche Theory
Many plant ecologists (e.g., David Tilman) maintain that plant species specialize with respect to their ability to capture different resources. This is probably not true, however, for crops and weeds.
Competitors (Weeds)
Weeds reduce crop yield by reducing the supply of resources through competition. Plants use common resources--Light, C02, Water, Nutrients. Plants obtain resources from resource depletion zones, which depend on root and shoot architecture, and on resource mobility. Intensity of competition depends on the degree of overlap of resource depletion zones.
Pathogens Diseases reduce ecological resource use efficiency by reducing resource uptake by various mechanisms: obstructing vascular tissues, damaging roots, restricting root growth, or removing leaf area. Plants possess morphological and chemical defenses against pathogens:
Pathogens
Morphological-- cuticle Chemical-both constitutive and inducible (inducible defenses against pathogens are called phytoalexins) These defenses most effective for aboveground pathogens.
Diversity has been defined as: Richness-number of species Equitability-number and relative abundance Connectance or complexity-usually as foodweb complexity Ecosystem function is usually defined in terms: energy capture (i.e., productivity-yield inagriculture) nutrient cycling population regulation (including food web structure) stability
Crop Rotation
Prior to development of agrichemicals, rotations were the standard practice to control pests and diseases and maintain soil fertility. Development of pesticides and herbicides made continuous monoculture possible. Thus continuous monoculture is a relatively recent agricultural practice.
Crop Rotation
Short rotations vs Long (Extended) Rotations: Short rotation: Usually just 2 years Objective is typically pest control Corn-soybean is the commonest crop system in the US-both crops have a high demand Long (extended) rotations: 3 years or longer Objectives are pest control, maintain soil organic matter, reduce agrichemical inputs Usually includes hay, pasture, or "green manure" to improve soil fertility.
Crop Rotation
Rotation Effect! This term refers generally to the higher yields of most crops when grown in rotation, and more specifically to the yield increases that cannot be compensated for by input substitutions. Most crops produce higher yields in rotation than in continuous cultivation, usually 10-15% higher in maize (Singer & Cox, 1998).
Intercropping
Intercropping involves growing two crops in the same field at the same time. The following are different ways of intercropping, in order of increasing degree of association between crop components: Relay-intercropping-planting a second crop before harvesting the first crop. Continue
Intercropping
Strip-intercropping-growing 2 or more crops in alternating strips. Smith & Carter (1997) found that maize grown in a strip intercrop with alfalfa produced yields 6% higher in 40-ft wide strips, 11% higher in 20-ft wide strips, and 17% higher in 10-ft wide strips. May be due to extra light in border rows of maize.
Continue
Intercropping
Between-row intercropping -growing 2 or more crops in alternating rows. Within-row intercropping -growing 2 or more crops in the same rows. Between-row and within-row intercrops may be either additive or replacement designs.
Intercropping Concepts.
Additive vs. replacement intercrops. In an additive intercrop both species are planted at the same density as in their respective monoculture; in a replacement intercrop a row of one crop "replaces" a row of the second crop in forming the intercrop. Additive intercrops double the density, and therefore may use resources more completely.
Intercropping Concepts.
Duration refers to the temporal overlap of the intercrop components: Differing duration-usually combines a short season crop and a long season crop. Intercrops of differing duration are usually additive. Similar duration-competition more intense because both components are using resources at the same time. Intercrops of similar duration tend to be replacement types.
Intercropping Concepts.
Dominant vs. subordinate components. Typically, one crop component of the intercrop is more competitive and hence dominates the mixture in terms of growth and yield. Dominance may be due to: Rapid initial growth Height Photosynthetic pathway (C4 crops tend to be dominant when grown with C3 crops) Legumes are usually subordinate
Example of case
Pengelolaan agroekosistem
Pola Budidaya
Tahap subsisten Pada tahap ini petani mengusahakan lahan pertanian untuk memenuhi kebutuhan hidupnya dan kaum kerabatnya. Masukan seperti yang dilakukan pada pertanian modern belum dikenal, belum ada penggunaan bahan kimiawi sintetik. Umumnya produktivitas rendah, dan petani mengerjakan tanah garapannya dengan mengikuti irama musim dan daur alami.
Pola Budidaya Tahap eksploitasi Segala usaha diarahkan untuk mencapai nilai ekonomis terbaik atau tertinggi. Input atau masukan baik berupa sarana produksi atau masukan lainnya diusahakan tidak hanya optimal, kadang-kadang bahkan maksimal. Penggunaan teknologi amat intensif, bahan kimia pertanian amat diutamakan. Efisiensi juga diusahakan dengan mengatur daur bertanam dan perlakuan mengikuti kalender dan rutinitas manusia, karenanya
Pola Budidaya
Tahap eksploitasi Pertimbangan kondisi alami di pertanaman sering tidak diperhatikan. Pada awalnya usaha seperti ini memang memberikan hasil yang baik, sehingga petani atau penanam semakin dirangsang untuk terus menggunakan saranaprasarana produksi berbasis teknologi tinggi.
Pola Budidaya
Tahap kritis
Yang selanjutnya terjadi adalah tidak imbangnya agroekosistem, sehingga biaya usaha tani tidak akan dapat menghasilkan laba. Sebaliknya, biaya menjadi terlalu besar tetapi produk hanya sedikit, serta harganya bisa jatuh karena kualitasnya jelek/tidak disukai konsumen karena mengandung residu bahan kimia pertanian.
Pola Budidaya Tahap bencana Apabila tahap kritis tidak diantisipasi dengan baik, maka tahap berikutnya yang terjadi adalah bencana. Modal usaha tani tidak akan mendatangkan keuntungan, biaya yang dikeluarkan besar tetapi usaha yang dijalankan tidak mampu mengembalikan modal.
Pola Budidaya
Tahap pertanian ekologis/organik/sustainable
Jika petani/penanam ingin keluar dari bencana yang menimpa usaha taninya, maka harus dilakukan perubahan metode dalam penyelenggaraan budidaya tanamannya. Penerapan metode yang lebih akrab lingkungan seperti misalnya model PHT (Pengelolaan Hama Terpadu), pertanian organik, sistem tumpang gilir atau tumpang sari, budidaya lorong, sistem surjan dan berbagai cara bercocok tanam lainnya harus dan perlu dilakukan.
Pola Budidaya
Tahap pertanian ekologis/organik/sustainable
Cara bercocok tanam itu haruslah merupakan cara yang lebih mengabaikan sifat dan watak tanamannya, menjaga kelestarian media tanam, berorientasi pada produk berkualitas (dalam arti kandungan bahan bebas residu dan sisa yang bahan kimia pertanian membahayakan kesehatan).
CROP ECOLOGY